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Review of the Caribbean species of Dohrniphora Dahl (Diptera: )

Article in Journal of Natural History · February 2008 DOI: 10.1080/00222930601046493

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The user has requested enhancement of the downloaded file. Downloaded By: [University of Southern California] At: 21:52 24 January 2008 otro aeo h idfmraddaig fteml eiai.Bt ftenwseisare species new the of the Both species of genitalia. The micrographs male four. usual the electron the of scanning of drawings instead with setae and scutellar all femur six illustrate having hind in and the unusual of species, face known posterior previously three the aiba pce of species Caribbean with synonymized present: are species Five Trinidad. including not islands, Caribbean History Natural of Journal O:10.1080/00222930601046493 online DOI: 1464-5262 print/ISSN 0022-2933 ISSN CA 2006 Angeles, December Los 4 Boulevard, Exposition Published 900 County, [email protected] Angeles Email: Los of USA. Museum 90007, History Natural Kung, Giar-Ann Correspondence: genus The Introduction Keywords: of species the review we (Enderlein), material, collected newly and records previous on Based Abstract USA CA, Angeles, ( Los County, Angeles Los of Museum History Natural BROWN V. BRIAN & KUNG GIAR-ANN of species Caribbean the of Review hssuy iems oenrvsosof revisions Female modern most like study, This South materials essentially and is Methods newly and which material older projects. found of on survey species based fauna recent the 1995), from Disney (the treat by specimens herein covered Trinidad collected we partly excluding been revision, has this islands, and of American Caribbean part As the species. in described 80 causing over and with food, infesting houses, entering (Disney importance, myiasis. economic parasitoids human of and occasionally are is predators, that and larvae have facultative species, some One unknown, kleptoparasites, 1994). is species fungivores, most of scavengers, history life the Although worldwide. Dpea Phoridae) (Diptera: cetd2 etme 2006) September 28 Accepted eaecretyrvsn h e ol pce ftegns(ugadBon2005), Brown and (Kung genus the of species World New the revising currently are We Dohrniphora .divaricata D. Dohrniphora Caribbean, .divaricata D. Dohrniphora .cornuta D. r uhls itntv hntemls n aktems important most the lack and males, the than distinctive less much are Dohrniphora Adih n w e species, new two and (Aldrich) ali ag ru crety17seis fpoisfound phorids of species) 167 (currently group large a is Dahl 06 03–4:1931–1945 40(32–34): 2006; , n h loyeml dniidas identified male allotype the and , sprovided. is Bgt,hsbe cietlyitoue rudteworld, the around introduced accidentally been has (Bigot), # hrde Phoridae, , 06Tyo Francis & Taylor 2006 Dohrniphora .seriata D. pce,i ae nml specimens. male on based is species, Dohrniphora .cornuta D. and .cornuta D. .sexspinosa D. .cavifemur D. e otemlsof males the to key A . Dohrniphora (Bigot), eredescribe We . Dahl ogee is Borgmeier .dispar D. fthe of Downloaded By: [University of Southern California] At: 21:52 24 January 2008 ftealtp aeof male allotype with it the synonomize of hereby and Dominica, from treatments Museum, Species National States of United Museum MCZC, USNM, County; University; Institution. Angeles Smithsonian Harvard Los of Zoology, Museum Comparative History Map.’’ Natural ‘‘Global then LACM, and History; interest, of species the of name the on ‘‘ click type then can page, Users search www.discoverlife.org. the at available are records, their Caribbean facilitate to holotypes for barcode), cited a are with numbers (often barcode recognition. number usual The later unique the database. critical to a a addition with into alcohol, In entered labelled into pins. used and were collected specimens femur) to all were hind glued labels, study the or specimen point-mounted of this card face in either inner specimens and the point-dried, Most of recognition. sculpturing species and for setae (the characters diagnostic 99Dmnc ae,Brmirdsrbstemdcx sdr rw.W have We brown. same dark the yellow, as clearly legs. is coxa the coxa of mid mid his rest the In the and the 2). as localities, 1, (Figures same describes colour setae the peg-like from Borgmeier ventrobasal specimens five paper, examined to four Dominica of group 1969 a to addition in femur, cornuta Dohrniphora Diagnosis 1932 orihr divaricata Dohrniphora flaviventris Apocephalus bequaerti Dohrniphora venusta Dohrniphora mordax Phora chlorogastra Phora cleghorni Phora navigans Phora cornuta Dohrniphora cornuta (Dohrniphora) Diploneura cornuta Phora Syntypes: cavifemur Dohrniphora willowsi Dohrniphora crockeri Dohrniphora opposita Dohrniphora fulva Dohrniphora divaricata Dohrniphora h olwn olcinabeitosaeue:CN,Crei uemo Natural of Museum Carnegie CMNH, (www.morphbank.org). used: MorphBank are in abbreviations deposited collection be non- following will other, The species all as of well illustrations as SEM herein, cited records the on based distributions species of Maps eas xmndtehltp eaeof female holotype the examined also We .Kn n .V Brown V. B. and Kung G. „ , R io 87 4 Fec dto:15,p827). p 1857, edition: (French 348 p 1857, Bigot re 91 3,Fgr 1. Figure 531, p 1911, Brues ua(ai uem ntexamined). (not Museum) (Paris Cuba . io 80 191. p 1890, Bigot runed16,p154. p 1867, Frauenfeld ekr10,p32. p 1901, Becker atsAru12,p1,Fgr B lt iue1. figure Plate AB, Figure 12, p 1921, Abreu Santos ogee 90 7,Fgr 28. Figure 277, p 1960, Borgmeier : alc 92 432. p 1912, Malloch : a itntv iedroaa ea ntepseirfc ftehind the of face posterior the on setae dorsobasal fine distinctive has a ue 93 70. p 1933, Duzee Van a ue 93 70. p 1933, Duzee Van ogee 95 0,Pae2 iue6 90 277. p 1960, 6; Figure 2, Plate 107, p 1925, Borgmeier cmt 94 0,Fgr 1. Figure 105, p 1914, Schmitz ogee 99 – i at aeol) misidentification. only), male part, (in 1–2 p 1969, Borgmeier var. var. .cavifemur D. iv iura11,p1,Fgr 8. Figure 15, p 1916, Figueroa Silva ohihr cornuta Dorhniphora basalis obscura cmt 96 44. p 1926, Schmitz : Fgrs1 ,2,24) 23, 2, 1, (Figures atsAru12,p11. p 1921, Abreu Santos atsAru12,p11. p 1921, Abreu Santos oee,rsmlsta of that resembles however, , orihr cavifemur Dohrniphora .divaricata D. Bgt 1857) (Bigot, seblw.Tehn femur hind The below). (see .cornuta D. ogee,described Borgmeier, Dohrniphora xetfrthe for except , ’on ’’ Downloaded By: [University of Southern California] At: 21:52 24 January 2008 1 95 7 ( 1965, Survey Biodiversity Smithsonian Bredin-Archbold Spangler, J. 01 pi 92 .My (USNM). Mayr E. 1952, April 10–14 Bahamas examined Material Cosmopolitan. mm; 0.22 laterally, length cercus yellow Distribution Mean 1 white. Tergite abdomen 24). medially. of 23, Venter dark thicker (Figures markings yellow. five mm mostly 0.19–0.28 orange or 2 tergites range to four tergite Abdominal of of yellow setae. group half without a isolated anterior to or without addition with tibia with in 2) Hind brown, third, 1, setae. basal (Figures femur in peg-like hind setae 0.47–0.53. ventral of dorsal face range sparse Posterior length fine, length; yellow. yellow. short, Halter wing small, wing lobes 6.80–11.67:1.50–3.00:1.00. Mean 0.51 Postcoxal range yellow. length 8.96:2.14:1.00; posteriorly. Legs ratio costal pleuron dark sector of Mean costal slightly half mm. Mean Dorsal 1.62–2.25 posterior Anepisternum pair. present; range yellow. anterior setae mm; apical of 1.69 scutellar half five thickness of and ventral with pairs length orange, brown, Two times Palpus brown. 3.5 brown. dark approximately 1 pair same scutellum approximately Flagellomere and at setae. Scutum setae interfrontal setae. fronto-orbital ventral Ventral black. as to level brown dark Frons 0.50–0.59. Male. Description preparation. of artifact an be to shown lYnu,Ser eLqil,18.32 Luquillo, de Sierra Yunque, El (LACM). orchard citrus of character defining the thus and phenol, in placed when femora hind of the specimen A excavation’’. ‘‘dorsal – ac 90 .W it,Mlieta (USNM). trap Malaise Wirth, W. 62.57 W. 1970, March 1–8 Lucia aisn .Kige,W ao,J aln,S hmsn 5 (CMNH). m 355 Thompson, S. Rawlins, J. 17.35 Zanol, W. Klingler, M. Davidson, pigil sae . mEECnfed 15.35 1 Canefield, River, ENE 1 Figues km 2.5 Gutter, Fond Estate, Manets BASBS; Springfield ft; trap, 400 Malaise Gagne, Wirth, J. W. R. W. 1965, March 20 aln,S .Topo,40m ot hlenEtt,1 Estate, Chiltern South m; 450 Thompson, A. S. Rawlins, .Rwis .A hmsn 0m(CMNH). m 50 Thompson, A. S. Rawlins, E. 18.10 Rojo, Cabo N km 26 m, 540 forest, deciduous trap, intercept 71.63 1 (USNM). Hill, BASBS Woodford BASBS; Wirth, uy18,J .Rwis .L aisn 20m(CMNH). m 18.15 1250 Rojo, Davidson, L. Cabo R. NNW Rawlins, km C. 38 J. Abeja, 1987, La July m, 730 forest, 1 deciduous 1990, July 20 „ 02 pi 94 .S ln,lgtta altp of (allotype trap light Flint, S. O. 1964, April 10–20 , u u u neGlt mSWAs aRy,13.93 Raye, La Anse SSW km 1 Galet, Anse : ,1 W, ,62.73 N, ,2 W, enbd egh17 m ag .222 m enfotlrto05;range 0.54; ratio frontal Mean mm. 1.62–2.25 range mm; 1.70 length body Mean „ 13 aur 94 3 1964, January 21–31 , ot iiiIln,1 Island, Bimini South : „ „ 31 uy19,11 1990, July 13–19 , 8Ags o2 etme 95 .Coe .Bacet,Mlietrap, Malaise Blanchette, T. Coote, L. 1985, September 21 to August 28 , u ,1 W, „ 32 uy19,L anr .Rwis .Yug necp trap, intercept Young, C. Rawlins, J. Masner, L. 1990, July 13–25 , oiia Republic Dominican „ –7Ags 95 .Coe aas rp 0 (LACM). m 300 trap, Malaise Coote, L. 1985, August 1–27 , uroRico Puerto „ 12 eray16,1 1965, February 11–20 , „ „ „ uy15,2 1951, July , .divaricata D. 92 uy19,L anr .Rwis .Young, C. Rawlins, J. Masner, L. 1990, July 19–25 , 7Fbur 95 .W it,sapforest, swamp Wirth, W. W. 1965, February 27 , „ Dominica i rne lVreSain . mWWPico WNW km 2.1 Station, Verde El Grande, Rio : ac 95 .W it,BSS an Paul, Saint BASBS; Wirth, W. W. 1965, March 5 , u ,65.82 N, eenls 35k aoRj,18.10 Rojo, Cabo N km 23.5 Pedernales, : aiba pce of species Caribbean u neBueu 1 Bouleau, Anse : u ,61.37 N, eeaie a iia itrinof distortion similar a had examined we ,1 W, „ u uut15,C n .Vui,1 Vaurie, P. and C. 1951, August , ,61.05 N, „ .cavifemur D. Nevis – ue19,C on,R. Young, C. 1996, June 3–6 , u ,1 W, „ „ Jamaica u –0Mrh16,1 1965, March 1–10 , ,1 W, eray16,W W. W. 1965, February 2 , aet sae 17.12 Estate, Dasents : „ 5 11 ue19,J E. J. 1991, June 11–18 , „ „ 1933 Dohrniphora AB) lreHall, Clarke BASBS); u 0Otbr16,P. 1964, October 10 , u ,71.63 N, 13 ue19,J. 1991, June 21–30 , ,4 ), ,71.63 N, uaa a,1 Bay, Runaway : „ 12 January 11–20 , .cavifemur D. „ pi 1966, April , u an Kitts Saint ,9 W, u ,1 W, „ „ Saint „ 13– , 11– , ,15 u u N, N, „ „ is , : , Downloaded By: [University of Southern California] At: 21:52 24 January 2008 .204.Fosbon eta rnoobtlstea prxmtl aelvlas level same approximately at setae fronto-orbital Ventral brown. Frons 0.42–0.48. 60 Male. approximately 4). of 3, (Figures group species a other Description to from posterior it femur, distinguish hind help circular the also dark, setae of distinct peg-like The face 19). ventrobasal posterior (Figure 5 the tarsomere on fore carina darkened and broad unusually the dispar Dohrniphora Diagnosis examined). (not Poland LC N 219829]. ENT [LACM 1934 iue –.Pseirfc fhn eu.(,2) (1, femur. hind of face Posterior 1–4. Figures Holotype: ilnua(orihr)dispar (Dohrniphora) Diploneura dispar Dohrniphora hr dispar Phora enbd egh23 m ag .728 m enfotlrto04;range 0.46; ratio frontal Mean mm. 1.97–2.81 range mm; 2.33 length body Mean .Kn n .V Brown V. B. and Kung G. „ rzl at aaia Lu Catarina, Santa Brazil: . neli 92 9( 29 p 1912, Enderlein cmt 93 55–56. p 1923, Schmitz : srcgie yteoag,elre lgloee1(iue 1 2 and 22) 21, (Figures 1 flagellomere enlarged orange, the by recognized is ohihr dispar Dorhniphora Fgrs3 ,1,2,2,2,26) 25, 22, 21, 19, 4, 3, (Figures ogee 95 3,Fgrs2 ,5 lt ,Fgr 4. Figure 5, Plate 5, 3, 2, Figures 431, p 1935, Borgmeier : „ Hamo ; drad,Pls cdm fSine Warszawa, Science, of Academy Polish ¨derwaldt, orihr cornuta Dohrniphora na at Catarina). Santa ˆnia, Edren 1912) (Enderlein, LC N 831.(,4) (3, 186301]. ENT [LACM .dispar D. Downloaded By: [University of Southern California] At: 21:52 24 January 2008 6 7 1965, 1 BASBS, Wirth, ( Survey 1 Biological Smithsonian Hall, Bredin-Archbold Gagne, R. 1966, April Dominica examined Material Indies. West Ecuador, Panama, Brazil, range setae. Distribution dark mm; 0.27 medially, isolated length cercus yellow Mean without margins, yellow. 26). to posterior 25, tibia (Figures white black, and abdomen mm dark, 0.23–0.30 of Hind anterior to with 19). Venter (Figure basal, yellow 4) laterally. 60 setae. brown brown with 3, enlarged approximately (Figures of tergites peg-like dark femur pulvilli group Abdominal hind to 5 curve-tipped, point; apical of concavity, face tarsomere apical apically-rounded, surrounding Posterior carina medial, Fore circular Halter length developed. ventral not with yellow. wing 7.50–12.50:1.57–2.50:1.00. lobes apex, Mean coxae, range Postcoxal 0.47–0.52. towards range pair. 10.47:2.22:1.00; including length; pair broadening anterior wing ratio Legs, Posterior 0.49 of sector setae. length yellow. thickness costal scutellar costal Mean and of mm. Mean 2.19–2.62 pairs length yellow. range pleuron two times mm; and 2.30 with Scutum 3.5 setae. yellow, apical approximately five sometimes with length brown, orange, one-half Palpus approximately Scutellum 22). enlarged, 21, (Figures orange, eye 1 of Flagellomere setae. interfrontal ventral 1 1 1966, March 1 6k aoRj,18.10 m; 540 forest, Rojo, deciduous trap, Cabo intercept Young, N C. Rawlins, km J. 26 Masner, L. 1990, July 19–25 leil (MCZC). illegible 97 .L .Kas (USNM). Krauss H. L. N. 1967, Pedernales, 6 18.10 forest; m, 540 lowland Rojo, Thompson, mesic Cabo Young, C. N Thompson, km S. 23.5 Rawlins, J. Davidson, R. 1992, aur 95 .W it,BSS 2 BASBS, Wirth, W. W. 1965, January 1 96 .J an,12m AB;muho ogsRvr 1 River, Hodges of mouth BASBS; m, 122 Gagne, J. R. 1966, it,sapfrs,BSS aesGte,2 Gutter, Manets BASBS; forest, swamp Wirth, .W it,sra e,3 bed, stream Wirth, W. W. aln,C on,itretta,wtdcdosfrs,70m(CMNH). m 730 forest, deciduous wet trap, 1 intercept Gap, Hardwar Young, C. Rawlins, ac 95 .W it,BSS aeuCif,L acet,1 Fanchette, La Cliffs, 2 Tareau Falls, BASBS; Trafalgar Gagne; Wirth, W. W. 1965, March 0Fbur 95 .W it,BSS(AM USNM). (LACM, BASBS Wirth, W. W. 1965, February 20 ao,Pru o atss mWCead rn,19.07 Arena, de Cueva W km 3 Haitises, Los Parque Mayor, „ „ „ „ 13 aur 95 .W it,Mlieta,BSS lr alEtt,1 Estate, Hall Clark BASBS; trap, Malaise Wirth, W. W. 1965, January 21–31 , 2My16,1 1966, May 12 , 0Jn o4Jl 02 .Kaoe,Mlieta,62f;WieRvr 1 River, White ft; 632 trap, Malaise Krakower, A. 2002, July 4 to June 30 , ue16,1 1966, June 7 , „ „ –0Jnay16,11 1965, January 8–10 , 12 eray16,3 1965, February 21–28 , nrm 1 Antrim, : „ pi 96 5 1966, April 6 , „ „ 0Mrh17,WrhadFr;RnwyBy 12 Bay, Runaway Farr; and Wirth 1970, March 10 , Montserrat 12 aur 95 1 1965, January 11–20 , „ „ „ 9My16,1 1966, May 29 , 3Jn 96 .Sesa,BSS odFge ie,1 River, Figues Fond BASBS; Steyskal, G. 1966, June 13 , „ 31 uy19,5 1990, July 13–19 , 0Mrh15,J .G lre 0 ;pt oCbis 1 Cabrits, to path m; 305 Clarke, G. F. J. 1956, March 10 , „ – pi 96 .J an,36m ot hlenEtt,3 Estate, Chiltern South m; 366 Gagne, J. R. 1966, April 5–6 , „ u ,71.63 N, u – ac 90 .W it UN) 1 (USNM); Wirth W. W. 1970, March 1–8 , ,71.63 N, asv hu,BateHue 16 House, Beattie Ghaut, Cassava : „ „ 4Arl16,5 1966, April 24 , „ 12 ac 95 3 1965, March 11–20 , 13 ac 95 .W it,lgtta,BASBS, trap, light Wirth, W. W. 1965, March 21–31 , uroRico Puerto „ u 0My16,7 1966, May 30 , u ,20 W, „ ,2 W, pi 96 1 1966, April , „ 32 uy19,12 1990, July 13–25 , „ 13 aur 95 2 1965, January 21–31 , „ „ „ aiba pce of species Caribbean 3Jl 90 .Yug .Rwis S. Rawlins, J. Young, C. 1990, July 13 , 32 uy19,15 1990, July 13–20 , ac 95 6 1965, March 1 , sbl,Bsu saa eGuajataca, de Estatal Bosque Isabela, : „ 7Arl16,R .Gge BASBS, Gagne, J. R. 1966, April 27 , „ „ ue16,1 1966, June 4 , „ 2Arl16,3 1966, April 12 , 13 ac 95 .W. W. 1965, March 21–31 , oiia Republic Dominican „ u 7Fbur 95 .W. W. 1965, February 27 , ,69.48 N, „ u „ orihr 1935 Dohrniphora „ 45.91 92 uy19,8 1990, July 19–21 , ac 95 3 1965, March 5 , 3Arl16,R J. R. 1966, April 13 , „ „ 5 „ u eray1969, February , 9 –0February 1–10 , ,6 W, AB) Clarke BASBS); .Mse,J. Masner, L. , ,62 N, „ ue1966, June 6 , „ „ te data other , November , „ „ u 9April 29 , – July 7–9 , Jamaica 12.95 Hato : „ „ „ ,25 „ ,28 9 W, ,2 ,1 „ „ , : , Downloaded By: [University of Southern California] At: 21:52 24 January 2008 otnsAmmn 18.42 Aymamon, Montanas 1936 iue –.Pseirfc fhn eu,vrainin variation femur, hind of face Posterior 5–8. Figures LC N 156.(,8 oiia[AMET210539]. ENT [LACM Dominica 8) (7, 210546]. ENT [LACM aisn .Yug .Kige,S hmsn oet 1 CN) aauz 1 Mayaguez, (CMNH); m 210 forest, Thompson, S. Klingler, M. Young, C. Davidson, –2My13,G .Tloh(SM;RoGad,E ed tto,31k WNW km 3.1 Station, Verde El Grande, 10.32 Rio Luquillo, de (USNM); Sierra Tulloch Yunque, S. El Pico G. 1930, May 5–12 aisn .Kige,W ao,J aln,S hmsn 5 CN) Utuado, (CMNH); 18.33 m 355 Bocas, Dos Thompson, W S. km 4.9 Rawlins, Abajo, Rio J. de Zanol, Estatal W. Bosque Klingler, M. Davidson, (examined). synonomy. m 380 Klingler, M. Thompson, S. Zanol, W. Davidson, R. Rawlins, J. (CMNH). Young, C. Holotype: cavifemur Dohrniphora diffusa Dohrniphora cornuta Dohrniphora venusta Dohrniphora divaricata Dohrniphora divaricata Phora .Kn n .V Brown V. B. and Kung G. „ an icn:n te aa(SMn.76;LC N 219927) ENT LACM 7765; no. (USNM data other no Vincent: Saint . lrc 86 437. p 1896, Aldrich ogee 90 8,Fgr 35. Figure 281, p 1960, Borgmeier cmt 99 2 5 32. 25, 22, p 1929, Schmitz : re 95 95. p 1915, Brues : ogee 91 111. p 1961, Borgmeier : ogee 99 –,Fgrs1 i at eaeol) new only), female part, (in 2 1, Figures 1–2, p 1969, Borgmeier ohihr divaricata Dorhniphora u ,66.96 N, Fgrs51,2,2,28) 27, 20, 5–12, (Figures u ,1 W, u ,65.82 N, „ 41 ue19,J aln,W ao,R. Zanol, W. Rawlins, J. 1996, June 14–15 , orihr divaricata Dohrniphora Adih 1896) (Aldrich, u ,2 W, u „ ,66.72 N, – ue19,C on,R. Young, C. 1996, June 3–6 , 5 )Prtp,SitVincent Saint Paratype, 6) (5, . u ,1 W, „ 7Jn 1996, June 17 , „ , Downloaded By: [University of Southern California] At: 21:52 24 January 2008 mn aiba species, Caribbean Among Diagnosis 210822]. ENT [LACM in Rico variation Puerto femur, 12) hind (11, of 210487]. face Posterior 9–12. Figures ogee stemi hrce fti pce,i natfc fpeaain(e Diagnosis (see preparation of artifact an is species, this of of section character main the as Borgmeier pattern) triangular or curved a of in female a probable (sometimes to a row addition in the margin, to ventral peg-like dorsal the of along setae pattern 5–12). rows (Figures scattered and of two number 22–34 The usually group approximately femur. are larger to hind there the apical although of depression vary, face setae posterior shallow the a on of setae peg-like presence the and setae, ee svnrlitrrna ea.Faelmr rw.Plu rne ihfv apical five with orange, Palpus brown. 1 Flagellomere same setae. approximately at interfrontal setae ventral fronto-orbital Ventral as brown. level dark to brown Frons 0.47–0.54. Male. Description h ubro eglpae n ooro h i oaalwdu omthi ihother with it match to us allowed coxa mid the of of colour specimens and plates Caribbean tergal of number the eeaie h ooyefml of female holotype the examined We enbd egh20 m ag .625 m enfotlrto05;range 0.51; ratio frontal Mean mm. 1.66–2.50 range mm; 2.09 length body Mean .cornuta D. ). .divaricata D. .divaricata D. orihr divaricata Dohrniphora lhuhtxnm of taxonomy Although . orihr cavifemur Dohrniphora h itrino h idfmr ie by given femur, hind the of distortion The . orihr divaricata Dohrniphora aiba pce of species Caribbean sdfeetae yisfu scutellar four its by differentiated is Dohrniphora ogee n on tt be to it found and Borgmeier 9 0 an it LC ENT [LACM Kitts Saint 10) (9, . orihr 1937 Dohrniphora sbsdo males, on based is Downloaded By: [University of Southern California] At: 21:52 24 January 2008 95 .W it,lgtta;BSS(SM;1 (USNM); BASBS trap; light Wirth, W. W. 1965, hmsn .Rwis osa eeaino ietn,1 CN) aEtelt,Rio Estrelleta, La 18.10 near (CMNH); Rojo, m Haitises, 10 limestone, Los 1 on Parque Limpio. vegetation coastal Mayor, Rawlins, Hato J. 19.67 Thompson, LACM), Arena, (CMNH, de m 20 Cueva forest, lowland mesic 19.67 Republic Dominican AB UN) an al pigil sae . mEECnfed 15.35 Canefield, ENE km 2.5 Estate, Springfield Paul, 1 Saint 61.37 Cabrits, to (USNM); path (USNM); BASBS BASBS Steyskal, G. 1966, May 2 (USNM); 1 (USNM); BASBS 1 (USNM); BASBS cavifemur eray16,D .Ba UN) 1 (USNM); Bray F. D. 1964, February rhodSihoinBoiest uvy( Survey Biodiversity Smithsonian Archbold Dominica examined Material Indies. West Trinidad, Rica, Costa Brazil, Argentina, here shown are Distribution variants the of Some islands. individual whether to and 5–12). component, (Figures restricted is geographical and are Sampling strong 6), a Rica). Specimens patterns 5, has Costa (Figures extensive. variation from certain setae this example is of if (for femur number determine seen to have largest hind we insufficient the the specimens have of mainland Vincent to face Saint similar posterior are in the locality type of the setae from the in variation The length cercus Mean white. Variation abdomen of 28). 27, Venter (Figures with Abdominal mm medially. 0.20–0.30 setae. also range isolated yellow below); mm; without 0.26 Variation, brown, tibia in Hind and dark setae. discussion femur to of peg-like tergites further margin posterior basal (see ventral ratio depression of along setae round rows group shallow peg-like sector two triangular dorsal with yellow. large usually costal more developed, but with well several varies, lobes 5–12) Mean setae (Figures Postcoxal of femur number brown. 0.48–0.54. setae; hind dark coxa of range mid face brown; white, Posterior light length; mm. coxae 1.50–2.25 mostly hind range Legs wing twice and white. mm; fore pair 1.92 Halter 0.51 pleuron length posterior 7.00–11.33:1.67–2.40:1.00. of wing range present; length half 9.50:2.24:1.00; Mean setae Ventral pair. costal scutellar brown. anterior of to Mean of pairs brown thickness Two light and brown. pleuron length dark of Scutellum half dorsal white. and Scutum setae. 4 ,2 mNCb oo 18.10 Rojo, Cabo N km 26 m, 540 1 1938 „ 92 uy19,L anr .Rwis .Yug necp rp eiuu forest, deciduous trap, intercept Young, C. Rawlins, J. Masner, L. 1990, July 19–25 , u u ,69.48 N, ,2 W, ,2 ), .Kn n .V Brown V. B. and Kung G. „ artSap 1 Swamp, Cabrit : 5Ags 90 .Nrbm 5 CN) eenls 35k Cabo N km 23.5 Pedernales, (CMNH); m 650 Norrbom, A. 1980, August 15 , u „ „ ,71.63 N, „ ue16,1 1966, June 4 , 11 ue19,J .Rwis .A hmsn 5 (CMNH). m 450 Thompson, A. S. Rawlins, E. J. 1991, June 11–18 , 13 aur 95 1 1965, January 21–31 , u ,14 W, „ „ u 0Mrh16,1 1966, March 30 , 12 eray16,1 1965, February 21–28 , „ ,4 N, aoMyr aqeLsHiie,3k uv eArena, de Cueva W km 3 Haitises, Los Parque Mayor, Hato : – uy19,R aisn .Rwis .Topo,C Young, C. Thompson, S. Rawlins, J. Davidson, R. 1992, July 7–9 , u ,69.47 N, „ 31 uy19,2 1990, July 13–19 , „ ue16,G tykl AB UN) ao,1 Layou, (USNM); BASBS Steyskal, G. 1966, June 7 , „ 3Fbur 95 .W it,lgtta,Bredin- trap, light Wirth, W. W. 1965, February 23 , u u ,6 W, ,71.63 N, „ „ 13 ac 95 .W it,Mlietrap, Malaise Wirth, W. W. 1965, March 21–31 , ue16,O .Fit ih rp(ooyeof (holotype trap light Flint, L. O. 1964, June , „ „ – uy19,C on,R aisn S. Davidson, R. Young, C. 1992, July 7–9 , pi 96 .J an,a ih,BASBS light, at Gagne, J. R. 1966, April 3 , u ,1 W, 5 „ AB)(SM;Cak al 1 Hall, Clarke (USNM); BASBS) „ 32 uy19,4 1990, July 13–25 , –0Mrh16,1 1965, March 1–10 , „ „ 32 uy19,itretta,wet trap, intercept 1990, July 13–20 , –0Fbur 95 .W Wirth, W. W. 1965, February 1–10 , „ pi 96 .J Gagne, J. R. 1966, April 2 , „ 92 uy1990, July 19–21 , „ 13 March 21–31 , „ „ ,29 u ,4 N, D. Downloaded By: [University of Southern California] At: 21:52 24 January 2008 eiuu oet 3 CN,LACM). (CMNH, m Jamaica 730 forest, deciduous 5Fbur 99 .W it,mrh(SM;RnwyBy 1 Bay, Runaway (USNM); marsh Wirth, W. W. 1969, February 25 99 .W it UN) uaa a,11 Bay, Runaway (USNM); Wirth W. W. 1969, encru egh02 m ag .903 m(iue 9 30). isolated 29, without (Figures yellow. tibia mm abdomen 0.19–0.30 Hind of range setae. Venter mm; small, of 0.26 medially. four yellow row length or brown, ventral 14). cercus three 13, dark of Mean (Figures and mostly apical line trochanter tergites straight slightly near Abdominal a and tooth setae. in to basal not dorsal with row, or also the setae one in femur with thin setae specimens hind only of some of specimens rest 16); face to straight 15, some Posterior (Figures dorsal relatively or setae in with ventral peg-like femur although setae five hind two or dark, of four face femur of Posterior row hind developed. ventrobasal of length; not lobes wing Apex Postcoxal 0.51 yellow. faintly. length Legs 6.50–9.50:1.83–3.25:1.00. costal range white. 7.50:2.18:1.00; Mean subequal. ratio Halter pairs mm. times sector 1.44–2.09 posterior costal 3.5 range Mean and approximately 0.49–0.54. anterior mm; range 1.83 pair setae. pairs; length posterior middle apical and wing setae; five anterior Mean of with of pairs yellow, thickness three and Palpus length with brown. Scutellum to ventral yellow. orange as Thorax 1 level same Flagellomere approximately setae. at setae interfrontal fronto-orbital Ventral brown. Frons 0.50–0.54. Male. Description neGlt mSWAs aRy,13.93 Raye, La Anse (LACM). SSW m 400 km 1 field/forest, old 17.35 Galet, trap, Level, Anse Malaise Phillips Coote, m; 300 D. rainforest, L. trap, Malaise Coote, D. 62.77 L. 1985, August 1–27 3 Bay, Runaway (USNM). bed; stream USNM), aln,S .Topo,5 (CMNH). USNM). m 50 (LACM, Thompson, A. S. Rawlins, rne lVreSain . mWWPc lYnu,Ser eLqil,10.32 Luquillo, de Sierra LACM). Yunque, (CMNH, El m 355 Pico Thompson, WNW km 3.1 65.82 Station, Verde 1 El Maricao, Grande, (MCZC); m 640–670 Rico Puerto orihr seriata Dohrniphora Diagnosis rue Fgrs1,1) o narw(sin the in (as setae row from a peg-like in differs ventrobasal not six it 18), 17, the (Figures figures), and on grouped There setae row, (including setae. peg-like straight peg-like ventrobasal description ventrobasal five relatively or scattered the four 14 only approximately in are having in information species the Caribbean on based scutellaris World New known u u ,4 W, ,5 W, enbd egh19 m ag .125 m enfotlrto05;range 0.52; ratio frontal Mean mm. 1.41–2.50 range mm; 1.95 length body Mean ada a,1 Gap, Hardwar : nBrmirs16 e.W i o xmn n pcmn of specimens any examine not did We key. 1960 Borgmeier’s in Nevis „ „ 95 .D ot,Mlieta;WnfedMn,2 Mtn., Wingfield trap; Malaise Coote, D. L. 1985, , – ue19,C on,R aisn .Kigr .Znl .Rwis S. Rawlins, J. Zanol, W. Klinger, M. Davidson, R. Young, C. 1996, June 3–6 , lYnu,2 Yunque, El : ecsl,2 Newcastle, : , Dohrniphora .sexspinosa D. „ orihr seriata Dohrniphora 0Mrh17,WrhadFr UN) not a,1 Bay, Annotto (USNM); Farr and Wirth 1970, March 10 , „ 52 eray16,T n .Haa,L emnJr., Herman L. Hlavac, B. and T. 1969, February 15–24 , „ uyt uut19,B ukad a rp(LACM). trap pan Buckland, B. 1994, August 3 to July 9 , pce ihsxsuelrste l he pce e to key species three All setae. scutellar six with species Fgrs1–6 9 30) 29, 13–16, (Figures e pce,and species, new „ uy16,J adnd . T(SM;Rio (USNM); LT C., Maldonado J. 1960, July , an Kitts Saint „ – ac 90 .W it,Mlietrap Malaise Wirth, W. W. 1970, March 1–8 , an Vincent Saint .seriata D. u ,61.05 N, Grenada aiba pce of species Caribbean „ .seriata D. eray16,W .Wrh(LACM, Wirth W. W. 1969, February , rehl,14 Greenhill, : e species new .scutellaris D. o ctee a in (as scattered nor ) u :1 ,2 W, :3 Fgrs1–6,arne na in arranged 13–16), (Figures „ an Lucia Saint „ oohrdt (MCZC). data other no , „ oohrdt (paratypes) data other no , 13 ue19,J E. J. 1991, June 21–30 , „ „ .sexspinosa D. ogee r h only the are Borgmeier 23 uy18,88 1985, July 12–31 , –0Nvme 1985, November 1–30 , orihr 1939 Dohrniphora „ 62 February 16–28 , .scutellaris D. neL Raye, La Anse : .scutellaris D. r closely are but , u u N, N, D. „ „ ). , , Downloaded By: [University of Southern California] At: 21:52 24 January 2008 1940 otro aeo h idfemur. hind the of face posterior word Latin the From Etymology Islands. Virgin US Rico, Puerto Islands, Virgin British Distribution 14) (13, femur. hind of face Posterior 13–18. Figures seriata LC N 189.(7 18) (17, 210889]. ENT [LACM .Kn n .V Brown V. B. and Kung G. series enn o,rfrigt h etoaa o fsteo the on setae of row ventrobasal the to referring row, meaning , .sexspinosa D. LC N 179771]. ENT [LACM orihr seriata Dohrniphora LC N 855.(5 16) (15, 184595]. ENT [LACM D. Downloaded By: [University of Southern California] At: 21:52 24 January 2008 Holotype: uy1 etme 94 .Ii ta.FT(AM broe AMET027609]. ENT LACM [barcode: (LACM) FIT al. et Ivie M. Paratypes: 1994, September July–15 oet e ee,08 UN) lnainCsen 18.49 Cistern, Plantation (USNM), m 0–80 level, sea forest, 1 Bay, North Miller, 93 .R nlig aas rp ht ec,18.47 Beach, White trap, Malaise Snelling, R. R. 1993, aeo h idfemur. hind the of face words Latin the of Combination Etymology Republic. Dominican mm 0.23–33 range Distribution line mm; tooth, 0.29 yellow apical length thin cercus and without Mean tooth, or yellow. tibia basal with 32). Hind 31, abdomen brown, margin, (Figures microscopy). of dark light ventral under tergites Venter see of faces. Abdominal medially. to base posterior difficult setae. (teeth at and isolated setae setae without of anterior group peg-like on with the 18) to six black, 17, dorsoapical (Figures of almost femur hind group of brown, face a Posterior dark rounded. range yellow, femur present; mm; yellow, 2.22 lobes hind Postcoxal mostly sector length of costal Legs and wing Mean white. apex Anterior 0.49–0.53. Halter Mean except range 7.67–10.75:2.67–3.75:1.00. pairs. yellow. length; range other wing Pleuron setae. 8.54:3.11:1.00; 0.52 of ratio scutellar size. length thickness costal of in Mean and pairs equal mm. 2.00–2.41 length three at about times with pairs setae 2.5 Scutellum posterior fronto-orbital brown. approximately apical Ventral five light pair or scutellum four brown. Middle with and yellow, to Palpus Scutum setae. orange interfrontal setae. ventral 1 as level flagellomere same approximately and Frons 0.48–0.51. Male. (See Description differs. femur hind the on setae peg-like of ventrobasal section the Diagnosis of setae. number peg-like and ventrobasal arrangement six of group scutellaris close and setae, scutellar . mWWPc lYnu,Ser eLqil,18.32 Luquillo, de Sierra Yunque, (LACM). El trap Pico WNW Malaise km 3.1 Liao, P. W. 2002, orihr sexspinosa Dohrniphora Diagnosis m 355 Thompson, S. (LACM). Rawlins, FIT al. J. et Ivie Zanol, M. 1994, W. September 15 Klingler, to M. July Davidson, R. (CMNH). Young, C. enbd egh24 m ag .227 m enfotlrto05;range 0.50; ratio frontal Mean mm. 2.12–2.72 range mm; 2.44 length body Mean nBrmirs16 e eas ftepeec fsxsuelrste u the but setae, scutellar six of presence the of because key 1960 Borgmeier’s in „ SVri Islands Virgin US rts ignIslands Virgin British . SVri Islands Virgin US „ .seriata D. 52 uy18,S .adP .Mle,Mlietrap, Malaise Miller, M. P. and E. S. 1985, July 15–23 , sesl itnuse rmother from distinguished easily is orihr sexspinosa Dohrniphora .) an hms saeBtn a,18.35 Bay, Botany Estate Thomas: Saint : sex Fgrs1,1,3,32) 31, 18, 17, (Figures an hms saeBtn a,18.35 Bay, Botany Estate Thomas: Saint : un sad 1 Island: Guana : and spinosus uroRico Puerto aiba pce of species Caribbean eern otesxsteo h posterior the on setae six the to referring , e species new „ –4Jl 94 .E n .M. P. and E. S. 1984, July 1–14 , u u i rne lVreStation, Verde El Grande, Rio : ,65.82 N, ,64.67 N, Dohrniphora u ,64.58 N, .sexspinosa D. u u orihr 1941 Dohrniphora ,1 N, ,1 W, u ,1 W, „ u pce yissix its by species „ ,64.92 N, – ue1996, June 3–6 , –0October 2–10 , „ u 51 April 15–18 , ,64.92 N, esto keys Coccoloba u ,2 W, u W, D. „ , Downloaded By: [University of Southern California] At: 21:52 24 January 2008 381,632]. iue 9 0 oetroee5 (19) 5. tarsomere Fore 20. 19, Figures 1942 .Kn n .V Brown V. B. and Kung G. iue 1 22. 21, Figures orihr dispar Dohrniphora orihr dispar Dohrniphora ed 2)Fotve.(2 aea view. Lateral (22) view. Front (21) head. , LC N 173.(20) 219743]. ENT [LACM .divaricata D. [CMNH Downloaded By: [University of Southern California] At: 21:52 24 January 2008 oet 4 CN)[acd:LC N 178.Paratypes: 219718]. ENT LACM [barcode: (CMNH) Republic m 540 forest, 28) 1 71.63 Holotype: 24) (23, views. lateral left and right genitalia, Male 23–32. Figures ctlu ihsxste...... setae . six with Scutellum . setae four with – Scutellum 1. LACM). (CMNH, m 540 forest, deciduous trap, intercept Young, „ 32 uy19,6 1990, July 13–25 , .divaricata D. u ,1–1Jl 90 .Mse,J aln,C on,itretta,deciduous trap, intercept Young, C. Rawlins, J. Masner, L. 1990, July 19–21 W, eenls 35k aoRj,18.10 Rojo, Cabo N km 23.5 Pedernales, : „ . 2,30) (29, . oiia Republic Dominican .seriata D. „ 92 uy19,4 1990, July 19–21 , e omlso Caribbean of males to Key 3,32) (31, . .sexspinosa D. eenls 35k aoRj,18.10 Rojo, Cabo N km 23.5 Pedernales, : ...... 4 ...... 2 „ 92 uy19,L anr .Rwis C. Rawlins, J. Masner, L. 1990, July 19–25 , . aiba pce of species Caribbean Dohrniphora u ,71.63 N, orihr cornuta Dohrniphora u ,2 W, orihr 1943 Dohrniphora „ 2,26) (25, . 31 uy1990, July 13–19 , Dominican .dispar D. (27, . u N, Downloaded By: [University of Southern California] At: 21:52 24 January 2008 .Pseirfc fhn eu ihsas oslste etoaa einwt only with region ventrobasal setae; dorsal sparse with femur hind of face Posterior 2. 1944 lrc M 86 oihpdde hrde n ilso W dtr nteDpeao t icn (West Vincent St. of Diptera the On editor. SW, Williston In: Phoridae. Dolichopodidae; 1896. JM. Aldrich re T 95 yoyi aaou ftedpeosfml hrde ultno h icni Natural Wisconsin the Hall. & of Chapman Bulletin London: Phoridae. Phoridae. —the family Hungarici Scuttle dipterous Nationalis 1994. the Musei RHL. of Annales Disney catalogue Sauter. synonymic H. A Mr. 1915. by CT. collected Formasa Brues Dominica of of Phoridae Phoridae The the 1911. Dominica: CT. of Brues Survey Biological Bredin-Archbold-Smithsonian 1969. T. Borgmeier su beschriebenen 1912 Jahre im Enderlein G. Prof. Nacional Museu von do der Archivos typen Phoridae. Die familia 1935. da T. conhecimento 7. Borgmeier o Volume Cuba. para de subsidios Isla Novos la de 1925. natural T. y Borgmeier politica [ fisica, 1890. Historia J. editor. Bigot la, 1:1–100. de Wien R in Sagra Gesellschaft In: Zoologisch-Botanischen Dipteros. der 1857. Abhandlungen J. Phoriden. Bigot Die 1901. T. Becker ogee .16.WieeBeitra Weitere 1961. T. Borgmeier ogee .16.Geflu 1960. T. Borgmeier otro aeo idfmrwtotdra ea;vnrbslrgo ihmany with region ventrobasal setae; dorsal without femur hind of face Posterior – .Faelmr rne nagd(iue 1 2;fr asmr ak broad dark, 5 tarsomere fore 22); 21, (Figures enlarged orange, 1 Flagellomere 3. lgloee1bon o nagd oetroee5sm oora eto leg, of rest as colour same 5 tarsomere fore enlarged; not brown, 1 Flagellomere – rn E-262) n .Coefrhslrednto fseiesfo an Kitts Saint recent from the specimens of of on donation much work large Our for his for Nevis. Museum (NSF) for Coote Foundation and Carnegie L. Science Berezovskiy National and the by material V. DEB-0206520), (supported of of grant and Republic donation Dominican Young or drawings the loan C. from line for material especially individuals the several herein, preparing to grateful studied skillfully row are for We a assistance. Koehler in technical B. not thank together, We grouped setae six Acknowledgements with femur hind of five or face four of Posterior row ventrobasal straight relatively with – femur hind of face Posterior 4. rw n .Sih ucaeo E tteLC a uddb S rn DBI- grant NSF by funded References was LACM the at SEM a of Purchase 0216506. Smith. P. and Brown itr oit 12:85–152. Society History 9:530–559. 23:1–69. Zoology to Contributions Smithsonian (Diptera). 25:85–281. 328–349. p Bertrand, A. Paris: 108–119), Figures 12, (Plate 1896:309–345 London of Society Entomological 435–438. the of Transactions Indies). Dohrniphora ibnneu sieben 5:427–470. Entomologia de Revista Phoridae). (Dipt. Phoriden oro iesot hc e-ieste(iue ,2 . . . . 2) 1, (Figures setae peg-like thick short, five or four mr hn2)ln,tikpglk setae peg-like thick long, 20) than (more Fgr 9;pseirfc fhn eu ihdr,crua eta aiaaia to apical carina ventral 4) circular 3, dark, (Figures with setae femur ventral hind of face posterior 19); (Figure hlo eta ersinaia ovnrlste(iue –2 . . . . with but . carina, without femur 5–12) (Figures hind setae ventral of to face apical depression posterior ventral 20); shallow (Figure broad not ea Fgrs1–6;pscxllbsntdvlpd... present lobes . postcoxal 18); 17, (Figures . developed not lobes postcoxal 13–16); (Figures setae ...... Kn n .V Brown V. B. and Kung G. nGtugn(itr,Poia) tdaEtmlgc 3:257–374. Entomologica Studia Phoridae). (Diptera, Gattungen ¨n hr cleghorni Phora Atnasdrid-utaice ein(itr,Poia) tdaEtmlgc 4:1–112. Entomologica Studia Phoridae). (Diptera, Region indo-australischen der aus -Arten gleudungeflu und ¨gelte .s..Ida uemNts111 Ntseen). (Not 1:191. Notes Museum Indian sp.]. n. Dohrniphora g u ennsdrnorpshnPoie,nbtBshebn einiger Beschreibung nebst Phoriden, neotropischen der Kenntnis zur ¨ge glePoie u e etoice ein es ecriugvon Beschreibung nebst Region, neotropischen der aus Phoriden ¨gelte ...... sspotdb S rn E 562 oB. to 0516420 DEB grant NSF by supported is ...... 3 ...... sexspinosa D. .divaricata D. .seriata D. .dispar D. .cornuta D. ¨damerikanischen e species new species new (Enderlein) (Aldrich) (Bigot) Downloaded By: [University of Southern California] At: 21:52 24 January 2008 View publication stats a ue C 93 h epeo rce xeiino h aionaAaeyo cecs 92 o 7, No. 1932, Sciences, of Academy California the of Expedition Crocker Templeton The 1933. MC. Duzee Van Contribucio 1916. Du F. C. Berlin: 43–48, Figueroa Phoriden. 35–36, der Silva Revision 15:19–24, 1929. Maandblad H. Natuurhistorisch Schmitz Phoridentypen. an Untersuchungen 1920– 1926. Limburg in H. Genootschap Schmitz Natuurhistorisch het van Jaarboek Phoriden. an Typenstudien 1923. H. Schmitz cmt .11.DemreohlnPoie e amn’ce amug olgsh Jahrbu Zoologische Sammlung. de Wasmann’schen Academia Real der la Museum. Phoriden de National Memorias myrmecophilen States Canarias. Die Islas United las 1914. the de H. in Phoridos los Phoridae Schmitz de family Monografia dipterous 1921. the E. Abreu of Santos The 1912. JR. Malloch ugG rw V 05 e pce of species New 2005. BV. in Brown Gesellschaft G, Zoologisch-Botanischen der Kung Verhandlungen XI. Miscellen. 6(1993):417–436. Zoologische Entomologia 1867. di von. Italiano su G Giornale Phoridenfauna Frauenfeld Trinidad. Die of 1912. (Diptera) G. Phoridae Enderlein Cave 1995. RHL. Disney oihpddeadPoia.Poednso h aionaAaeyo cecs21:65–74. Sciences of Academy California the of Proceedings Phoridae. and Dolichopodidae 9:5–17. Santiago Nacional 55–57. 23:49–59. 37:509–566. Systematik inisyAtsd acln 17:1–90. Barcelona de Artes y Ciencias 43:411–529. Museum National the of Proceedings 98:55–62. America of Society in17:425–502. Wien na ooiinod afmlaPoia nCie Boletı Chile. en Phoridae familia la de conocimiento al ´n dBaiin.SetnrEtmlgsh etn 73:16–45. Zeitung Entomologische Stettiner ¨d-Brasiliens. Dohrniphora mlr 1 p. 211 ¨mmler. eae to related aiba pce of species Caribbean .longirostrata D. naso h Entomological the of Annals . orihr 1945 Dohrniphora ndlMuseo del ´n ¨cher