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First record of Hyla parviceps on the lower step of a central Venezuelan table mountain

Andreas Schlüter1, Karin Mägdefrau2 1 StaatlichesMuseum für Naturkunde, SchlossRosenstein, Rosenstein 1, D-7000Stuttgart 1, FR Germany 2 ZoologischeStaatssamlung München, Münchhausenstr.21, D-8000 München 60, FR Germany

The Venezuelan sandstone table mountains () are remnants of an extensive sandstone , the Plateau, which in the Tertiary covered a big part of the Guianan Region (Haffer, 1974). The Guianan Region, bordered by Rio Orinoco, Cassiquiare Canal, and Rio Negro in the west, by the Rio Amazonas in the south, and by the Atlantic Ocean in the north and east (Hoogmoed, 1979), is geologically known as the (Gansser, 1954, Fittkau, 1974). Over 160 anuran species occur in the Guianan Region (Hoogmoed, 1979); distribution records are valuable to improve our fragmentary zoogeographical knowledge on that area. Previous distribution records for Hyla parviceps Boulenger, 1882 suggest it is con- fined to the upper part of the and is absent from the Guiana Shield (comp. Rivero, 1961, 1971, Hoogmoed, 1979). Boulenger (1882) described H. par- 218

Fig. 1. Distributionof Hyla parviceps(Guiana Shieldshaded). Arrow indicateslocality Guaiquinima-. aiceps from Sarayacu, Provincia Pastaza, Amazonian Ecuador. Melin (1941) reported it from Taracui, Amazonas, , Cochran and Goin (1970) from Rio Apoporis, Amazonas, and Ceildn, Caqueti, both in southern Colombia, and from Acre, Brazil. Duellman and Crump (1974) and Duellman (1978) reported it from different sites in Amazonian Ecuador, and Toft and Duellman (1979) from different localities in Amazonian Peru. McDiarmid and Paolillo (1988) provided the first evidence for its presence in - from the Cerro de la Neblina area, the southernmost rem- nant of the Roraima Formation. During a recent expedition to the Guaiquinima-Tepui, we collected a female Hyla parviceps (17 February 1990). Situated along the eastern side of the Paragua River, Estado Bolivar (fig. 1), the Guaiquinima-Tepui is one of the largest Venezuelan sand- stone table mountains (approximately 1600 kml including talus slopes). Notable features include an abundant vegetational cover with a high percentage of tall forest, a fairly level topography and a deficiency of rock outcrops (Steyermark and Dunster- ville, 1980). The specimen was caught in a tall forest of the southern lower step of the tepui, ca. 100 km SW of Canaima (approx. 5°37'N,63°34'W) at an altitude of approximately