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uttnz 7leans, Sqpict Soc. Jdpan 49 (1): 73-84, February l998

A contribution to the knowledge of the larger fauna (, Macrolepidoptera) of Mt Litoyka (Primorye Territory, )'

Yu. A. [I]smgrJAKovD, Keitaro EDA2) and E. A. BEuAEv])

i)Institute of Biology and Pedology, Far Eastern Branch of Russian Academy of Scineces,

Vladivostok, 696022 Russia 2]4-3-24-303, Kita-ando, Shizuoka, 420-088I Japan

Abstract Three hundred and forty-three species of larger from 15 farnilies (Cossidae, Limacodidae, , t}eometridae, Thyatiridae, Lasiocampidae, Sphingidae, Saturniidae,

Brahrnaeidae, BombyCidae, Notodontidae, Lymantriidae, Nolidae, , Arodidae) are recorded foT the first, time from Mt Litovka (Southern Sikhotae-Alin' Range, Primorye Terri- tory), of which 2g5 species are indicated for mixed broad-leaved-coniferous fbrest, 71 species for

spruce-forest and 101 species for the subalpine zone, As the result of observations, more than 160 species of the lower fioristic belt are found to be penetrating up to mountain taiga forest or even into the subalpine zone, among which 44 species turned out to be new fbr the high altitude zone of the Sikhotae-Alin' Range. Among the Geometridae 3 species (Scopula asthena, J?Zinusia laria, H:Iidnefia adlasma) are recorded as new fbT Russia and 4 species (Idaea avensata, Scopula tenuisocius Perizema taeniaturn, Eulithis prunata) us new for Primorye Territory; among the Noctuidae 3 species (1)-ichosea ludlrfaa, Poita hepatica, Apamea sodads) are also recorded here ft)r the first time for this region,

Key words Moth fauna, Macrolepidoptera, Primorye Territory, Russia, new record, mixed

forest, spruce forest, subalpine zone, biotope.

Introduction

Mt Litovka (fbrmerly-Khualaza, befbre it was renamed in 1967) is the socond highest summit (1,279 m a. s. 1.) of the Livadiiskii Range-a south-eastern spur ofthe Sikhotae-Alin' Range. It is situated about 12 km southward from Anisimovka (better known from the old entomological 1iterature, under its previous name Kangauz)-a small railway station only l20 km from Vladivostok and, due to this position, it is one ofthe most accessible surrmiits in the Southern Primorye with well marked zonation, typical fbr this region-from mixed coniferous-broad-leaved fbrest at the foot up to the subalpine zone with arctic vegetation and stony deposits at the top.

The first entomologist to visit this mountain (first in 1923 and then in 1928) was the young naturatist A. I. Kurenzov (later to become the famous explorer of the entomofauna of the and a professor), who collected here some remarkable species of including moths, some of which turned out to be new to science-Eunyydoxa advena Fil. (Tortricidae) and Areqgnatha sichotensis Kurenz. (Noctuidae). The results ofthese observa- tions (obtained mainly during the second ascent, because the first one took place in early spring when the top of this summit was still covered with snow) he published in two scientific papers (Kurenzov, 1929, 1937) and he later described both excursions in his memoirs based on his field note-books (Kurenzov, 1953, 1961, 1973).

'Report No. 54 frorn Russian/Japan Cooperative East Asian Entomological Program.

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74 Yu.A. TsHrsTJAKov. Keitaro EDA and E. A. BEuAEv

Since that time many naturalists and professional entomologists, including the first author of this paper, visited this meuntain. But these were mostly one-day excursions, and the pardcipants had no equipment for night{ollecting, so even those overnighting at the top could collect almost nothing and all collections of moths were taken in the closest vicinity

ofAnisimovka only, For this reason the moth fauna ofthe Litovka Mt. still remains almost unknown, except for a little data scattered throughout the literature, mentioned above and some other papers (Filipjev, 1927 ; Tshisljakov, 1984),

In July, 1995 a joint Russian-Japanese expedition consisting of 7 entemologists had the opportunity to carry out a three-day (from 19th to 21st efJuly) field survey close to the foot of this mountain. Using this opportunity we decided to organize a special excursion for

night trapping in the high altitude zone of the summit. During this excursion 2 light-traps were installed : one of them, with mercury vapor lamp, was operated from a portable "Honda generator EX 3oo" in the subalpine zone at an altitude of 1,240 m above sea level, and the other, with a super-Eictinic fiuorescent light tube 8 W, was operated from a 12 Volts DC power source in the spruce fbrest, at an altitude of 1,150 m above sea level. Thus, using the transect method of collecting, we obtained some interesting data about the moth population within three vertical belts of vegetation. The results of these observations with brief accounts of the cotlecting sites are given herein.

Collecting sites

Biotope 1. 8km SSE Anisimovka, the upper Beresovyi stream, 400m a, s. 1, Small meadow by the foot of the northern slope in the mixed coniferous-broad-leaved fbrest (MF, below). The clearing stretches along the stream, the banks of which are occupied by a narrow belt of riparian forest with old trees of Populas maxinzowiczz7, UZmus propinqua, en(nsenia anbutij?)1la, Abius hinsuta and Syringa amurensts. The opposite side of the rneadow is bordered by dense secondary forest on the site of an old fe11ing, consisting of various broad-leaved trees : PopultLs dovicliana, IleaxiniAs manctshurica, UZmus laciniata,

Betula mandshurica, B. c(rstata, Acer manclychurieum, A. mona Maackia amurensis, Mains mandshuiiica, lvglans manshutiba and others, The underwood is composed of various shrubs (Lonicera maackiZ L. maximowizeii Pltilacleiphtts tenuijbfius Rthes inaximovic- a'anum, Eieutheroeoceus sentitrastLs, Acanthopanex sessilijlorum, Clo7:ylths manshurica, Euonymus paucijlora, EL sacrasaneta), lianas (Vitis amurensis, Schisandia chinensis, Actinidia colomicta, A. arguta) and by young growth of coniferous trees. The end of the meadow adjoins the northern slope of the mountain, which is covered, like other nearby slopes, mainly by euercus mongoiita 7-Zeia amurensts T manctshurica and coniferous plants: Pinus koraiensis, Abies hoiojphyila with an admixture of Pibea yesoensts, Abies nqphroiepis and isolated trees of TZixus cuspidata.

Biotope 2. I2km S Anisimovka, fern-moss spruce fbrest on the northern slope of the rnountain, at an eleyation of 1,150 m abeve sea leyel (SF, below). Predominant species are ceniferous-Pieea cijanensis, PL jesoensis and Abies nephroftzpts with isolated trees of Pinus koraiensis. Deciduous trees are represented mostly by Betula caytata and B. ftznata, with isolated trees of 7}iia taeuetii, Ubnus laciniaa, Acer tqgmentasum and A. ukutunduense. Underwood is very sparse, with scattered shrubs of Lonicera gthbijlbra, Sambueus sthiri a, Rasa acieulaiiy, Rthes manschurieum and Ethinopanax elatus. Grass cover is sparse also, composed mainly of ferns and mosses (Dryopteris amurensis, D. bushiana, Athyrium ntbnl' ]es, A. pyenasontm, Hylocomium splendens,, Rlij,tidiadeiphus biguetrtLs, Pleumampsis ntthenica), and some grass as well (Canex xiphium, C]bcads acetoseha, uraZctstein'ia teniata, CZintonia

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Larger MothFauna of Mt Litovka, the Primorye Territory 75

udensis),

Biotope 3. About 12 km S Anisimovka, stony deposit in the subalpine zone (SZ, below) on the southern slope near the eastern summit ofthe mountain, at an e]evation ef l,240 m above sea level. The stony deposit is occupied partly by a tapestry ofthe prostrate shrub Microbiota ciecussata and some dwarf shrubs (Ledum decumbens, Rhododend)nyon aureum, dassiope ericoides, Vaccinium uliginayuni, Arctous aipina) or by isolated shrubs of Rasa acieulaiis and sparse plots of grassy yegetation, including mostly Artemtsia lagocephala. This open area stretches far down almost up to the fbot, and is surrounded on both sides by orophyllic scrub

fbrest consisting of scattered Ribea cu'anensis, Betula kinata and various shrubs, such as Duschekia fruticosa, SYringa woijZi, SPiraea betuiZfoila and Lonicera eduds. The clearings cons{st of subalpine meadows composed of Ctilamagtzrstig langsdoizt77i, P'lerntXtzm aipestre, Hieracium coreanum, ([Ienanium erianthum and other tall grasses. Some plots at the boundary of the stony deposit with spruce fbrest are covered by a tapestry ofBeringia pacifica with other dwarf shrubs,

Species composition and zonal distribution of the larger moths

The data about species composition and zenal distribution of the larger moths were ebtained

mainly during three nights of trapping, namely : 19, 21.VII-in the MF, 20,VII-simultane-

eusly in FS and in SZ. In addition some material, taken 10 days later-31.VII and ]. VIII in the same site by the foot of the mountain (MF) are included as well (this additional "+;' material is marked with in the list). Species recorded fbr the first time for the high

altitude zone ofthe Sikhotae-Alin' Range are indicated in the list with asterisk,

No, specimens No. specimens collected in collected in List of species List of species each biotope each biotope MF SF SZ MF SF SZ - - COSSIDAE T consimids 3+1

Lamedoccrssus terebtus' 2 1 T athicestata 2-- - - LIMACODIDAE Parapsestts argenteopicta +1

Parasa sinica 11 GEOMETRIDAE

Cbvatenema christophi (ORTHOSTIXINAE) DREPANIDAE 7Vbxa seriaiia )

Agnidea seabiosa .fixeni 1+1+1+3+2 (GEOMETRINAE) Rseudothara parvula Geometra papifiOnaria 1 Norctstromia gntyearia' 4 12 G, sponsaria +1+1

Sabra harpagula G, ussuriensis otivacea G, dieckmanni 1 curvatula aeuta 2+3+1 1 G, valida 11+1 adu .t211ojevi G. glaucaria' 2 THYATIRIDAE Cbmthaena amoenaria" 2 10 Hdbrosy,ne intermedia +2+17+3 C tancrei 2 ft dieckmanni C ingrata* 1 2 Tlethea oculaits amutL Hemtthea aestivaria t3+1 ensisZ 7Vipponegelasma lucia or terresa 5 1

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76 Yu ． A ． TsHIsTJAKov， Keitaro EDA and E ． A ． BELJAEv

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Largcr Moth Fauna of Mt Litovka， the PrimoTye Territory 77

Bupalus ye5taliS＊ 10 Odonestts pruni rafescens ， 5 十 610 一 Antic）petla di：t7itsaria ＋ 2 1）en 〃 olimtas seroehans 十 6 2 sibiricus A 尸た ノ〜anna 〃le1dna 厂ia 17 33461 2

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78 Yu. A. TsHisTJAKov, KeitaroEDA and E. A. BEuAEv

IVerice leechi 13+1 MdeochTista miniata 2+2

Leucodonta bicoloria 1 M aberrans askoidensis +2

Phalera bucephala 15+2 M rasaria +1+1+1

Shaka atrovittatus id rosaeea

Pterostoma gniseum 1 l 1 M paVida

R sinicum 1 Midina artaxidia 1

Fletsaptet:yx ladislai 1 Mlelanaema venata 2

Tbgupteotr vehttina 1 Macrobrochis staudingen' +3+1 Lophontosia cuculas +14+35+5+3 Pelosia muscerda tetra- sticta .Hlexqfrenum leucodera' 14 1 Lith(rsia 15+273+13 1 Semidbnta biloba quadea' Ghoria gigantea 7 EPodonta aneata G, couttoidles* Pttibdbn hoqgei 1 Edema vetusta 7 SPatatia dives 1+2 grtseoinm E deplanum pavescems 58 S. pinsiotis E crthratum 6+152+2 4 S. doerriesi 1 PericaMa matronula Micromelak4)ha trqg= 1 , lodyta amurensas RhMparioidea amurensis 1+2 Gimphisia crenata japon- 9+6 2 iea R. nebulbsus 2

C7bstera athesigma cuF 5+5 SPdosoma urticae mancZei 1･+13+2 tuloidles Chionarctia nivea a anaehoreta +10 SPilarctia seriatopunc- LYMANTRIIDAE tata

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A. -nignim ussuricum coenobita 2

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Larger Moth Fauna of Mt Litovka, the Primorye Territory 79

- - C junkowskii 11 Anthagrata 2 - - C athonigra dnmia restituta' 1

(HELIOTHINAE) C spurcopyga 32 - llyrrhia umbra 1 Ccamptastigma () Dimorphicasmia vari- 2 egataEnargia Aetebia praecurrens 11]2 paleaeea 11 Protexarnts balanistts'

contusa' 51 1 sigma 3 lpimorpha Chasminotles athonitens Sineugrapha bipartita ' C sugii 11 Paraiarsia punicea 12 Chytonix athonotata 1111 Diarsia brunnea 2433 618542 Eucarta arctidles l\kestia sineera Ilrometopus JC dbscripta" 8 .t7tivicol2Zs

sigillata XZ ditraperium 5PhrcLgCt?ira (SARROTHRIPINAE) Anoplectoidbs prasina Ab,cteola asiatieaS 1 A. virens 2 (HADENINAE) (CHLOEPHORINAE) Parhylophila celsiana 71 Poha nebuttrsa L 21 13323 Rseudoips prasinantts' 2 R gohath

A hepatica (ACONTIINAE) 7}Jisatefies emertuaiis' 1 Melanchva persicariae 1 Kqyaga numisma 1 Lacanobia contigua Anterastria auata' Il L. contrastata 1 Bryophilina mollicula* Lasionyeta hospita' 6

Hadena deathata 1 () Lamprotes mikadina 1 ?Llythimna divergens' 22 AutQgrapha urupina' 11 Aletia radiata' (CUCULLINAE) A. buraetica' Diachrysia shrysiti 1 CtltXierges ramosuin* 1 (AMPHIPYRINAE) (CATOCALINAE) mancischurica 11 Apamea soduds* 11111 QJsgonia Mbcis ancltki A. striata

A. aquiZd (OPHIDERINAE) [tygephila pastinum 1411111 A, remistsa hercules A. sordlens SJ;pnoidles Pangrapta lunulata A. scokrpaeina 11 Hbpatica anccprs Leucapamea askoidZs

Diomea Parastiehtis suspecta' 1196 J'ankowskii

Laspeyria 2 (]elaena leucostigma .fexula* Paragabara secunda Emplarcia lvcipara 1311l1

PhtOgophora iattstrata (HYPENINAE)

1[lypena tristads 1 11 T)"achea atripficis

Bomolocha bipartita* Z punkthonis B. ztila(HERMINIINAE) 1 Athetis athisignata

Amphipyra schnenekii 1 1

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Larger Moth Fauna of Mt Litovka, the Primorye Territory gl

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Discussion

In spite of the short period of research, we obtained very interesting results, including some remarkable faunistic data. For example, from 132 collected species of the Geometridae, 3 (Sbopula asthena, Venusia laria, H]ycbeha aciesma) turned out to be new fbr Russia, and 4 species (Idqea aversata, Sbopula tenutsocitLs, Pentoma taeniatum, Eufithis pntnata) are new fbr Primorye Tenitory ; among 98 collected species of the Noctuidae, 3 species (7}ichasea iitthMa, Poha hepatica, Apamea sodods) are absent from the current check-list of the Noctuidae ofthe Primorye Tenitory by Kononenko (1990) and are recorded here fbr the first time for this region also.

However, the list of species presented above is far from complete even for that season. The real number of species, representing a middle summer phenological aspect in the local fauna, must be much higher. For this reason the obtained correlation of species composition as 285:71:101 in the discussed biotopes must be sen as a provisional one and refiects only the results of this short observation. If we had more complete data, this correlation would be changed substantially. Moreover, about 60 species, mentioned in the list from MF only are known (Kurenzov, 1937; Tshisljakov, 1975a, b; Vasyurin, 1978; Vasyurin & Tshisejakov, 1979 ; Kononenko, 1986) to be more or less constantly occuring (Agnitbra scabiasa .fixeni, Dmpana curvatula acuta, Hatvvsyne dZeckmanni, 7lethea oculatis amurensts, Venusia blomeri, Eutithis conveilgenata, Xdenthonhoe fiectantemediana, Lenicrgrapha temerata, dabera exanthemata, Plagodis putveran'a, Rtagodis dolabran'a, iFletreula jureula sangnrka, Stauropus fogi persiMiZis, Arbtodbnta dembowskii, Leucodonta bicoloha, 5?)ataha dtves, Cihstera ana- choreta, Elphrageidus simizas, Edema cmbratum, SPilarctia seriatopunctata, Panthea coenobita and others) or penetrating from time to time to the mountain taiga forests (Cbmthaena tancnei, EPb:rhoe supergressa, DeiZ!:ptenin mancischuriaria, Rhthonasema tendinosaua, CZxbetzi schaE777eri, Endeopiodey indutinatia, Malacasoma neustrium testaceum, CZidombuiix tatarinovi, Ampekzph(rga retbiginasa, Achas artemts, Shaka atrovittatus, Tbgeptet:yx vehitina, Arttoniis athum, Euproctis pipenta, PlambalZia matronula and others). Quite probably all of them will be found in future in the high alitude zone of Mt Litovka also, But even on this limited data it is possible to see that species composition of Macroheterocera in SF and SZ consists of a high number of species, more typical fbr the lower floTistic belt. Thus, the overwhelming majority of al1 discussed species from the Drepanidae, Thyatiridae, Lasiocampidae, Sphingidae, Notodontidae, Lymantriidae, Arctiidae and Geometridae recor- ded here from the high altitude zones actually are connected trophically with broad-leaved

Figs 1-3, Collecting sites ofMt Litovka. 1. Biotope 1 (mixed broad-leavedtoniferous forest). 2. Biotope 2 (spruce-forest). 3. Biotope 3 (top ofMt LttQvka). Figs4-16. Some moths collected at Mt Litovka. 4. Lamedocessus terebrus ([Denis & SchifiermUller]). 5. Rroblepsis phoebean-a Erschoff. ,6. Paralebeda plagijZira femorata Menetrids, 7. 5Phingulus mus Staudinger. 8. Kentrochnpsads streckeri Staudinger. 9, Brahmaea tancrei Austant. 10, Oberthueria caeca (ObemhU'T). 11. J5restia sincera (Herrich-Sch'a'ffer). 12. Poila hettu)ica (Clerck). 13, Lasionycta haspita Bang-Haas, I4. CaZ2ilarges ramasula (Staudinger). 15. Hlepatica anceps Staudinger. 16. Rara- gabala secunda Remm.

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82 Yu. A. TsHisTJAKov, Keitaro EDA and E, A. BELJAEv

trees and shrubs, occuning in the priamurian mixed forests. The greater part of the Acronictinae, Catocalinae and Ophiderinae from our list of the Nectuidae also have close

relations with deciduous vegetation.

The number of species recorded from SF is smaller than from any other biotope. Typical

species in this bietope are as fbllows: Cbmthaena amoenaua, Ubnusia cambrica, Asthena

sachaiinensis, 7}iphasa dubitata, Pentoma saxeum, T7iera vatiata, Slemiothisa signaria,

Tbenmphora unio,' Bupalhas vestads, Dedeptenia nbeata, GaraeiAs mirandus from Geometridae and IYbstia sinceTzz from Noctuidae. Besides, we collected 1 male ofX sincera in SZ. Meanwhile Fibiger (1993) regards JYI sincera as an inhabitant of primeval coniferous fbrest and as a very rare species everywhere. One ef us (Eda), obtained evidence of that a week 1ater, coilecting X sincera in the subalpine zone of Mt Kita-dake (at an altitude of about 2,200 m) in Japan (Kobayashi & Eda, 1995), All the rest indicated for this biotope, are species which inhabit only MF constantly or are common to both these biotopes.

The situation is almost the sarne with species composition of Macroheterocera in SZ,

composed totally by moths from the lower floristic zones, while the typical subalpine species are not fbund here. On the other hand most ofthe Noctuinae and some Apamea (A. aeuim, A. remissa, A. striata) and several Geometridae (Rheumaptera inanata, Angerona prttnaria, thebomorpha falgt{raria) in our case are recorded frorn SZ only. A curious fact is that only a smal1 number of species penetrate from valley to mountain taiga through the dense forest along northern slope (such as IVbrdstromin gtiseana, H:)/pameeis, roboran'a, Eustroma melancholibum, .\enarthoinha propinguata, Euch,ismphia cumuima, Iiletreula bictLspis, StaurL opus fogi persimids, Notodonta dembowskii, Rirerastoma griseum, fibesaptenyyx ladisini, Hexopenum leucodera, Slemidbnta biloba, Gh(m'a gigantea, Edema cleplanum paveseens), while many ofthem (LamedocasstLs terebnLs, Sabra harpagula olb7acea, Dnepana curvatula acuta, 712thea or terrcrsa, Geometra giZTucan'a, Pbrkoma minihiatum, 7)iftrptera bedo, Hkiteranuia dissiniiZZs, Agaraeus parya, Gastsopacha papuiijbEa angzLstioennis, G. clanata, Dendeoiimus superans sibincus, Malacasoma netLstrium tesldceum, Laothoe amurensts, Dedephila eipenor, Brahmaea tancrei, Oherthueria eaeca, derttra enuinea eandidti, Hemij2intonia mandshurica, thopyia meticulodina, feridea glgantea, Glmphisia crenata iaponica, Lithasia quadea, Ghon'a conttoides and many others) have not been fbund in SF, but were observed in SZ near the top of the summit, which they reached through an open

area, along a stony deposit on the southern slope, spreading far to the foot of the mountain.

Acknowledgement

We wish to express our thanks to Dr T. Saigusa Kyushu Univ,, Fukuoka and Dr N, V. Kurzenko, Institute of Biology and Pedology, Vladivostok fbr great help with our prQject. Thanks are also due to Mr M.Tashita, Nagano and IVtr N.Nishio, Ueda fbr their co-

operation in carrying out night collecting at Mt Litovka, and Mr S, Sugi, Tokyo, for his advioe and infbrmation on literature.

References

Fibiger, M., 1993. Noctuinaell. IVbctuidde Europaeae 2. 230pp., 11 pls. Entomologlcal Press, Sdiro, Denmark. Filipjev, N. N., 1927. Zur kenntniss deT Heteroceren (Lepidoptera) von Sutshan (Ussuri Gebiet). drheg, zoot Mua rs : 219-264, pls l2-16, Kobayashi, H. & K. Eda, 1995. Abundance of Xestia sincera (Herrich-Schlifier) (Noctuidae) in the

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Larger Moth Fauna of Mt Litovka， the Primorye Territory 83

− conifer zone of Mt ． Kitadake ． ノ＠ 砌 H 罐 アoc8 厂醜 ▽ ，（186）： 167 169， − Kononenko ， V ． S，，1986． ［Noctuid moths （Lepidoptera， Noctuidae）of the dark coniferous taiga of the − ’ − Sou 由 ern Sikhotae Ahn Range ］． 加 SiStematika i 盈 o わgjα nase たomyh D α 1短 go Vostoka： 44 53． Vladivostok．〔in Russian ）． ，1990． Synonymic Ch k List of the Noctuidae of the Primorye Territory， the Far East of U ， − S．S．R ． Tinea 13 （SuppL D ： 1 40 ． − Kurenzov， A ．1．，1929． ［About zonal distribution of Lepidoptcra in the southern SikhotaeAliゴ ］． 一 − ZapiSki Vladivost∂ k． otdeL Geogr ． ob ッa 19 ： 41 50． Vladivostok ．（In Russian ）， − ’ ，1937． ［New and interesting Lepidoptera from the Sikhotae AHn ］， Vestnik Z）厂 FiZ AiV − ∬ SR （26）： ll5 132 （in Russian）． 厂 ，1953． レ gorakh Tac乃in−Guana ［ln the mountains qブ Tachin−Guan ］．

lg61 VubazhiShch α 配∬ urii reliktov ln the refuges the u ∬ urian relicts 182 ， ， h kih ［ qプ ］、 pp ． ’ Primorskoe knizhnoe isdatelstvo ， Vladivostok．（In Russian ）、

’ ， 1973． M （）ノ puteshestvia ［ル4ン travelS］，623 PP ， 1）al nevostochnoe knizhnoe isdaterstvo， Vladivostok．（In Russian）． − Remm ， H ．，1983． New species on Noctuidae（Lepidoptera）from the U SSR．． Ent ．0 う（）lr ．62 ： 596 600 （in Russjan ）． Skou， P．，1991． Nordens Ugler． Danm α rks Dyreliv 5．565 pp ．（37 p 且s．）． Apollo Books． Stenstrup． − Sugi， S．＆ M ． Owad 馮 1982． Noctuidae． 動 Inoue， H ． et al ．［Eds ］， Moths qプ 」の 砌 1 ： 669 935，2 ： − − 344 408 164 226 ， pls ． Tshistjakov， Yu ， A ，，1975α ． ［To the knowledge of the lepidoptera of darkconiferous taiga in the − ’ − Southem and Middle SikhotaeAlin］． Entomologicheskie L∬ tedoレania na D4 加 刑 Yostoke： 160 172． Vladivostok．（ln Russian）． ，1975 わ． ［An in且uence of the secondary factors on the faunictic complexes f（）rming in the − − ’ ’ spruce forests of the Middle Sikhotae Alin ］， Entomologicheskie lssledoyania na Dal nem V ∂stoke ： − 173 179． Vladivostok．〔ln Russian）． 1984． of the rnoths efthe superfamilies and ， ［Distribution larger Bombycoidea， Notodontoidea some Noctuoidea within main fbrest fbrmations of the Southern Primorye ］． Fauna i Ekologia ’ − Bespozvonoehn ．アh 1）al nego Vostoka（yrediteli i entomop 乃agi ）： 80 99． Vladivostok ．（In Russian ）． − Vasyurin， V ． D ．1978． Pyadenitsy（Lcp．， Geometridae）sgor Yuzhnogo SikhotaeAlinya．［Geomctrid − ’ − moths （Lep．， Geomc 亡ridae ）from mountains of the Southern SikhotaeAlin Range］． UI∬ 〃π醜 ： 1 24 In Russian ．Manuscript in VINITI No l 549−78 ． （ ） ［ ，deposited ， ］ ． − − Vasyurin， V ， D ．＆ Yu ． A ． Tshistjakov，1979， ［Materialson the high altitude moths fauna （Macrole − ’ − pidQptera）of the Southern SikhotaeAlin ］． β わ logたheskie lssledoyania刀 α D α 肋 e ’η Vost∂ke ： 95 104． Vladivostok．（In Russian）．

摘 要

ロ シ ア 沿 海州 Litbvka山 〔シ ホ テ ア リ ン 山 脈） に お け る 蛾 （Yu ， A ．Tshistjakov，枝 恵太 郎， E A ．Beljaev） − 1995 7 19 21 ロ ン Ljtovka 筆者 ら は 年 月 日 に か け て ， シ ア 沿海州 ， シ ホ テ ア リ 山脈 南西端 に 位置 す る 1 279 m つ 山 （標高 ， ， 沿 海 州 第 2 の 高峰） に て ， 3 の 異 な る 環 境 で 夜 間採集 を実施 した． ・ そ の 結果，小蛾類 を の ぞ く 15 科 343 種 の 蛾類 が 確認 さ れ た ．採集 地 の 環境別 に 見 る と ， ド ロ ノ キ ハ ・ ・ ・ ・ ル ニ レ ケ シ ョ ウ ヤ ナ ギ シ ナ ノ キ 類 モ ン ゴ リ ナ ラ な ど の 落葉広葉樹 に チ ョ ウ セ ン ゴ ヨ ウ エ ゾ マ ツ な ど の 針葉 樹が 混 じ る 混 交林 （標 高 400 m ） で 285 種 ， ト ウ ヒ ・モ ミ類 の 原 生 林 内 （標 高 1，］50m ）で ー 71 種 ，岩 稜 に ウ ス リ ヒ バ が 覆 う山 頂 部 （標 高 1，279m ）で 101種 が 確 認 さ れ た ． こ の う ち ，日本 に な じ み の 薄 い 蛾類 数種 に っ い て 図示 し た ．

新知 見 とし て ， ロ シ ア 初 記録 とな る の は以 下 の シ ャ ク ガ科 3 種 で あ る ，

Seopula asthena lnouc （キ ス ジ シ ロ ヒ メ シ ャ ク 〕 Ven usia laria OberthUr 〔ク ロ ス ジ カ バ イ ロ ナ ミ シ ャ ク）

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84 Yu ． A ． TsHlsTJAKOv Kejtaro EDA and E ． A ． BE 凵 AEv ，

Hydrelia adesma Prout （カ バ イ ロ ヒ メ ナ ミ シ ャ ク ）

また ，沿海州 に お い て初 記録 とな る種 は，以 下 の シ ャ クガ 科 4 種 と ヤ ガ 科 3 種 で あ る．

Idaea aversata （Linnaeus）（エ ゾ キ ヒ メ シ ャ ク） Scopula tenutSocius lnoue （ア メ イ ロ ヒ メ シ ャ ク） PeriZoma taeniatum （Stephens）（ウ ス カ バ ス ジナ ミ シ ャ ク） Euiithis prunata （Linnaeus）（チ ョ ウ セ ン ハ ガ タ ナ ミ シ ャ ク ） Trichosea 如 雌 α （Linnaeus ）（キ タ キ バ ラ ケ ン モ ン ） Polia hepatiαa （Clerck） Apamea sodalts （Butler）（チ ャ イ ロ カ ドモ ン ヨ ト ウ）

（A epted OCtober l l，1997）

Published by the Lepidopterologica】 Society of Japan，

5−20 Motoyokoyama Hachi （’ Tokyo 1920063 Japan ， 2， ji， ，

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