%$ &9 J?9.(0$8SM     *!-N/-MI Cytospora R$? &2WO--J/J?5.8c- New records of form-genus Cytospora in Iran (II)

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 & J "/NV8 ( O-.Vb/!!&1&8T1-)-.'  M* *,7) Cytospora R$? &2W-.' &#*&2W .VGN1.7MJV#).!.'.0.GN19N < N0 )N < !.8GMI-)O)Nb#.W%J$$aI.@/- -* " I)JVC M Biggs 1989, Christensen 1940 J$8VT'.V0#INVS.7NSMI&VT `V7!.V'.0 6VS-%J/INUM-P8H !.'.0-)M-P/M. 01 -*!-*$&1R$? &2W"/-MI&#* &2W ) N0b$0VVS M Saccardo 1882-1931, Grove 1935, Gutner 1935, Gvritishvili 1982  !.V1P0 &#* O-V01 Farr et al. 1989 !-M.b ') M. ) Sinclair et al . 1987 !-M.b '  &VVV#*)I.VVV($7 M JVVV#- %I*VVV #6VVVSN(-M.VVV' YM.VVVO-%)NVVV"VVV/-&VVV1Q.VVVTC.'.VVV0  C. sacchari E.J. Butler ) Triticum aestivum L. -)M Cytospora tritici Punith.  6VVVVS-%JVVVVWUM-PVVVV&VVVVT `VVVV7!.VVVV'.0O- Saccharum officinarum L. -)M M.0VT1b/-NVW.VGN1MI.V' YM."/-O-.W.#-I.\8-5M.TG M Adams et al. 2005, 2006   -.VV' R$VV?-.VV' M* .VV#! Cytospora R$VV? &2VVW-.VV' &VV#* &2VVW M JVVW.1 .VV I.VV/O 9 Leucostoma (Nitschke) Höhn. ) Valseutypella Höhn. 9 Valsella Fuckel 9 Fr. "V/-.V1 M J$VW.1 .V  &G.VW) &8VS-M9 Valsaceace %N07&18 &8G.$VW.M* *,7 Cytospora R$? &2WO-%JWUM-P-.' &#*&2WO--M.0T1--N19I*?)  M Adams et al. 2005 6S-%JX# .V' !!&V1&8VT1-)-.V'  M*V  *,7) Cytospora R$V? &2VW-.V' &V#* &2VW-J$1 %IM  ) von Höhnel 1919, 1923, 1927, 1928  V#*'!*V M 6S-%I*1-I./O5-N00 7U*H8SI !*V "0V$\ ' M J#I*1.' YM.O-%)N"/-MIJ/J?0'. M-L &/.S Nannfeldt 1932  6#.# MI%I M.VV1-JVV#--.VV' %NVV@CVVV/-M!&VVaJVVW&VV?*8    9  9  9   VV#*' ) Gvritishvili 1982  ./*VX08/M* 9-NV0G- M J$8VT'5).V8 R$V? &2VW"/--.' &#* &2W R$V? &2VWMIR$V?O-NV7 "0/.VSA*@S-J$1 %IM6(? Spielman 1983, 1985  " 0TS- J#-M %I* #%I.8S-9 #*'!* bS*7%JW./.S.$W8H -.' R$?O-9 Cytospora &8G.$VW-.V' &#* &2WO-. .?0[*7 Gutner 1935  N$7* ) Grove 1935  ))N *.VI "0V$\ ' M JV#- %J#.SMQ.[&1-M&8T1-)-.' R$? &2W) Cytospora R$? &2WO-%JW ) Urban 1957, 1958  !.V1M- 9 Kern 1955, 1957, 1961  !NVa 9 Défago 1935, 1944  -.' .^/)O-R$? &2W"/-MI.' &#* & 2W`0b7M*$ &1 Hubbes 1960a, 1960b  R1*' 0VV[*7" VV_    ./*VVX08/M* M JVV#- %INVVa%I.8VVS-.b/]*VV*/P0).VVS.$W 6VVH/M -M&V#* &2W  91.S-)M*WN0'. ?I.D7-MI Cytospora R$? &2WO-I*?* -.' &#*&2W .VS.$W 6VH/M-. ' .^/)0[*7" _   !-M.b ') P -I! M 6N/LSR$? &2W"/-MI .1.V/OM-IM*V .*b* -.' U)M` a&1-M.' !!61-N9Q*8T0.a--)MI*?* -.' &#* &2W   !-N/-MI Cytospora R$? &2WO--J/J?5.8c-

J#-M %I-IM-N &V1a*V1N !-NV/-MI Cytospora R$V? &2VW-.V' &#*&2W%M.1MI&.@ "/N7 . /J Silene boryi Boiss. -)M-M C. silenes Gonz. Frag. &#*&a6S- Fragoso 1918  )O*-N 9 Petrak & Esfandiari 1941  -M./J$VS- )_-NV8S NV0#9"/N/.VSRTVS M I*V #UM-PV )/NVW ) Steyeart 1953  5M.08VS- 9 Petrak 1949  _-NV8S 9  )   -M./J$S- UM-PV!-NV/-V8H !.V'.0-)M-MR$V?&2W"/-O--N/I-.' &#*&2W   I.WM- -.V' &V#* &2WO-.GN1./-P/M. 01) . *#*Ta.7    )    I)M.@C )!.bW- M J#- %I* # 9   !.bVVW- M JVV#- %I*VV #&VV.@ !-NVV/-%*VV0 !.VV8GMI-)M-M Cytospora R$VV? &2VVW MINVa.N  Salix zygostemon Boiss. -)M-P/M. 01 .!-*$&1-M C. translucens Sacc. V .!-*V$&V1 S. officinarum -)MO--M C. sacchari PV0#   !-M .Vb ') .#.GN'.c . VW.' .V$1 )-JV C- M JV#- %I* #UM-P!.8SO*G!.8S-MINbX0# 8 .J0S*S-M. 01 !.V8GMI-)OV .9 C. juglandicola ) C. juglandina -.' &#* &2W.SMN1" _    9 C. ambiens  .VVWR$VV? &2VVW"VV/-O--NVV/I- .VV' &VV#* &2VVW9!-NVV/-/*VV$?MI)INVV 9M.$[9J$900S!.8GMI-)MO-007N7&1-M C. cincta ) C. atra 9 C. platani 9 C. ocellata  -.VV' &VV#* &2VVW   !-M.VVb ') NVV .C*VV8 -NVV0G- M JVV#- %I*VV #-O.VVS-J?*VV')5*VV7  -M C. rosarum ) C. nivea 9 C. gutnerae 9 C. carbonacea 9 Cytospora atrocirrhata J#-M %INa UM-P!-N/- - YM.M*--N1-J/J?-.`-.' !-*$&1 -.V' &V#* &2VW .VS.$W 6VH/M -.' .^/)0[*7)./.S.$W M*$ &1 N_.C0D7 M 6N.@#-!-N/-MIR$? &2W"/-O-I*?*    .SMN1U)M -.V' .V^/).VSMN1 9.V' &V/-J? -O.S Z.G)-O.S-J? 9.[M.- &#*.' #-M)!  ? !-M.Vb ') NV .C*V8 U)MV2c.V' &V/-J? .VS.$W 6H/M - .^/).SMN1.' ) YM.&$NS J#-M %JWZHX -M)!  ?-.' %M. W.1.' &/-J? M 6N.@#-      :D1)&@08# R$V? &2VW-.V' &V#* &2VW.b0 *#*VTa.7&V.@ &V1a*1N >/.8#9N_.C&8W*#MI -.V`-&V#* &2VWM.([9"01"/-O- M IIN . &/-M-9&#* &2W6' .W!-N/-MI Cytospora -.V' !.V1P0 PV0#N/I&#* &2W)I--N1)J#*W . /*TD !-N/--.' YM.M*--N1-J/J? 0 J$W.1 . /KANW&1.' &/-M!"/- M J#IN . .N .'.0J/J? !-M. ')N .C*8-IN1 0G 

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                  M `T/I M c9N1 c.' Mb9.' c.' Ma0   &/-J? Cytospora intermedia ƒ bW Fig. 1. Cytospora intermedia : a. Conidia, b. Conidiophore, c. Disc.   )  9 -.V'. IMI9   &V/-J? 08H -.'. IMI YM.&$NS JWM-.' .^/) I-NV .8#.S&?MI  -. IMI)INaJWMN8 .0  2 )  29 007N7&1I-N .8#.S&?MI N/.VSO-N8VX01I-N .8# .S&?MI  -. IMIYM.&$NSJWM9>/.8#&1&?*7.1 M I*1JWMJ. M I*1V/. O!IM* -.'. I  %.V0"0V$\ ' M JVW.1 .V !-NV/--.V' YM.VM*V--NV1-JV/J?*V`&#* &2W"/- M IIN . UM-P.0#IMI&#* &2W"/---N1-J/J?!.1P0  Quercus brantii  !-M. ')N .C*8-IN1 0G

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              9`VVT/I M c9NVV1 c.VV' Mb9.VV' c.VV' Ma0  ƒ &VV/-J? Cytospora kantschavelii ƒ bVVW %N@CV/-M!.'M M d Fig. 2. Cytospora kantschavelii : a. Conidia, b. Conidiophore, c. Disc, d. Locules arrangement.    Cytospora leucosperma (Pers.) Fr., Syst. mycol. 2(2): 543, 1823 :    N8 *0a9!.8SN!.8S-9 Platanus orientalis L. -)M 9  &/-J? 0 %JW.SMN1-.' &#* # 9  &V/-J? ƒNV .C*V8-M)! V ?9  2 2 9N8    .7M-9I.1! NG&1IN?)N1%I.? .7M-9!-*/N &1=J$$S%I.?  N8 *0a9!.8SINa!.8S-9 Amygdalus communis L. -)M 9 Robinia pseudoacacia L. -)M 9  &/-J? ƒ .C*8-M)!N  ?9  2 2 9N8    -M)! V ?9  2 2 9NV8    .7M-9!-N(7&1!-J '%I.?  N8 *0a9!-J '!.8S- &1%)N%I.?  N8 *0a9!.8SINa!.8S-9 Juglans regia L. -)M 9   ƒ &/-J? ƒ .C*8N .C*8-M)!NM  ?9  2 2 9N8   .7M-9!-J ' -.'. )N8S-N8 .0  ƒ  N@&1./.' &G.W-)M 0 !.1P0 6.1- )MYM.-.' .^/) B@SMI.'. )N8S- M J#*W . 0bX7!.1P0 6.1!)MI 8@ . a./)%J$a-NSM*c&1YM. %NV07-N8VTa.G.V7-N8VTa.GV#M&V1&8T?N15M*[&1)%INaJWM/*G!.1P0 6S*S MI)!.V1P0 B@VSMI)%JW%J0W*S!.1P0 6S* SbS*78 .a.'. )N8S- M J$W.1 . ZHX )!.V1P0 6V.1B@VSMI. )N8S-N@ M J#*W . %J/I#M%.0Sa.#5M*[&1*08S-D !-M. ')N .C*8-IN1 0G

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               VV/-M! M d9`VT/I M c9NV1 c.' Mb9.' c.' Ma0  &/-J? Cytospora leucosperma ƒ bW %N@C.'M Fig. 3. Cytospora leucosperma 0 a. Conidia, b. Conidiophore, c. Disc, d. Locules arrangement.   &V?MI  )  -.V'. IMI)INVaJVWMNV8 .V0  2 ) 0V07N7&V1I-NV .8#.S&?MI &V?MI  -.V IMI.V' &V/-J?&V0a&V$NSJVWM9>/.V8#Q.VS-N1 M I*1JWMJ.I-N .8#.S M I*1V/. O!IM* -.'. IN/.SO-N8X01I-N .8#.S !!--NV1-M-NV7 0S)&#**( 9 &#* &2W"/-0[*7" _    ./*X08/M* -.V' &V#*O--M.0VT19.VS.$W6VH/M-.' .^/)Q.S-N1&a-M*c&1 M 6S-&8NN#MI .#.V1P0 &V$ -I 9-)0[*72c M 6S-%I-IM-N&#* &2W"/-  synonyme  I-N8 -M`/IP# )NV < !.V8GMIV8H -.V' R$V?O--IJV8 -.' &#*)%I*10S)M.0T1&#* &2W"/- M JW.1 . !-N/--.' YM.M*--N1-J/J?*`N_.C&# &2W* M IIN .  .W-MN < N0    Cytospora ribis Ehrenb., Sylv. mycol. berol. : 28, 1818 : 9-PVVaN !.8VVS-9 Elaeagnus angustifolia L. -)M 9  &VV/-J? 0 %JVW.VVSMN1-.VV' &V#* # !-M. ')N .C*8-IN1 0G

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               %N@CV/-M!.'M M d9`T/I M c9N1 c.' Mb9.' c.' Ma0  &/-J? Cytospora ribis ƒ bW Fig. 4. Cytospora ribis : a. Conidia, b. Conidiophore, c. Disc, d. Locules arrangement.  "VW)M-N8Ta.G&1/. 8 -- *(#M&1%  &/-J?&$NS 0YM.  &$NS- .' .^/) %NV07%.0VS ƒ-N8VTa.G&1/. 8  P2S#M&1-N8S`8X76XSMI)%I*1 greyish sepia  NS-M-N8S`8X7 .aM*c&1)&8W-IW6.1YM.&$NS M I*W . %J/I greenish black  b0VD B@VSMI%JV$a-NSM*Vc&V1)M*)&1)%I*1%.0S#M &1YM.%IM.1-.' -J#- M J$a .  )NV@ M J#*VW .V %JV/I-- &#-*8VS-.V7-)Na.bW-&1%IM.1-.' -J#- M J#*W . 0bX76Xa )%I*V1N0 8 M.0T1%IM.1-.' -J#-%M-*/I6 .H_ M 6S-N8 .0  ƒ 2 %IM.1-.' -J#-.7M- M J#INb#J0*7c.' ./-L b0D MI%IM.1-.' -J M#- J$8T'IN$ %N@C`/J?-) )  9 -.V'. IMI9  ƒ &V/-J? 0V8H -.'. IMI YM.&$NS JWM- .' .^/) I-NV .8#.S&?MI  -. IMI)INaJWMN8 .0  )  9 007N7&1I-N .8#.S&?MI  ) 2 92  0V07N7&V1I-NV .8#. S&?MI  )  9 -.'. IMI9  &/-J? MI*1JWMJ.  9  ƒ &V/-J? MI*V1JVWMJV.I-NV .8#.VS&V?MI  -.V IMI)INVaJVWMN8 .0  2 )INVaJVWMN8 .0  2 ) 9 2  2 007N7&1I-N .8#.S&?MI  )  9 -.'. IMI !-M. ')N .C*8-IN1 0G 

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NEW RECORDS OF FORM-GENUS CYTOSPORA IN IRAN (II)

* KH.B. FOTOUHIFAR , GH.A. HEDJAROUDE, D. ERSHAD, S.M. MOUSSAVI, S.M. OKHOVVAT and M. JAVAN-NIKKHAH

University College of Agriculture & Natural Resources, University of Tehran and Iranian Research Institute of Plant Protection

Received: 25.08.2007 Accepted: 16.06.2008

In a taxonomic study on form-species of form-genus Cytospora , about 400 fungal specimens were collected from all over the Iran during 2003-2005. In this paper, the second part of the obtained results, including seven form-species are presented. Cytospora intermedia , C. kantschavelii , C. leucosperma and C. ribis are reported as new members for mycoflora of Iran. Salix aegyptiaca for C. aurora and Pistacia khinjuk for C. terebinthi are reported as new hosts (matrix nova) in the world. Also, previously reported form-species C. sacchari is redescribed in detail.

Key words: Fungi, , Biodiversity, Coelomycetes

To observe the figures, refer to the Persian text.

* Corresponding author (E-mail: [email protected]) NEW RECORDS OF FORM-GENUS CYTOSPORA IN IRAN (II) 39

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Addresses of the authors: Dr. KH.B. FOTOUHIFAR, Dr. GH.A. HEDJAROUDE, Dr. S.M. OKHOVVAT and Dr. M. JAVAN-NIKKHAH, Department of Plant Protection, College of Horticulture Science and Plant Protection, University College of Agriculture and Natural Resources, University of Tehran, Karaj 31587-11167, Iran, and Dr. D. ERSHAD and S.M. MOUSSAVI, Department of Botany, Iranian Research Institute of Plant Protection, P.O. Box 1454, Tehran 19395, Iran.