A Checklist of Fishes of Kerala, India

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080

A checklist of fishes of Kerala, India

A. Bijukumar 1 & Rajeev Raghavan 2 ISSN 0974-7907 (Online) Short Communication Short ISSN 0974-7893 (Print) 1 Department of Aquatic Biology and Fisheries, University of Kerala, Thiruvananthapuram, Kerala 695581, India 2 Center for Taxonomy of Aquatic Animals, Department of Fisheries Resource Management, Kerala University of Fisheries and Ocean Studies (KUFOS), Kochi, Kerala 682506, India OPEN ACCESS 2 Laboratory of Systematics, Ecology and Conservation, Zoo Outreach Organization (ZOO), Coimbatore, Tamil Nadu 641035, India 1 [email protected] (corresponding author), 2 [email protected]

Abstract: A checklist of the fishes of Kerala State is presented, along of India recognizes an estimated 3,231 valid species of with their scientific and common names (English and Malayalam), freshwater and marine fishes, of which the total diversity endemism, IUCN Red List status, listing under different Schedules of the Indian Wildlife (Protection) Act and in the Appendices of of marine species comprise 75.6% (2,443 species) (Gopi Convention on International Trade in Endangered Species of Wild & Mishra 2015). Flora and Fauna (CITES). Nine Hundred and five species of fishes are recorded from the inland and marine waters of Kerala comprising Despite over 200 years of exploration and research, of 41 orders and 172 families. Close to 30% of the freshwater fish the fish fauna of peninsular India and Kerala continue species found in Kerala are endemic to the State. Only 8% of the total to be poorly known. Freshwater ichthyological research fishes of Kerala are listed as threatened in the IUCN Red List, of which th the majority are freshwater species. Several hundred fish species in Kerala started in the late 18 century with the occurring in the marine waters of Kerala have not yet been assessed description of Cirrhinus cirrhosus and Labeo fimbriatus for their conservation status by IUCN. by Bloch (1795) from the erstwhile Malabar, followed by Keywords: Brackishwater, fish, freshwater, Kerala, marine, species. the description of Wallago attu from the same region by Bloch & Schneider (1801). Over the next century (1800–1900), several naturalists including Georges Fish diversity known from the fresh and marine Cuvier (1828–1831), Achilles Valenciennes (1840–1847), waters of India constitutes 9.7% of the total number Thomas Jerdon (1849), Albert Günther (1864) and Francis (~33,059 species) of fish species known from the world Day (1865–1889) advanced the knowledge of freshwater (Eschmeyer & Fong 2014). Of these, marine fishes alone fishes of erstwhile Malabar District and Cochin State. account for 7.4%. An assessment of the ichthyodiversity One of the earliest checklists of freshwater fishes of the

Hypselobarbus periyarensis

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DOI: http://dx.doi.org/10.11609/jott.2004.7.13.8036-8080 | ZooBank: urn:lsid:zoobank.org:pub:624B60B8-E050-4C63-BEA4-85B67DEAEB75

Editor: Mewa Singh, University of Mysore, Mysuru, India. Date of publication: 17 November 2015 (online & print)

Manuscript details: Ms # o4308 | Received 11 May 2015 | Final received 23 September 2015 | Finally accepted 29 September 2015

Citation: Bijukumar, A. & R. Raghavan (2015). A checklist of fishes of Kerala, India. Journal of Threatened Taxa 7(13): 8036–8080; http://dx.doi.org/10.11609/ jott.2004.7.13.8036-8080

Copyright: © Bijukumar & Raghavan 2015. Creative Commons Attribution 4.0 International License. JoTT allows unrestricted use of this article in any medium, reproduction and distribution by providing adequate credit to the authors and the source of publication.

Funding: Rajeev Raghavan thanks the Critical Ecosystem Partnership Fund (CEPF), Western Ghats Program and the Mohammed Bin Zayed Species Conservation Fund for funding various projects during which the data for the checklist was compiled.

Conflict of Interest:The authors declare no competing interests.

Acknowledgements: Rajeev Raghavan thanks Neelesh Dahanukar, J.D. Marcus Knight, Siby Philip, Anvar Ali and Unmesh Katwate for providing necessary help and suggestions during the compilation of this checklist.

8036 Fishes of Kerala Bijukumar & Raghavan region - On the fresh water fishes of southern India was fishes of Kerala State. Since the publication of the most published by Jerdon (1849), in which he gave an account recent comprehensive checklist (Kurup et al. 2004), of the fishes of Madras Presidency (including Malabar around 20 new freshwater species have been described, State). in addition to revalidation of the species status of several Francis Day, a medical surgeon of the British army, others. There have also been many new genera and a who later turned to ichthyology, laid the foundation new family of freshwater fish identified from Kerala for fisheries research in Kerala. His book The Fishes during this period (2004–2014). The last one decade of Malabar (Day 1865a) was the first comprehensive witnessed renewed interest in marine fish taxonomy, account on the diversity of fishes off the Kerala coast. with many new species described from the Kerala coast He describes that “while staying at Cochin on the and more records of deep-sea fishes from the country’s Malabar Coast of India, from 1859 to 1864, the annually exclusive economic zone of India. Recent information increasing importance of its fisheries came most compiled by Gopi & Mishra (2015) reveals that 2,443 prominently to my notice. Desirous of ascertaining what species of marine fish are recognized from India, species of fish were most abundant, and which were represented under 230 families and 927 genera; though best adapted for different manufacturers, I was induced state-wise species records and list of species were not to make a collection of those procurable from the sea, mentioned. estuaries or freshwaters, with notes on their habits and The last two years has also witnessed an increasing uses, to elucidate which, plain and coloured drawings of trend of dubious ‘taxonomic’ practices creeping into each species were made as soon as possible after the fish the ichthyological literature of Kerala State thereby had been captured”. Francis Day subsequently published disrupting an otherwise renewed phase in freshwater pioneering accounts on the fishes of the region including fish taxonomy in the region (see Raghavan et al. 2014). On the Fishes of Cochin, on the Malabar Coast of India For the purpose of the present checklist, we have (Day 1865b,c), On the Fishes of the Neilgherry Hills and not considered any description that has appeared Rivers around their Bases (Day 1867a) and On some in a ‘predatory journal’ in which the peer review is Fishes from the Wynaad (Day 1867b). compromised (see Beall 2012, 2015; Raghavan et al. The H.M.S. Challenger Expedition in the late sixties 2014). In this checklist, freshwater fishes are listed in and seventies of theth 19 century triggered interest in brown colored font. The column marked EC (Ecosystem) marine resource surveys in the Indian seas and recorded denotes whether the habitat of the species is marine the occurrence of several marine fishes along the Kerala (M), freshwater (F) or primarily marine but secondary coast. Taxonomic investigations on deep-sea fishes of freshwater (S). India began with the publications of A. Alcock, based on the samples collected during the voyage of Indian References marine survey steamer H.M.S. Investigator (Alcock 1899); and the descriptions also include a few deep- Alcock, A. (1899). A descriptive catalogue of the Indian deep-sea fishes sea fishes off the erstwhile Travancore State. In the 20th in the Indian Museum. Being a revised account of the deep-sea fishes collected by the Royal Indian marine survey ship Investigator. century, several ichthyologists compiled information on Indian Museum, Calcutta, 211pp. marine fish fauna of the Indian Ocean, including those Beall, J. (2015). Scholarly Open Access - critical analysis of scholarly in the western ocean, notable in this series being The open-access publishing. http://scholarlyoa.com/. Accessed on 14 April 2015. Fishes of Indo-Australian Archipelago (Weber & de Beall, J. (2012). Predatory publishers are corrupting open access. Beaufort, 1916–1936; de Beaufort 1940; de Beaufort & Nature 489: 179; http://dx.doi.org/10.1038/489179a Chapman 1951). Bijukumar, A., T.K. Sirajudheen, L. Rajesh & T. Radhakrishnan (2010). Marine fish and fisheries of Kerala- Heritage, diversity and The freshwater fishes of Travancore region remained conservation paradigms, pp. 32–102. In: Leonard, S. & A. Benizger poorly known until the checklists of Pillai (1929), Hora & (eds.). Conservation of Fishes in Kerala, India. World Aquarium, St. Law (1941), Raj (1941) and Silas (1951) were published. Louis, USA. Bloch, M.E. (1795). Naturgeschichte der ausländischen Fische. Berlin. During the last 50 years, several localized checklists v. 9: i-ii + 1-192, pls. 397–429. and compilations confined to smaller regions such as a de Beaufort, L.F. (1940). The Fishes of Indo-Australian Archipelago. E. J. Brill, Leiden 8, 508pp. district, a protected area or a river system of Kerala have Bloch, M.E. & J.G. Schneider (1801). M.E. Blochii, Systema been published. Ichthyologiae Iconibus cx Ilustratum. Post obitum auctoris opus Until this day, very few comprehensive checklists inchoatum absolvit, correxit, interpolavit Jo. Gottlob Schneider, Saxo. Berolini. Sumtibus Auctoris Impressum et Bibliopolio (Chhapgar & Manakadan 2000; Gopi 2000; Kurup et al. Sanderiano Commissum. i-lx + 1-584, Pls. 1-110pp. 2004; Bijukumar et al. 2010) have been published on the Chhapgar, B.F. & R. Manakadan (Eds). (2000). Ecology of Hill Streams

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8037 Fishes of Kerala Bijukumar & Raghavan

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Vernacular Name Vernacular Āṟucekiḷa Srāv Ēḻucekiḷa Srāv Makarasrāv Srāv, Timiṅgala Muḷasrāv Aṟēbyan Muḷasrāv Cāra Īrkkil Muḷasrāv Muḷasrāv Veḷḷappuḷḷi Muḷasrāv Taviṭṭuvarayan Srāv Sībra Nēḻs Srāv Kapilavarṇṇa Srāv Mutala Mākkēāsrāv Cheṟuciṟakan Nilantallisrāv Puṟaṅkaṭal Nilantallisrāv Peruṅkaṇṇan Nilantallisrāv Nāṭan Pūccasrāv Kēāṟal

Malayalam Name Malayalam BdpsNInf {kmhv GgpsNInf {kmhv XnanwKe {kmhv, aIc{kmhv Atd_y³ apf{kmhv Nmc apf{kmhv CuÀ¡nÂ apf{kmhv shÅ¸pÅnapf{kmhv Xhn«phcb³ apf{kmhv ko{_ {kmhv I]nehÀ® t\gvkv {kmhv apXe {kmhv sNdpNndI³ amt¡m{kmhv ]pdwISÂ \newXÃn{kmhv s]cpwI®³ \newXÃn{kmhv \mS³ \newXÃn{kmhv tImdÂ ]q¨{kmhv- (Lowe, 1841) (Lowe, (Matsubara, 1936) (Matsubara, 1810 Rafinesque, 1935 Nakamura, 1788) (Bonnaterre, Authority 1788) (Bonnaterre, Smith, 1828 1783) (Hermann, (Lesson, 1831) 1830) [Bennett], (Anonymous Müller & Henle, 1838 Müller & Henle, (Bonnaterre, 1788) (Bonnaterre, 1838 Müller & Henle, 1830) [Bennett], (Anonymous

Gubanov, 1980 Gubanov, (Gmelin, 1789) (Gmelin, Alopias superciliosus Pseudocarcharias kamoharai Pseudocarcharias Isurus oxyrinchus Alopias pelagicus Alopias vulpinus Species Name griseus Hexanchus typus Rhincodon fasciatum Stegostoma Nebrius ferrugineus marmoratus Atelomycterus Chiloscylilum punctatum Heptranchias perlo Heptranchias Chiloscyllium griseum Chiloscylilum plagiosum

Chiloscyllium arabicum Chiloscyllium indicum Bigeye Thresher Shark Thresher Bigeye Crocodile Shark Crocodile Shark) (Shortfin Mako Shortfin Mako Shark) Shark (Whiptail Thresher Pelagic (Thresher) Common Thresher English Name English I. ORDER HEXANCHIFORMES sharks) (cow Hexanchidae 1. Family Shark) Sixgilled Shark (Cow ORDER ORECTOLOBIFORMES II. (whale sharks) Rhincodontidae 2. Family Whale Shark sharks) (zebra Stegostomatidae 4. Family Shark) Shark (Leopard Zebtra sharks) (nurse Ginglymostomatidae 5. Family Shark) Sleepy Shark (Giant Nurse Tawny ORDER III. LAMNIFORMES (crocodile Pseudocarchariidae 6. Family sharks) ) sharks Lamnidae (mackerel 7. Family sharks) Alopiidae (thresher 8. Family ORDER CARCHARHINIFORMES IV. sharks) Scyliorhinidae (cat 9. Family Shark) (Marbled Cat Catshark Coral Brownspotted Bambooshark Brownspotted Sharpnose Sevengill Shark Sharpnose Sevengill Bamboo Shark Grey Bamboo(Whitespotted Shark Whitespotted Bamboo Shark) 3. Family Hemiscylliidae (bamboo sharks) 3. Family Shark Carpet Arabian Slender Bamboo Shark (Indian Cat Shark, Shark, Slender Bamboo Shark (Indian Cat Bamboo Shark) Ridge-back

1 3 9 8 2 5 7 4 6 14 11 12 13 15 10 16 Table 1. Checklist of fishes of Kerala of fishes 1. Checklist Table

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Vernacular Name Vernacular Intyan Vīkkasrāv Intyan Pūccasrāv Kvāgga Pūccasrāv Rēāma Pūccasrāv Ṟibbaṇvālan Kuḷḷan Vēṭṭanāyasrāv Aṟēbyan Srāv Cūṇṭappallan Srāv Kuṟṟippallan Srāv Śāntan Srāv Pannikkaṇṇan Spinner Srāv Srāv Veḷḷaccekiṭan Silkk Srāv Srāv Peṭṭi Srāv Veḷḷavālan Srāv Mūkkan Srāv Kaṟuttavālan Srāv Karimpuḷḷi Srāv Puḷḷivālan Puḷḷi Srāv Tekkan Srāv, Valiyachiṟakan Srāv Srāv Nīṇṭakaṇṇan Nāraṅṅasrāv Arivāḷciṟakan Srāv Nīlacciṟakan Srāv Pāl Srāv, Peruṁ Srāv Kūrttamūkkan Cāra

Malayalam Name Malayalam C´y³ ho¡{kmhv- Izm¤ ]q¨{kmhv- tcma ]q¨{kmhv- IpÅ³ dnº¬hme³ ]q¨{kmhv Atd_y³ th«\mb{kmhv- Nqï¸Ã³ {kmhv- Ipän¸Ã³ {kmhv im´³ {kmhv ]¶n¡®³ {kmhv kv]ns¶À {kmhv shÅs¨InS³ {kmhv knÂ¡v {kmhv s]«n {kmhv shÅhme³ {kmhv aq¡³ {kmhv Idp¯hme³ {kmhv Icnw]pÅn {kmhv ]pÅnhme³ {kmhv ]pÅn {kmhv henbNndI³ {kmhv, sX¡³ {kmhv \oïI®³ {kmhv AcnhmÄNndI³ \mc§{kmhv \oe¨ndI³ {kmhv s]cpw {kmhv, ]mÂ {kmhv Nmc IqÀ¯aq¡³ {kmhv Quoy & Gaimard, 1824) & Gaimard, Quoy (Müller 1839) & Henle, (Rüppell, 1837) (Rüppell, 1837) (Müller 1839) & Henle, (Alcock, 1899) (Alcock, (Müller 1839) & Henle, (Müller 1839) & Henle, 1861) (Poey, (Klunzinger, 1871) (Klunzinger, (Alcock, 1891) (Alcock, 1899 Hemprich & Ehernberg, 1852) (Bleeker, (Müller 1839) & Henle, (Müller 1839) & Henle, (Müller 1839) & Henle, 1913) (Pietschmann, 1822) & Lesueur, (Péron 1839 Müller & Henle, (Linnaeus, 1758) 1964 Springer, Authority (Müller 1839) & Henle, Smith, 1913 1934) (Whitley, (Talwar, 1974) (Talwar,

Carcharhinus melanopterus Carcharhinus sorrah acutus Rhizoprionodon Negaprion acutidens Carcharhinus macloti Halaelurus quagga Carcharhinus brevipinna Carcharhinus falciformis Carcharhinus longimanus Hemipristis elongata Hemipristis Bythaelurus hispidus Bythaelurus mosis Mustelus Chaenogaleus macrostoma Carcharhinus amboinensis Carcharhinus dussumieri Carcharhinus limbatus Carcharhinus sealei cuvieri Galeocerdo macrorhinus Loxodon glauca Prionace oligolinx Rhizoprionodon Species Name Lamiopsis temminckii Lamiopsis Eridacnis radcliffei Eridacnis Carcharhinus amblyrhynchoides

Cephaloscyllium silasi )

Blacktip Reef Shark Reef Blacktip Shark Spottail Shark) Milk Shark (White-eyed Sicklefin Lemon Shark (Indian Lemon Shark) Sicklefin Hardnose Shark (Maclot's Shark) Hardnose Quagga Catshark Quagga Spinner Shark Silky Shark (Blackspot Shark) Silky Shark (Blackspot Shark) Shark (Whitetip Whitetip Oceanic Snaggletooth Shark (Fossil Elliot'sShark, Shark (Fossil Snaggletooth Shark ) Grey Bristly Catshark Bristly Catshark Ribbontail Pygmy Smoothhound (Hardnosed Arabian Smoothhound) Shark Hooktooth Shark) Shark (Java Pigeye Shark (Widemouth Blackspot Whitecheek Shark) Shark Blacktip Shark Blackspot Shark) Shark (Ground Tiger Shark Sliteye Blue Shark Dog Shark ) Shark (Grey Sharpnose Grey English Name English catsharks) (finback Proscylliidae 10. Family (hound sharks) Triakidae 11. Family sharks) (weasel Hemigaleidae 12. Family sharks) (requiem Carcharhinidae 13. Family Shark Shark (Queensland Graceful Broadfin Shark Broadfin Indian Swellshark (Ground Shark) (Ground Indian Swellshark

32 34 40 38 31 18 26 28 30 23 19 20 21 22 25 27 29 33 35 37 39 41 24 36 17

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Vernacular Name Vernacular Tūmpamūkkan Srāv, Pūḻi Srāv Srāv, Tūmpamūkkan Srāv Kaḷḷa Paviḻasrāv, Veḷḷavālan Chuṟṟikathalayan Kaṇṇankēāṭi Srāv Srāv Cuṟṟikattalayan Taraṅga Srāv Cuṟṟikattalayan Vampan Srāv Cuṟṟikattalayan Caṭṭi Alaṅkārasrāv Maṭiyan Velvaṟṟsrāv Mūkkan Nīṇṭa Srāv Grasana Grasanasrāv Cheṟuciṟakan Ceṟumuḷḷan Nāyasrāv Muḷḷan Srāv Keāmpansrāv Munayan Keāmpansrāv Valiyapallan Keāmpansrāv Valiya Vaidyutatiraṇṭi Taviṭṭu Puḷḷi Vaidyutatiraṇṭi Mārbiḷ Vaidyutatiraṇṭi

Malayalam Name Malayalam Xq¼aq¡³ {kmhv, ]qgn {kmhv shÅhme³ ]hng{kmhv, IÅ {kmhv I®³tImSn NpänI¯eb³ {kmhv XcwK NpänI¯eb³ {kmhv h¼³ NpänI¯eb³ {kmhv N«n NpänI¯eb³ {kmhv aSnb³ Ae¦mc{kmhv- \oï aq¡³ shÂhäv{kmhv {Kk\ {kmhv sNdpNndI³ {Kk\{kmhv sNdpapÅ³ \mb{kmhv apÅ³ {kmhv ap\b³ sIm¼³{kmhv henb]Ã³ sIm¼³{kmhv henb sIm¼³{kmhv Xhn«p sshZypXXncïn ]pÅn sshZypXXncïn amÀ_nÄ sshZypXXncïn Bleeker, 1851 Bleeker, (Linnaeus, 1758) (Barbosa du Bocage & de Brito Brito & de (Barbosa du Bocage 1864) Capello, (Griffith & Smith, 1834) (Griffith & Smith, (Rüppell, 1837) 1816) (Cuvier, (Rüppell, 1837) 1889) (Alcock, 1860 Bleeker, 1903 & Snyder, Jordan 1788) (Bonnaterre, 1794 Latham, Annandale, 1909 Authority 1801) (Bloch & Schneider, 1794) (Latham, 1831 Olfers, (Bloch & Schneidar, 1801) (Bloch & Schneidar,

Müller & Henle, 1838 Müller & Henle, Pristis zijsron Pristis Sphyrna zygaena Sphyrna Centroscymnus crepidater Centroscymnus Sphyrna lewini Sphyrna Triaenodon obesus Triaenodon blochii Eusphyra mokarran Sphyrna ornatum Centroscyllium moluccensis Centrophorus Squalus mitsukurii brucus Echinorhinus microdon Pristis brunnea Narcine Species Name granulosus Centrophorus cuspidata Anoxypristis sinuspersici Torpedo Narcine timlei Narcine

Scoliodon laticaudus Scoliodon Longcomb sawfish Longcomb Smooth Hammerhead (Round-headed Smooth Hammerhead (Round-headed Hammerhead ) Longnose Velvet Dogfish Longnose Velvet Scalloped Hammerhead Scalloped Whitetip Reef Shark Reef Whitetip Winghead Shark Hammerhead Great Dogfish Ornate Smallfin Gulper Shark Shortspine Spurdog Shark Bramble Sawfish Largetooth Numbfish Brown English Name English (hammerheads) Sphyrnidae 14. Family sharks) (sleeper Dalatiidae 15. Family (gulper sharks) Centrophoridae 16. Family Gulper Shark Squalidae (dogfish sharks) 17. Family sharks) (bramble Echinorhinidae 18. Family VI. ORDER PRISTIFORMES (sawfishes) Pristidae 19. Family Narrow Sawfish, (Knifetooth Sawfish Pointed Sawfish) VII. ORDER TORPEDINIFORMES (numbfishes) Narcinidae 20. Family (electric rays) Torpedinidae 21. Family Marbled Electric Ray Spotted Numbfish Spotted V. ORDER SQUALIFORMES V. Spadenose Shark Spadenose

56 47 49 45 43 44 46 48 51 52 53 55 57 50 54 59 58 42

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V ‌ Vernacular Name Vernacular Bhīman Kēārimūkkan Bhīman Kēārimūkkan Gittārmatsyaṁ Vill Gittārmatsyaṁ āyan Gittārmatsyaṁ Annanṇṭele Gittārmatsyaṁ Mūkkan Kūrtta Vītimūkkan Gittārmatsyaṁ Gittārmatsyaṁ Kēārimūkkan Bhīman Gittārmatsyaṁ Muḷḷantiraṇṭi Ñeāṟivālan Muḷḷantiraṇṭi Mūkkan Cāṭṭavālantiraṇṭi Cempāṭan Muḷḷantiraṇṭi Nīṇṭamūkkan Cāṭṭavālantiraṇṭi Kaṇṭal Cāṭṭavālantiraṇṭi Śalkka Cāṭṭavālantiraṇṭi Jālika Nīlappuḷḷi Muḷḷantiraṇṭi Muḷḷantiraṇṭi Paśuvālan Minusa Citraśalabhatiraṇṭi, Tapputiraṇṭi Citraśalabhatiraṇṭi Nīṇṭavālan Puḷḷi Kākkattiraṇṭi Kākkattiraṇṭi Alaṅkāra Bhīman Cekuttāntiraṇṭi Cekuttāntiraṇṭi Nīṇṭakeāmpan Cekuttāntiraṇṭi Aṭappumūkkan

Malayalam Name Malayalam òa³ tImcnaq¡³ Kn¯mÀaÕyw hnÃv- hmb³ Kn¯mÀaÕyw A¶ïse Kn¯mÀaÕyw IqÀ¯ aq¡³ Kn¯mÀaÕyw hoXnaq¡³ Kn¯mÀaÕyw tImcnaq¡³ Kn¯mÀaÕyw òa³ Kn¯mÀaÕyw sRmdnhme³ apÅ³Xncïn aq¡³ apÅ³Xncïn sN¼mS³ Nm«hme³Xncïn \oïaq¡³ apÅ³Xncïn IïÂ Nm«hme³Xncïn iev¡ Nm«hme³Xncïn PmenI Nm«hme³Xncïn \oe¸pÅn apÅ³Xncïn ]iphme³ apÅ³Xncïn an\pk Nn{Xie`Xncïn, X¸pXncïn \oïhme³ Nn{Xie`Xncïn ]pÅn Im¡¯ncïn Ae¦mc Im¡¯ncïn òa³ sNIp¯m³Xncïn \oïsIm¼³ sNIp¯m³Xncïn AS¸paq¡³ sNIp¯m³Xncïn (Blyth, 1860) 1883) (Macleay, 1852) (Bleeker, 1859) (Bleeker, Cuvier, 1829 Cuvier, 1789) (Gmelin, 1775) (Forsskål, Bloch & Schneider, 1801 Bloch & Schneider, 1804) (Shaw, (Anonymous [Bennett], 1830) [Bennett], (Anonymous 1926 Norman, 1841 Müller & Henle, 1775) (Forsskål, (Müller 1841) & Henle, 1851) (Gray, 1801) (Bloch & Schneider, (Müller 1841) & Henle, 1804) (Shaw, 1790) (Euphrasen, 1792) (Walbaum, Authority (Müller 1841) & Henle, 1801) (Bloch & Schneider, Müller & Henle, 1841 Müller & Henle,

Himantura bleekeri Himantura granulata Himantura vespertilio Aetomylaeus Mobula eregoodootenkee Rhinobatos granulatus Rhinobatos uarnak Himantura sephen Pastinachus Rhina ancylostoma thouiniana Rhinobatos Glaucostegus typus Glaucostegus annandalei Rhinobatos obtusus Rhinobatos djiddensis Rhynchobatus zugei Dasyatis gerrardi Himantura imbricata Himantura Neotrygon kuhlii poecilura Gymnura narinari Aetobatus birostris Manta Species Name bennettii Dasyatis micrura Gymnura Rhinoptera javanica Rhinoptera

Bleeker’s Whip Ray Bleeker’s Whipray Mangrove Eagle Ray) (Reticulate Ray Eagle Ornate Longhorned Mobula Sharpnose Guitar Fish Sharpnose Guitar Whipray) (Reticulate Stingray Honeycomb Tailed Stingray) (Frill Stingray Cowtail Bowmouth Guitarfish Bowmouth Fish Guitar Shovelnose Shaw's Giant Shovelnose Ray (Common Shovelnose (Common Shovelnose Ray Shovelnose Giant Ray) (Annandale's Annandale's Guitarfish Ray) Shovelnose Fish Widenose Guitar Shovelnose Spotted Fish (White Guitar Giant ) Ray ) Stingray (Sharpnose Stingray Paleedged Sharpnose Stingray Whipray Scaly Stingray Bluespotted Ray Butterfly Longtailed Eagle Ray Spotted Ray) (Devil Manta Giant Ray) Cownose (Javanese Flapnose Ray English Name English (guitarfishes) Rhinobatidae 22. Family ORDER IX. MYLIOBATIFORMES (stingrays) Dasyatidae 23. Family Stingray) (Frilltailed Stingray Bennett's rays) (butterfly Gymnuridae 24. Family Ray Smooth Butterfly and manta (eagle Myliobatidae 25. Family rays) VIII. ORDER RAJIFORMES

69 71 79 81 63 73 75 61 65 60 62 64 66 68 70 72 74 77 78 80 82 67 76

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Vernacular Name Vernacular Mūkkan Kimēṟa Mūkkan Ampaṭṭanvāḷa Vaḷḷippūmīn Pālānkaṇṇi Eli Mīn Puḷḷi Maliññīl Maliññīl Kaṟutta Meāṟemaliññīl Veḷḷa Meāṟemaliññīl Veḷḷamukhan Meāṟemaliññīl Sībra Meāṟemaliññīl Alaṅkāra Meāṟemaliññīl Kaṟuppkeṭṭan Meāṟemaliññīl Mañña'arikan Meāṟemaliññīl Ṭarkki Meāṟemaliññīl Jālikā Meāṟemaliññīl Varayan Meāṟemaliññīl Kaṭukka Meliñña Bhīman Meāṟemaliññīl

Malayalam Name Malayalam aq¡³ Intad A¼«³hmf hÅn¸qao³ ]mem³I®n Fen ao³ ]pÅn aenªoÂ Idp¯ aenªoÂ shÅ samsd aenªoÂ shÅapJ³ samsd aenªoÂ ko{_ samsd aenªoÂ Ae¦mc samsd aenªoÂ Idp¸vsI«³ samsd aenªoÂ aªAcnI³ samsd aenªoÂ SÀ¡n samsd aenªoÂ PmenIm samsd aenªoÂ hcb³ samsd aenªoÂ ISp¡ samsd aenªoÂ saenª `oa³ samsd aenªoÂ (Pallas, 1769) (Pallas, (Linnaeus, 1758) McClelland, 1844 1856) (Kaup, 1797) (Shaw, & Randall, 1982) (McCosker 1795) (Shaw, (McClelland, 1844) Authority 1931) (Schnakenbeck, 1775) (Forsskål, 1782) (Broussonet, 1831) (Gray, Schultz, 1943 1801 Bloch & Schneider, (Rüppell, 1830) Bloch, 1795 (Lacepède, 1803) (Hamilton, 1822) (Hamilton,

Notopterus notopterus Notopterus Albula vulpes Anguilla bicolor delicatula Echidna zebra Gymnomuraena enigmaticus Gymnothorax meleagris Gymnothorax rueppelliae Gymnothorax Species Name pinnata Neoharriota machnata Elops cyprinoides Megalops Anguilla bengalensis Echidna leucotaenia Echidna favagineus Gymnothorax flavimarginatus Gymnothorax reticularis Gymnothorax undulatus Gymnothorax Strophidon sathete

Bronze Featherback Bronze Bone Fish Eel )¹ Eel (Shortfin Indonesian Shortfin Mottled Moray Moray) (Reticulated Moray Zebra (Banded moray) moray Enigmatic Moray) (Painted Moray Turkey Moray) (Rupell’s Banded Moray English Name English X. ORDER CHIMAERIFORMES Rhinochimaeridae (longnose 26. Family chimaeras) (Longnose Chimaera) Chimaera Sicklefin ORDER XI. OSTEOGLOSSIFORMES (featherback) Notopteridae 27. Family ORDER ELOPIFORMES XII. Elopidae (tenpounders) 28. Family (Ladyfish ) Tenpounder (tarpons) Megalopidae 29. Family Tarpon) Tarpon (Oxeye Indo-Pacific ORDER ALBULIFORMES XIII. Albulidae (bonefishes) 30. Family ORDER ANGUILLIFORMES XIV. eels) Anguillidae (freshwater 31. Family Eel )¹ Eel (Indian Longfin Indian Mottled eels) (moray Muraenidae 32. Family Whiteface Moray Whiteface moray Laced Moray Yellow-Edged moray) (Dusky-banded Morey Reticulated Moray Undulated Slender Giant Moray (Gangetic Moray) (Gangetic Moray Slender Giant

84 87 89 90 92 94 96 98 83 85 86 88 91 93 95 97 99 100

8042 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

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̥ duvayaṟan Matti duvayaṟan Vernacular Name Vernacular Ciṟakan Pāmpmaliññīl Ciṟakan Maṇal Maliññīl Jīni Pāmpmaliññīl Maliññīl Nīṇṭaciṟakan Maliññīl Vayal Mīśa Kēāṅṅar Mañña Kēāṅṅar Pāmpu Kēāṅṅar, Paikk Intyan Mīn Veḷḷi Kēāṅṅar, Paikk Nāṭan Maliññīl Kēāṅṅar Paikk Kattippallan Uruḷan Nettēāli Ḍēyuṭe Nūna Matti Maḻavil Malabār Matti Cūṭa Vēḷūri, Matti Nīlavarayan Nūna Nūlciṟakan Puḷḷi Matti Mr Chāḷa Vaṭṭi Cāḷa, Parappan Matti Valiya Matti Maṟiṣyan Matti Cetumpal Añcala Matti Svarṇavarayan

Malayalam Name Malayalam NndI³ ]m¼vaenªoÂ aWÂ aenªoÂ Po\n ]m¼vaenªoÂ \oïNndI³ aenªoÂ hbÂ aenªoÂ aoi tIm§À aª tIm§À C´y³ ss]¡v tIm§À, ]m¼p ao³ \mS³ ss]¡v tIm§À, shÅn aenªoÂ I¯n¸Ã³ ss]¡v tIm§À tUbpsS Dcpf³ s\t¯men \q\ aghnÂ a¯n ae_mÀ a¯n thfqcn, NqS \oehcb³ a¯n \qÂNndI³ \q\ ]pÅn a¯n arZphbd³ a¯n ]c¸³ Nmf, h«n Nmf henb a¯n audnjy³ a¯n A©e sNXp¼Â a¯n kzÀWhcb³ a¯n Authority 1853) (Bleeker, (McClelland, 1844) 1848) (Richardson, 1845) (Richardson, 1775) (Forsskål, 1929 Deraniyagala, (Rüppell, 1837) 1792) (Walbaum, 1847) (Valenciennes, (Lacepède, 1803) 1849) (Bleeker, Vahl, 1794 Vahl, 1839) & Bennett, Lay 1822) (Hamilton, Rüppell, 1830 1822) (Hamilton, 1873) (Day, 1847 Valenciennes, 1847) (Valenciennes, (Bloch, 1795) 1849 Bleeker, 1852 Bleeker, 1847) (Valenciennes, (Hamilton, 1822) (Hamilton,

Species Name talabonoides Congresox Lamnostoma orientalis Lamnostoma Pisodonophis cancrivorus lepturus Uroconger cinereus Muraenesox Ehirava fluviatilis quadrimaculatus Herklotsichthys sirm Amblygaster Sardinella albella Sardinella jussieu Sardinella gibbosa Caecula pterygera Leiuranus semicinctus Pisodonophis boro Conger cinerens bagio Muraenesox Dayella malabarica Dussumieria acuta Escualosa thoracata nasus Nematalosa clupeoides Amblygaster Sardinella brachysoma Sardinella fimbriata Anodontostoma chacunda Anodontostoma

English Name English eels) (snake Ophichthidae 33. Family and garden Congridae (conger 34. Family eels) ) congers (pike Muraenesocidae 35. Family Conger Indian Pike ORDER CLUPEIFORMES XV. Clupeidae (herrings, shads, 36. Family menhadens) sardines, Oriental Sand Eel (Oriental Worm Eel) Worm (Oriental Sand Eel Oriental Longfin Snake-eel Pike-Conger) (Yellow Slender Conger Conger Pike Daggertooth Malabar Sprat Herring Bluestripe Sardinella Spotted Sardinella White Mauritian Sardinella Sardinella Goldstripe Finny Snake Eel Snake Finny Saddled Snake-eel Eel Rice-Paddy Conger) (Moustache Conger Longfin African Eel) Siver Eel, (Pike Conger Common Pike Herring Round Day's Sardine Rainbow Sardine White Shad (Hairback) Bloch's Gizzard (Sharpnose Smoothbelly Sardinella Bleeker Sardine) (Indian Sprat) Deepbody Sardinella Sardinella Fringescale Chacunda Gizzard-Shad Chacunda Gizzard-Shad

108 102 104 107 110 114 116 118 120 122 124 101 103 105 106 109 111 113 115 117 119 121 123 112

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8043 Fishes of Kerala Bijukumar & Raghavan

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‌ ‌ Vernacular Name Vernacular Nalla Matti Matti Kaṟuppucuṭṭi Sindh Matti Hilsa Kēānettēāli Ḍēvis Kēānettēāli Ceṟutalayan Kēānettēāli Bukkānīr Nettēāli Bagān Nettēāli Keām'mēḻsan Nettēāli Intyan Nettēāli Hārḍenbarg Nettēāli Peāṭṭumukhan Neṭumaṇaṅṅ Maṇaṅṅ Hāmilṭṭan Malabār Maṇaṅṅ Mīśa Maṇaṅṅ Maṇaṅṅ Neṭuntāṭi Maṇaṅṅ Ōṟañcvāyan Pūmīn Peruvayamp Peruvayamp Ṟēba Paccilaveṭṭi Malabār Pāvukan Puḷḷi Pāvukan

Malayalam Name Malayalam \Ã a¯n Idp¸pNp«n a¯n knÔv a¯n lnÂk tUhnkv tIms\t¯men sNdpXeb³ tIms\t¯men _p¡m\oÀ tIms\t¯men _Km³ s\t¯men sImt½gvk³ s\t¯men C´y³ s\t¯men lmÀsU³_ÀKv s\t¯men s]m«papJ³ s\t¯men s\SpaW§v lmanÂ«³ aW§v ae_mÀ aW§v aoi aW§v s\SpwXmSn aW§v Hmd©v hmb³aW§v ]qao³ s]cphb¼v- s]cphb¼v- td_ ]¨nesh«n ae_mÀ ]mhpI³ ]pÅn ]mhpI³ Authority 1940) (Whitley, 1775) (Forsskål, (Cuvier, 1829) (Cuvier, 1822) (Hamilton, 1938 Fowler, 1803 Lacepède, 1933 Hardenberg, 1848) (Valenciennes, (Bloch, 1795) 1782) (Broussonet, 1849) (Jerdon, (Day, 1878) (Day, (Rüppell, 1837) 1933 Hardenberg, 1823) Hasselt, (van 1926 & Seale, Jordan 1835 Gray, 1801) (Bloch & Schneider, 1908) & Thompson, Gilchrist 1844) (Valenciennes, 1807) (Hamilton, 1865) (Day (Hamilton, 1807) (Hamilton,

(Bleeker, 1853) (Bleeker, Valenciennes, 1847 Valenciennes, Species Name devisi Encrasicholina Chanos chanos Sardinella melanura ilisha Tenualosa punctifer Encrasicholina commersoni Stolephorus insularis Stolephorus Thryssa dussumieri Thryssa malabarica Thryssa setirostris carnaticus Barbodes Sardinella sindensis heteroloba Encrasicholina baganensis Stolephorus indicus Stolephorus waitei Stolephorus Thryssa hamiltonii Thryssa mystax Thryssa vitrirostris Amblypharyngodon melettinus Bangana ariza Barilius bakeri Barilius bendelisis

Amblypharyngodon microlepis Sardinella longiceps English Name English (anchovies) Engraulidae 37. Family Anchovy Devis' XVI. ORDER GONORHYNCHIFORMES Chanidae (milkfish) 38. Family Milk Fish XVII. ORDER CYPRINIFORMES Cyprinidae (carplet) 39. Family Blacktip Sardinella Blacktip Hilsa (Hilsa Shad) Anchovy Buccaneer Anchovy Commerson's Anchovy Hardenberg's (Long Anchovy) Dussumier's Thryssa Malabar Thryssa Thryssa Longjaw Carp Carnatic Sind Sardinella Shorthead Anchovy anchovy Bagan Indian Anchovy Anchovy Spotty-Face Thryssa Hamilton's Thryssa Moustached Anchovy Orangemouth Carplet Silver Carp Ariza Baril Baker's Spotted Baril Spotted Indian Carplet Indian Oil Sardine

129 143 126 128 131 133 135 137 139 141 147 127 130 132 134 136 138 140 142 144 146 148 149 145 125

8044 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

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Vernacular Name Vernacular Varayan Pāvukan Varayan Pāvukan Jerḍanṟe Ḍāniyēā Varayan Paral Kāḷakkeāṭiyan Āścaryaparal Paral Pūvāli Paral Rēāhan Muppuḷḷi Paral Oḻukkilāṭṭi Oḻukkilāṭṭi Taṭiyan Nīlagiri Oḻukkilāṭṭi Īṟṟilakkaṇṭa Mīśappaṟava Mīśappaṟava Veḷḷi Mīśappaṟava Varayan Kalleāṭṭi Aruṇācalaṁ Kalleāṭṭi Kuḻivālan Kalleāṭṭi Veṇṇa Nīla Kalleāṭṭi Kalleāṭṭi Kuḷḷan Kalleāṭṭi Mlāppāṟa Kallunti Kalleāṭṭi Periyār Cuṇṭan Kalleāṭṭi Kaṟumpan Karimpācci Varayan Nīlagiri Vāḻakkā Varayan Vāḻakkā Kaṇaññēān Āṟṟu Kūral Malayalam Name Malayalam hcb³ ]mhpI³ sPÀU³sd ]mhpI³ hcb³ Um\ntbm Imfs¡mSnb³ ]cÂ BÝcy]cÂ ]qhmen ]cÂ tcml³ ]cÂ ap¸pÅn ]cÂ Hgp¡nem«n XSnb³ Hgp¡nem«n \oeKncn Hgp¡nem«n Cuäne¡ï aoi¸dh shÅn aoi¸dh hcb³ aoi¸dh AcpWmNew IsÃm«n Ipgnhme³ IsÃm«n sh® IsÃm«n \oe IsÃm«n IpÅ³ IsÃm«n ¾m¸md IsÃm«n IÃp´n s]cnbmÀ IsÃm«n Npï³ Idp¼³ IsÃm«n Icn¼m¨n \oeKncn hmg¡m hcb³ hmg¡m hcb³ Bäp IWtªm³ IqcÂ Authority (Hamilton, 1822) (Hamilton, 2005) & Kottelat, (Pethiyagoda & Knight, Indra Devi, (Rema 2010) (McClelland, 1839) 1867) (Day, 1849) (Jerdon, 1842) (Valenciennes, 2011 & Radhakrishnan, Kurup 1849) (Jerdon, 2011 & Radhakrishnan, Kurup Gopi, 2001 1996 Shaji, & Easa, Arun 1849) (Jerdon, 1992 & Menon, Devi Rema Jerdon,1849 1849) (Jerdon, 1849) (Jerdon, 1849) (Jerdon, (Raj, 1941) 1822) (Hamilton, 2001 Johnson & Soranam, 1849) (Jerdon, 1865) (Day, 1876) (Day, Rema Devi & Indra, 1984 & Indra, Devi Rema (Valenciennes, 1844) (Valenciennes, (Valenciennes, 1844) (Valenciennes, Silas, 1955 1839) (Sykes, 1996 Menon & Jacob, Species Name Danio rerio Dawkinsia exclamatio Dawkinsia rohani aequipinnatus Devario neilgherriensis Devario Esomus barbatus Esomus thermoicos Garra emarginata Garra mcclellandi Garra mlapparaensis Garra periyarensis Garra surendranathanii Haludaria fasciata Horadandia brittani Barilius malabaricus Dawkinsia assimilis Dawkinsia rubrotinctus malabaricus Devario ophicephalus Eechathalakenda Esomus danricus Garra arunachalami Garra stenorhynchus Haludaria melanampyx dobsoni Hypselobarbus Garra menoni Barilius gatensis Dawkinsia filamentosa Garra hughi Garra mullya periyarensis Gonorhynchus English Name English Zebra Fish Zebra Barb³ Exclamatio Barb² Rohan's Danio Giant Nilgiri Danio⁴ South Indian Flying Barb Flying Barb Sucker⁷ Stone Emarginate Sucker⁹ McClelland's Stone Sucker¹¹ Stone Mlappara Sucker¹¹ Stone Periyar Sucker⁸ Stone Surendran's Nilgiri Melon Barb¹³ Glass Carplet Malabar Baril Barb² Mascara Spot Barb² Three Malabar Danio Barb⁵ Snakehead Common Flying Barb Sucker⁶ Arunachalam's Stone Sucker Horned Stone Sahyadri Melon Barb¹³ Carp) Carp (Dobson's Krishna Menon's Stone Sucker¹⁰ Menon's Stone Emerald Baril Emerald Filament Barb² Filament Sucker⁸ Hughe's Stone Sucket Striped Stone Latia¹² Periyar 152 154 156 158 160 162 164 166 168 170 172 174 176 178 151 153 157 159 161 163 165 173 177 179 169 150 155 167 171 175

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8045 Fishes of Kerala Bijukumar & Raghavan

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ceviyan Paral ceviyan ̄ Vernacular Name Vernacular Tēnkūral Kūral Kānaṟa Mīn Kēāḻi Kūral Karivālan Kariyān Cempankūral Pullan Tūḷi, Lēbiyēā Ñeāṟiccuṇṭan Kākkamīn Puḷḷi Cīlan Cīlan Varayan Cīlan Matti Brāhmaṇakaṇṭa Mañña Kaṭanna Paral Tiriccaṟiyā Cem'muḷḷan Pāval Nīlagiri Muḷḷanparal Matsyaṁ Maccaḷ Mēāḍēān Maramīn Māmaḷ Paral Paisa Kaṟun Paral Pūkkēāṭan Svarṇavālan Paṭṭaruparal Paral Irupeāṭṭu Paral Kāvēri Paral Paral Ceṟumūkkan Malayalam Name Malayalam tX³IqcÂ Im\d IqcÂ tImgn ao³ Icnhme³ IqcÂ Icnbm³ sN¼³IqcÂ Xqfn, ]pÃ³ sRmdn¨pï³ te_ntbm Im¡ao³ ]pÅn Noe³ hcb³ Noe³ a¯n Noe³ {_mÒWIï aª IS¶ Xncn¨dnbm ]cÂ sN½pÅ³ ]mhÂ \oeKncn apÅ³]cÂ a¨fv aÕyw tamtUm³ acao³ amafv ss]k ]cÂ IdpwsNhnb³ ]cÂ ]qt¡mS³ ]cÂ kzÀWhme³ ]«cp]cÂ Ccps]m«p ]cÂ Imthcn ]cÂ ]cÂ sNdpaq¡³ ]cÂ Authority (Day, 1874) (Day, 1842) (Valenciennes, 1995 Devi, Menon & Rema (Raj, 1941) 1842) (Valenciennes, (Bloch, 1795) 1870) (Day 1952 Menon, (Silas, 1958) 1822) (Hamilton, Raj, 1941 2015 & Kumar Knight 1873) (Day, 1873) (Day, 1994 & Kottelat, Pethiyagoda 1877) (Day, 1822) (Hamilton, & Indra Devi, Rema (Knight, 2012) Arunachalam, 2007) & Jacob, (Mercy 1865) (Day, 1822) (Hamilton, 1863) (Bleeker, 1937) (Hora, 1822) (Hamilton, 1849) (Jerdon, (Day, 1870) (Day, (Day, 1874) (Day, 1873) (Day, 1873) (Day, 1869) (Day, Species Name Hypselobarbus lithopidos Hypselobarbus micropogon Hypselobarbus mussullah Hypselobarbus periyarensis Hypselobarbus Labeo dussumieri Labeo fimbriatus Labeo nigrescens dadiburjori Laubuca fasciata Laubuca laubuca Laubuca typus Lepidopygopsis incognito Oreichthys neilli Osteobrama brevidorsalis Osteochilichthys longidorsalis Osteochilichthys thomassi Osteochilichthys conchonius Pethia nigripinna Pethia pookodensis Pethia punctata Pethia ticto Pethia bimaculatus Puntius cauveriensis Puntius chola Puntius dorsalis Puntius Hypselobarbus jerdoni Hypselobarbus Hypselobarbus thomassi Hypselobarbus Neolissochilus wynaadensis bakeri Osteobrama nashii Osteochilichthys English Name English Canara Barb¹⁴ Canara Barb¹⁵ Korhi Barb Kurali Barb¹¹ Periyar Malabar Labeo¹⁶ Lipped Carp Fringe Black Labeo Burjor's Brilliance (Dadio) Malabar Leaping Barb Indian Glass Barb Hill Barb¹¹ Periyar High Fin Barb¹⁸ Kerala Neil's Barb⁹ Barb⁹ Kantaka Barb¹⁹ Long Finned Kerala Thomas' Barb Barb² Rosy Black Finned Barb² Barb² Pookode Barb² Sawfin Dotted Barb² Ticto Barb Redside Barb Cauvery Chola Barb Barb Long Snouted Jerdon's Carp Jerdon's Red Canarese Barb Canarese Red Mahseer¹⁷ Wayanad Barb Baker's Barb Nash's 181 182 183 184 186 187 188 189 190 191 192 194 196 197 198 200 201 202 203 204 205 206 207 208 209 180 185 193 195 199

8046 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

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Vernacular Name Vernacular Madhusūdana Paral Uruḷan Paral Paral Karimpuḷḷi Pāṟapparal Kuḷapparal Paral Kaypa Kuṭiyan Tuppal Cēārakkaṇiyān Ceṅkaṇiyān Paral Katti Paral Peṟumatti Cāḷa Paral Paral Kuṟuva Mīn Kuyil Kaṟṟi Mīn Kuyil Maṇalārēān Ceṟupūntārakan Jalappalli Kalnakki Muttuccuṭṭan Kalnakki Kalpūḷēān Veḷumpan Kalnakki Pacca Kalnakki Kaṟumpan Kalkkāri Silās Kalppūḷēān Kalnakki Neṭuṁ

Malayalam Name Malayalam a[pkqZ\ ]cÂ Dcpf³ ]cÂ Icn¼pÅn ]cÂ ]md¸cÂ Ipf¸cÂ Ibv] ]cÂ Xp¸Â IpSnb³ tNmc¡Wnbm³ sN¦Wnbm³ I¯n ]cÂ sNdpa¯n ]cÂ Nmf ]cÂ Ipdph ]cÂ IpbnÂ ao³ Iän IpbnÂ ao³ aWemtcm³ sNdp]q´mcI³ Pe¸Ãn IÂ\¡n ap¯p¨p«³ IÂ\¡n shfp¼³ IÂ]qtfm³ ]¨ IÂ\¡n Idp¼³ IÂ\¡n IÂ¡mcn knemkv IÂ¸qtfm³ s\Spw IÂ\¡n Authority 1846) (Valenciennes, Day, 1865 Day, & Thobias, Devi (Menon, Rema 1999) 1844) (Valenciennes, 1874) (Day, 1839) (Sykes, 2011 & Radhakrishnan, Kurup 1995) (Shaji & Easa, 1981) Devi, & Rema (Indra (Day, 1878) (Day, 1822) (Hamilton, 1844) (Valenciennes, 1865) (Day, 1839) (Sykes, 1842) (Valenciennes, 1849) (Jerdon, (Tilak 1972) Ali, Jadhav Tharian, Raghavan, 2012 & Dahanukar, Jerdon,1849 1945) (Herre, Johnson(Arunachalam, & 2002) Devi, Rema 1994 & Kottelat, Pethiyagoda

Kumar, Pereira & Pereira Kumar, 2012 Radhakrishnan, (Valenciennes, 1844) (Valenciennes, 1865 Day, 1941 Hora, & Radhakrishnan, 2011) (Kurup Species Name thermalis Lepidocephalichthys Puntius vittatus Puntius chalakkudiensis Sahyadria Salmophasia acinaces Salmophasia boopis khudree Tor remadeviae Tor Ghatsa menoni Ghatsa pillaii Puntius melanostigma Puntius sophore Puntius Rasbora dandia denisonii Sahyadria Salmophasia balooke subnasutus Systomus malabaricus Tor goaensis Pangio jalpalli Balitora Bhavania australis Ghatsa montana Ghatsa santhamparaiensis elongata Travancoria

Puntius madhusoodani Puntius Puntius mahecola Puntius parrah Puntius mysorensis Balitora Ghatsa silasi English Name English loaches) Cobitidae (spiny 40. Family Loach Common Spiny loaches) (stone Balitoridae 41. Family Green Stripe Barb Green Barb² Torpedo Redline Chalakudy Minnow²⁰ Razorbelly Silver Minnow²⁰ Boopis Razorbelly Mahseer Deccan Chinnar Mahseer²¹ Loach²³ Menon's Stone Loach²³ Pillai's Stone One spot Barb Barb Pool Black Line Rasbora Denison's Barb (Miss Kerala)² Minnow²⁰ Razorbelly Balooke Barb² Swamp Malabar Mahseer Indian Coolie Loach Loach²² Stone Valley Silent Loach Stone Bhavani Loach²⁴ Anamalai Stone Loach²⁵ Stone Santhampara Loach⁸ Stone Elongated Madhusoodan's Barb¹⁶ Mahe Barb Mahe Barb Parrah Loach⁹ Stone Mysore Loach¹¹ Silas's Stone

226 215 217 219 221 223 225 231 233 212 214 216 218 220 222 224 227 228 230 232 234 236 210 211 213 229 235

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8047 Fishes of Kerala Bijukumar & Raghavan

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KL KL KL KL KL KL KL KL WG WG WG WG WG WG WG WG WG WG WG WG WG WG WG WG WG END

LC LC LC LC LC LC LC LC LC LC LC LC LC LC LC CR CR NT EN EN EN VU VU VU VU VU VU IUCN Vernacular Name Vernacular Kuḷḷan Kalnakki Kuḷḷan Keāyma Caturavālan Keāyma Kēraḷa Keāytta, Kēraḷa Keāyma Pacca Ānamala Keāyma Keāyma Mēnēān Keāyma Pāmpār Keāyma Periyār Sundari Keāyma Keāyma Kunti Keāytta Pāṇṭan Keāyma Kaṟunvarayan Puḷḷi Keāyma Keāyma Varayan Nīlagiri Keāyma Ceṟupuḷḷi Ceṟupuḷḷi Keāyma, Keāytta Keāyma Oliyavarayan Nīlakkūri Ēṭṭakkūri Maññakkūri Maññakkūri Kariṅkaḻuttan Kūri Kuḷḷan Cakkamuḷḷan Malabār Kūri Cillankūri Malayan Cuṭṭikkūri Kūri Maññavayaṟan

Malayalam Name Malayalam IpÅ³ IÂ\¡n NXpchme³ sImbva tIcf sImbv¯, sImbva ]¨ sImbva B\ae sImbva tat\m³ sImbva ]m¼mÀ sImbva s]cnbmÀ sImbva kpµcn sImbva Ip´n sImbva ]mï³ sImbv¯ Idpwhcb³ sImbva ]pÅn sImbva hcb³ sImbva \oeKncn sImbva sNdp]pÅn sImbva, sImbv¯ Henbhcb³ sImbva \oe¡qcn G«¡qcn aª¡qcn IcnwIgp¯³ aª¡qcn IpÅ³ Iqcn N¡apÅ³ ae_mÀ Iqcn aeb³ NnÃ³Iqcn Np«n¡qcn aªhbd³ Iqcn Authority 1839) (Sykes, 1941 & Law, Hora (Day, 1867) (Day, & Minimol, 1999) (Zacharias Annandale, 1919 1867) (Day, 1867) (Day, 1864) (Günther, 1849) (Jerdon, 1840) (Valenciennes, (Rita & Nalbant, 1978) & Nalbant, (Rita 1982 & Bănărescu, Nalbant 1994) & Indra, Devi (Rema 1873) (Day, 1865) (Day, 1921 Hora, (Menon 1987) 1867) (Day, 1849) (Jerdon, 1994 & Kottelat, Pethiyagoda 1822) (Hamilton, 1849) (Jerdon, (Bloch, 1794) (Kurup & Radhakrishnan, 2005) (Kurup

Hora, 1941 Hora, Shaji, 2002 1865) (Day, Species Name mooreh Acanthocobitis travancoria Batasio Mesonoemacheilus guentheri Mesonoemacheilus menoni Mesonoemacheilus Nemacheilus anguilla denisoni Schistura semiarmata Schistura Horabagrus brachysoma malabaricus Mystus oculatus Mystus Indoreonectes keralensis Indoreonectes herrei Mesonoemacheilus pambarensis Mesonoemacheilus pulchellus Mesonoemacheilus triangularisMesonoemacheilus Nemacheilus monilis nilgiriensis Schistura striata Schistura Hemibagrus punctatus Horabagrus nigricollaris cavasius Mystus montanus Mystus vittatus Mystus Mesonoemacheilus periyarensis Mesonoemacheilus

Travancoria jonesi Travancoria Mesonoemacheilus remadeviae Mesonoemacheilus armatus Mystus English Name English Nemacheilidae (stream/river 42. Family loaches) Loach Mooreh XVIII. ORDER SILURIFORMES catfishes) Bagridae (river 43. Family Batasio Travancore Gunther's Loach Gunther's Loach¹¹ Menon's River Black Lined Loach Denison's Loach Loach Small-spotted Catfish) (Gunther's Catfish Yellow Malabar Mystus Mystus Spotted Cardamom Hills River Loach⁵ Hills River Cardamom Anamalai Loach²⁴ Loach²¹ Pambar Loach Spotted Pretty Loach Zodiac Black Bead Loach Nilgiri Loach⁴ Long Bodied Striped Loach²⁶ Catfish²⁷ Giant Cauvery Catfish²⁸ Collared Imperial Mystus Gangetic Mystus Wynad Striped Mystus Periyar Loach¹¹ Periyar Jone's Stone Loach⁸ Stone Jone's Remadevi's Loach²² Remadevi's Catfish Mystus Dwarf

238 254 240 242 248 250 252 256 260 262 239 241 243 245 247 249 251 253 255 257 259 261 263 244 237 246 258

8048 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

CITES

WPA

KL KL KL KL KL KL KL KL KL KL WG WG WG WG WG WG WG END Exotic *

LC LC LC LC NT NT NT NE EN NE EN EN EN NE EN EN NE DD DD DD DD LC IUCN

̥ ṣṇa Kuruṭanmuśi Vernacular Name Vernacular Bhīman Āṟṟuvāḷa Teāṇṇivāḷa Pulluvāḷa Vayanāṭanvāḷa Vāḷa Āṟṟu Miḍu Veḷḷivāḷa Muśi Peāṅṅan Pāṟakkūri Veḷḷikkeṭṭan Pāṟakkūri Naṭuvarayan Iruḷan Pāṟakkūri Pāṟakkūri Ēlakkāṭan Kalkkāri Pāṟakkūri Maññavayaṟan Malabār Pāṟakkūri Āṟṟukūri Tekkēṣyan Āsṭrina Muśi Vayanāṭan Muśi Nāṭan Muśi Āphrikkan Kuruṭanmuśi Abduḷkalāṁ Kuruṭanmuśi Alikkuññi Kr Malayalam Name Malayalam `oa³ Bäphmf sXm®nhmf ]pÃphmf hb\mS³hmf Bäp hmf anUp shÅnhmf s]m§³ apin shÅns¡«³ ]md¡qcn \Sphcb³ ]md¡qcn Ccpf³ ]md¡qcn Ge¡mS³ ]md¡qcn IÂ¡mcn aªhbd³ ]md¡qcn ae_mÀ ]md¡qcn Bkv{Sn\ sXt¡jy³ BäpIqcn hb\mS³ apin \mS³ apin B{^n¡³apin AÐpÄIemw IpcpS³apin Aen¡pªn IpcpS³apin IrjvW IpcpS³apin Authority (Bloch, 1794) (Valenciennes, 1840) (Valenciennes, 1801) (Bloch & Schneider, 2011 & Thomas, Vincent 1864 Günther, 1822) (Hamilton, Silas, 1952 1998 & Das, Manimekalan 1942 Herre, Gopi, 2010 Sureshkumar Radhakrishnan, & Ng, 2011 1936 Hora, 2004 Babu & Nayar, (Day, 1873) (Day, 1923 Hora, 1873) (Day, 1840 Valenciennes, Menon, 1950 Menon,

(Burchell 1822) (Burchell Babu, 2012 Babu, (Sykes, 1839) (Sykes, Plamoottil & Abraham, 2013 Plamoottil & Abraham, Species Name Ompok bimaculatus Ompok malabaricus attu Wallago shajii Kryptoglanis Pseudeutropius mitchelli pangasius Pangasius anamalaiensis Glyptothorax davissinghi Glyptothorax housei Glyptothorax malabarensis Glyptothorax Pseudolaguvia austrina Clarias dayi Horaglanis alikunhii Pterocryptis wynaadensis Pterocryptis annandalei Glyptothorax madraspatanus Glyptothorax Clarias dussumieri Horaglanis krishnai Clarias gariepinus Horaglanis abdulkalami Sperata seenghala Sperata Glyptothorax elankadensis Glyptothorax * English Name English catfishes) Siluridae (butter 44. Family Catfish Butter (blind catfishes) Kryptoglanidae 45. Family catfishes) Schilbeidae (river 46. Family catfishes) (pangasiid Pangasiidae 47. Family catfishes) Sisoridae (mountain 48. Family catfishes) (torrent Erethistidae 49. Family catfishes) Clariidae (river 50. Family (stinging Heteropneustidae 51. Family catfishes) Malabar Butter Catfish Malabar Butter Shark Freshwater Shaji's Blind Catfish²⁹ Catfish River Mitchell's Shark Catfish Catfish Anamalai Mountain Catfish³⁰ Nilambur Mountain Catfish Mountain Valparai Catfish³² Malabar Mountain Catfish Southern Indian Torrent Malabar Clarid Catfish African Catfish²⁹ Blind Cave Alikunhi's Wayanad Catfish Wayanad Catfish Annandale's Mountain Catfish Mountain Madras Clarid Valencienne's Blind Cave Catfish³³ Blind Cave Abdulkalam's Blind Cave Catfish²⁹ Blind Cave Abdulkalam's Giant River Catfish River Giant Elankadu Mountain Catfish³¹ Mountain Elankadu

265 266 268 269 270 271 272 274 276 278 279 280 282 284 267 273 277 281 285 283 264 275

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8049 Fishes of Kerala Bijukumar & Raghavan

CITES

WPA

END Exotic

LC LC LC NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE IUCN Vernacular Name Vernacular Kāri Tēṭ Nūlciṟakan Puḷḷi Tēṭ Tēṭ Karaṇṭimūkkan Tēṭ Mudra Bhīman Tēṭ Tēṭ Kaṟuppcuṭṭi Iruḷan Tēṭ Variccuṇṭanmuṣi Cāra Variccuṇṭanmuṣi Iruṇṭaciṟakan Variccuṇṭanmuṣi Varayan Sakkarmutsyaṁ Nīḷacciṟakan Munnūlan Kuṟṟippallan Vyāḷimatsyaṁ Pacchakkaṇṇan Cheṟumūkkan Araṇamīn Valiya Araṇamīn Braṣpallan Irupuḷḷi Araṇamīn Araṇamīn Intyan Gārmān Viḷakkmatsyaṁ Viḷakkmatsyaṁ Keāmpan Malayalam Name Malayalam Imcn \qÂNndI³ tXSv ]pÅn tXSv Icïnaq¡³ tXSv ap{Z tXSv `oa³ tXSv Idp¸vNp«n tXSv Ccpf³ tXSv Nmc hcn¨pï³ apjn CcpïNndI³ hcn¨pï³apjn hcb³ hcn¨pï³apjn \of¨ndI³k¡ÀaÕyw ap¶qe³ Ipän¸Ã³ hymfnaÕyw sNdpaq¡³ ]¨¡®³ henb AcWao³ {_jv]Ã³ AcWao³ Ccp]pÅn AcWao³ C´y³ AcWao³ KmÀam³ hnf¡vaÕyw sIm¼³ hnf¡vaÕyw Authority 1822) (Hamilton, 1822 Hamilton, 1840 Bonaparte, (Bloch, 1795) Gilbert, 1906 (Thunberg, 1792) (Thunberg, 1840) (Valenciennes, 1840) (Valenciennes, 1840 Valenciennes, 1984 & Haruta, Amaoka Gibbs, 1848) (Richardson, 1873) (Day, 1904) (Brauer, Valenciennes, 1840 Valenciennes, (Rüppell, 1837) 1822) (Hamilton, 1787) (Thunburg, Schultz, 1953

(Bloch, 1794) Species Name Arius arius canius Plotosus agassizi Chlorophthalmus Saurida tumbil Diaphus garmani Arius maculatus caelata Nemapteryx dussumieri Plicofollis limbatus Plotosus trifibulatus Astronesthes Saurida undosquamis indicus Synodus Diaphus splendidus Arius subrostratus thalassina Netuma Sciades sona lineatus Plotosus binotatus Synodus Pterygoplichthys spp Pterygoplichthys Heteropneustes fossilis Heteropneustes * English Name English (sea catfishes) Ariidae 52. Family Catfish) (Hamilton's Sea Catfish Threadfin ) catfishes (eeltail Plotosidae 53. Family ) (Canine Catfish-Eel Eel-Catfish Gray catfishes) (sucker Loricariidae 54. Family Sailfin Catfish Amzonian ORDER STOMIIFORMES XIX. (barbeled Stomiidae 55. Family dragonfishes) XX. ORDER AULOPIFORMES (greeneyes) Chlorophthalmidae 56. Family Shortnose Greeneye (lizardfishes) Synodontidae 57. Family Lizardfish Greater ORDER MYCTOPHIFORMES XXI. (lanternfishes) Myctophidae 58. Family Garman's Lanternfish Spotted Catfish Spotted Catfish Engraved Catfish Sea Blacktip Eel Catfish Darkfin Snaggletooth Triplethread Lizardfish Brushtooth Indian Lizardfish Horned Lanternfish Shovelnose Sea Catfish Shovelnose Catfish Giant (Dusky Catfish) Sona Catfish Sea Fish Cat Striped Eel Fish Lizard Two-Spot Stinging Catfish

287 294 297 299 300 304 288 290 292 295 298 301 303 305 289 291 293 296 302 286

8050 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

CITES

WPA

END

NE NE NE NE NE NE NE NE NE NE NE NE NE NE IUCN ‌

Vernacular Name Vernacular Tāyēāllīyāṟ Viḷakkmatsyaṁ Tāyēāllīyāṟ Viḷakkmatsyaṁ Vaṟṟasi Viḷakkmatsyaṁ Cappamūkkan Tāṭimīn Veḷḷikkaṇṇan Kēāḍ Oṟṟakkeāmpan Brēāṭṭula Āṭutāṭi Cheāṟittavaḷa Maññacchiṟakan Mīn Mīn Cheāṟittavaḷa Parappan Minusa Chūṇṭakkāran Chūṇṭakkāran Kaṟuttavāyan Tavaḷamīn Peāṭṭuciṟakan Tavaḷamīn Varayan Kaimīn Intyan Kallan Talayil Vakkan, Veḷḷi

Malayalam Name Malayalam XntbmÃnbÀ hnf¡vaÕyw häkn hnf¡vaÕyw N¸aq¡³ hnf¡vaÕyw shÅn¡®³ XmSnao³ Häs¡m¼³ tImUv- BSpXmSn t{_m«pe aª¨ndI³ sNmdn¯hf ao³ ]c¸³ sNmdn¯hf ao³ an\pk Nqï¡mc³ Idp¯hmb³ Nqï¡mc³ s]m«pNndI³ Xhfao³ hcb³ Xhfao³ C´y³ ssIao³ shÅn h¡³, XebnÂ IÃ³ &

Bineesh

2012

Authority 1840 Thompson, 1894) (Alcock, 1910 Annandale & Jenkins, Tåning, 1928 Tåning, 1794) (Shaw, Jordan & Starks, 1904 & Starks, Jordan 1877 Günther, 1846 & Schlegel, Temminck 1837) (Valenciennes, 1797) (Vahl, 1817) (Cuvier, 1835) (Valenciennes, Greenfield, Akhilesh,

Fowler, 1934 Fowler,

flavipinnis

Species Name macclellandi Bregmaceros Lophiodes mutilus indica Halieutaea Myctophum obtusirostre Myctophum striatus Antennarius Diaphus watasei japonica Polymixia Brotula multibarbata dussumieri Colletteichthys Lophiomus setigerus nummifer Antennarius duodecimalis Atherinomorus Colletteichthys

Diaphus thiollierei English Name English ORDER POLYMIXIIFORMES XXII. (beardfishes) Polymixiidae 59. Family ORDER GADIFORMES XXIII. (codlets) Bregmacerotidae 60. Family Codlet) Cod (Spotted Unicorn ORDER OPHIDIIFORMES XXIV. Ophidiidae (cuskeels) 61. Family ORDER BATRACHOIDIFORMES XXV. (toadfishes) Batrachoididae 62. Family toadfish Yellowfin XXVI. ORDER LOPHIIFORMES Lophiidae (goosefishes) 63. Family Smooth Angler (frogfishes) Antennariidae 64. Family (batfishes) Ogcocephalidae 65. Family Handfish) Indian Handfish (Starry XXVII. ORDER ATHERINIFORMES (silversides) Atherinidae 66. Family XXVIII. ORDER CYPRINODONTIFORMES Aplocheilidae (panchax) 67. Family Bluntsnout Lanternfish Bluntsnout Frogfish Striated Watases Lanternfish Watases Eye Silver Brotula Goatsbeard Toadfish Flat Blackmouth Angler (Blackmouth Goosefish) Spotfin Frogfish Silverside Tropical Thiolliere's Lanternfish Thiolliere's

310 312 314 318 308 317 307 309 311 313 315 316 319 306

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8051 Fishes of Kerala Bijukumar & Raghavan

CITES

WPA

KL END Exotic Exotic * *

LC LC LC LC LC NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE VU LC LC IUCN

Vernacular Name Vernacular Pacca Mānattukaṇṇi Pacca Mānattukaṇṇi Keātukmatsyaṁ Sārivālan Gappy, Kēālān Kēāli Pallan Sūcimīn, Parappan Sūcimīn Paṭṭa Sūcimīn Puḷḷivālan Kīltāṭi Sūcimīn Aracchuṇṭan Karinvarayan Aracchuṇṭan Luṭṭke Ḍus'sumīr Araccuṇṭan Aracchuṇṭan Kēāṅṅāṭṭuṟi Aṟṟaccuvappan Hūgli Araccuṇṭan Aracchuṇṭan Nīḷakkeākkan Malabār Aracchuṇṭan Paṟavamīn Karim Mīśa Paṟavamīn Paṟavamīn Iraṭṭacciṟakan Paṟavamīn Kēāṟamānṭel Ellan Paṟavamīn Malabār Nelmīn Malayalam Name Malayalam ]¨ am\¯pI®n am\¯pI®n sImXpIvaÕyw K¸n, kmcnhme³ tImem³ ]c¸³ kqNnao³, ]Ã³ tImen ]« kqNnao³ ]pÅnhme³ kqNnao³ IoÂXmSn kqNnao³ Icnwhcb³ Ac¨pï³ ep«vsI Ac¨pï³ UpÊpaoÀ Ac¨pï³ tIm§m«pdn Ac¨pï³ Aä¨ph¸³ lq¥n Ac¨pï³ \ofs¡m¡³ Ac¨pï³ ae_mÀ Ac¨pï³ Icnw]dhao³ aoi ]dhao³ Cc«¨ndI³ ]dhao³ tImdamïÂ ]dhao³ FÃ³ ]dhao³ ae_mÀ s\Âao³ Authority 1847) (Valenciennes, 1940) (Kulkarni, (Valenciennes, 1847) (Valenciennes, 1976 Collette, 1758 Linnaeus, (Valenciennes, 1846) (Valenciennes, 1822) (Hamilton, 1850) (Bleeker, 1850) (Bleeker, 1775) (Forsskål, 1847) (Valenciennes, 1847) (Valenciennes, 1847) (Valenciennes, 1846 Richardson, (Hornell, 1923) (Valenciennes, 1846) (Valenciennes, 1823) Hasselt, (van 1847 Valenciennes, 1852) (Bleeker,

(Baird & Girard 1853) & Girard (Baird 1859 Peters (Blyth, 1858) (Arnold, 1911) Species Name Cheilopogon cyanopterus Oryzias setnai Hyporhamphus limbatus malabaricus Rhynchorhamphus volitans Exocoetus Aplocheilus lineatus cancila Xenentodon Strongylura leiura acus Tylosurus Hemiramphus far Hyporhamphus dussumieri Hyporhamphus xanthopterus georgii Rhynchorhamphus monocirrhus Exocoetus coromandelensis Hirundichthys Ablennes hians Strongylura strongylura Hemiramphus lutkei oxycephalus Hirundichthys Gambusia affinis reticulata Poecilia Zenarchopterus striga Zenarchopterus Aplocheilus blockii * * English Name English fish) (mosquito Poeciliidae 68. Family ORDER BELONIFORMES XXIX. Belonidae (needlefishes) 69. Family (halfbeaks) Hemiramphidae 70. Family (flyingfishes) Exocoetidae 71. Family Flyingfish Margined (ricefishes) Adrianichthyidae 72. Family Indian Ricefish) (Miniature Malabar Ricefish ORDER BERICYFORMES XXX. Congaturi Halfbeak Congaturi Malabar Halfbeak Flyingfish Two-Wing Tropical Striped Panchax Fish Mosquito Needlefish Banded Needlefish Needlefish³⁴ Keel-Jawed Halfbeak Blackbarred Dussumier's Halfbeak Halfbeak Red-Tipped Long Billed Half Beak Barbel Flyingfish Flying Fish Coromandel Flat Needlefish Flat Needlefish Spottail Halfbeak Lutke's Flying Fish Bony Guppy Hoogly Halfbeak Green Panchax Green

337 342 332 336 339 321 322 324 326 328 329 331 333 335 338 340 325 327 330 341 323 334 320

8052 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

CITES App. II App. II App. II

WPA Sch. I Sch. I Sch. I Sch. I Sch. I Sch. I

END

LC LC NE NE NE NE NE NE NE NE NE NE NE NE NE NE VU VU DD IUCN

Vernacular Name Vernacular Ḍārvin Chēṟutalayan Aṇṇānmatsyaṁ Karimpuḷḷi Varayan Chuvapp Aṇṇānmatsyaṁ Pēārāḷimatsyaṁ Niḻal Cirakan Matsyaṁ, Pēārāḷi Painkāya Peruṅkaṇṇan Aṇṇānmatsyaṁ Muḷḷumūkkan Jappān Pēārāḷimatsyaṁ Ḍēāṟi Ṟēās Jēāṇ Ḍēāṟi Veḷḷi Chainīs Kuḻalmatsyaṁ Kuḻalmatsyaṁ Chuvapp Nīlappuḷḷi Kuḻalmatsyaṁ Chāl Kattimatsyaṁ Kaṭalkutira Cilka Puḷḷi Kaṭalkutira Muppuḷḷi Kaṭalkutira Paippmatsyaṁ Muttumaṇi Paippmatsyaṁ Mutalappallan Śud'dhajala Paippmatsyaṁ

Malayalam Name Malayalam UmÀhn³ tNdpXeb³ Icnw]pÅn A®m³aÕyw Nph¸v hcb³ A®m³aÕyw \ngÂ NndI³ t]mcmfnaÕyw ss]³Imb t]mcmfn aÕyw, s] cpwI®³ apÅpaq¡³ A®m³aÕyw P¸m³ t]mcmfnaÕyw tdmkv tUmdn shÅn tPm¬ tUmdn ssN\okv IpgÂaÕyw Nph¸v IpgÂaÕyw \oe¸pÅn IpgÂaÕyw Nmev I¯naÕyw NnÂ¡ ISÂIpXnc ]pÅn ISÂIpXnc ap¸pÅn ISÂIpXnc ap¯paWn ss]¸vaÕyw apXe¸Ã³ ss]¸vaÕyw ip²Pe ss]¸vaÕyw Authority (Linnaeus, 1766) 1803 Lacepède, (Forsskal, 1775) (Forsskal, (Forsskål, 1775) (Forsskål, 1829) (Cuvier, (Johnson, 1866) 1858) (Bleeker, 1853 Bleeker, & Yamakawa, Randall, Shimizu 1982 1843) (Lowe, 1852) (Lowe, Rüppell, 1838 1758 Linnaeus, Rüppell, 1838 1822) (Hamilton, (Cantor, 1849) (Cantor, 1822) (Hamilton,

Bleeker, 1852 Bleeker, Leach, 1814 Leach, Species Name chinensis Aulostomus petimba Fistularia Sargocentron rubrum Sargocentron Myripristis murdjan Myripristis japonicus Ostichthys Gephyroberyx darwinii Gephyroberyx melanospilos Sargocentron adjustus Myripristis acanthorhinus Ostichthys rosea Cyttopsis conchifer Zenopsis commersonii Fistularia scutatus Centriscus fuscus Hippocampus Microphis cuncalus Hippichthys penicillus Hippichthys carce Ichthyocampus

Hippocampus kuda Hippocampus Hippocampus trimaculatus Hippocampus English Name English (slimeheads) Trachichthyidae 73. Family (squirrelfish, Holocentridae 74. Family soldierfish) ORDER ZEIFORMES XXXI. (parazen) Parazenidae 75. Family (dories) Zeidae 76. Family ORDER SYNGNATHIFORMES XXXII. (trumpetfishes) Aulostomidae 77. Family Fish Trumpet Chinese (cornetfishes) Fistulariidae 78. Family Fish Cornet Red (razorfish) Centriscidae 79. Family and (pipefishes Syngnathidae 80. Family seahorses) Redcoat (Red Striped Squirrelfish) (Red Redcoat Pinecone Soldier Fish Pinecone Perch) Japanese Soldier Fish (Brocade Darwin's Slimehead Fish Black Spot Squirrel Shadowfin Soldier Fish Fish Spinesnout Squirrel Dory Rosy John DorySilvery Fish Cornet Blue-spotted Shrimpfish Grooved Seahorse) (Chilka Sea Pony Pipe Fish Tooth Crocodile Beady Pipefish Beady ¹ Sudhajala Pipe Matsyam Spotted Seahorse (Yellow Seahorse) (Yellow Seahorse Spotted Longnose Seahorse (Three-spot Seahorse) (Three-spot Longnose Seahorse

352 353 345 347 349 343 344 346 348 350 351 354 355 356 360 359 361 357 358

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8053 Fishes of Kerala Bijukumar & Raghavan

CITES

WPA Sch. I Sch. I Sch. I Sch. I

KL KL KL KL KL WG END

LC LC LC LC NE NE NE NE NE EN NE NE NE NE NE NE NE NE NE DD DD DD DD IUCN

Vernacular Name Vernacular Chīṅkaṇṇi Paippmatsyaṁ Chīṅkaṇṇi Paippmatsyaṁ Irutala Paippmatsyaṁ Vaṭi Paippmatsyaṁ Arivāḷ Pātaḷateāṇṭi Kaṭṭapuḷavan Kuḻipuḷavan Ceṅkalpuḷavan Madhurān Malabār Muḷḷārakan Malayārakan Panayārakan Tēḷmatsyaṁ Āḻakkaṭal Tēḷmatsyaṁ Aṟakkavālceviyan Muḷḷmatsyaṁ Chariññavarayan Tēḷmatsyaṁ Ālkēākk Muḷḷmatsyaṁ Cāra Tīmatsyaṁ Kariṅkālan Tēḷmatsyaṁ Vītivarayan Ṭarkimatsyaṁ Teḷivālan Tēḷmatsyaṁ Chuvapp Tēḷmatsyaṁ Guvāṁ Tēḷmatsyaṁ Kaḷa

Malayalam Name Malayalam No¦®n ss]¸vaÕyw CcpXe ss]¸vaÕyw hSn ss]¸vaÕyw AcnhmÄ ss]¸vaÕyw ]mXfsXmïn I«]pfh³ Ipgn]pfh³ sN¦Â]pfh³ a[pcm³ ae_mÀ apÅmcI³ aebmcI³ ]\bmcI³ Bg¡SÂ tXÄaÕyw Ad¡hmÂsNhnb³ tXÄaÕyw Ncnªhcb³ apÅvaÕyw BÂtIm¡v tXÄaÕyw Nmc apÅvaÕyw IcnwIme³ XoaÕyw hoXnhcb³ tXÄaÕyw sXfnhme³ SÀInaÕyw Nph¸v tXÄaÕyw Kphmw tXÄaÕyw If tXÄaÕyw Authority 1862 Johnson, 1846) (Richardson, Temminck & Schlegel, 1850) & Schlegel, Temminck (Bleeker, 1857) (Bleeker, 1896) (Alcock, 1831) (Bennett, 1824) & Gaimard, (Quoy Kaup, 1856 Kaup, Gopi, 2002 1951) (Nayar, McClelland, 1844 1865) (Day, 1849) (Jerdon Hallacher & Rama- Eschmeyer, 1979 Rao, 1801) (Bloch & Schneider, (Bloch, 1787) (Linnaeus, 1758) 1793) (Thunnberg, Talwar, 1991 Talwar,

(Lacepède, 1800) 1889 Alcock, (Bloch, 1785) Bailey & Gans, 1998 & Gans, Bailey Species Name guentheri Setarches serrulata Brachypterois Trachyrhamphus serratus Trachyrhamphus Trachyrhamphus bicoarctatus Trachyrhamphus macrura Parapterois russelli Pterois guamensis Scorpaenodes Trachyrhamphus longirostris Trachyrhamphus digressus Monopterus fossorius Monopterus bengalense Ophisternon Macrognathus guentheri malabaricus Mastacembelus Minous dempsterae Minous monodactylus antennata Pterois volitans Pterois cirrhosa Scorpaenopsis Monopterus eapeni Monopterus

Mastacembelus armatus Mastacembelus Minous inermis Syngnathoides biaculeatus Syngnathoides Monopterus roseni Monopterus English Name English ORDER SYNBRANCHIFORMES XXXIII. eel) (swamp Synbranchidae 81. Family eels) (spiny Mastacembelidae 82. Family ORDER SCORPAENIFORMES XXXIV. bristly (deep-sea Setarchidae 83. Family scorpionfishes) (Deepwater Channeled Rockfish Scorpionfish) or (scorpionfishes Scorpaenidae 84. Family rockfishes) Scorpionfish Sawcheek Saw Pipe Fish Saw Double-ended Pipe Fish (Bentstick Pipefish) Double-ended Pipe Fish (Bentstick Firefish Blackfoot Fish Turkey Plain Tail Guam Scorpionfish Straightstick Pipe Fish Straightstick Eel Swamp Eel Malabar Swamp Eel Swamp Bengal Eel Malabar Spiny Eel Malabar Tyre-Track Obliquebanded Stingfish Stingfish) Goblin Fish (Grey Grey Firefish Broadbarred Fish) Fire (Winged Lionfish Red Stingfish Weedy Eapen's Swamp Eel³⁵ Swamp Eapen's Zig-zag Eel (Tyre-track Eel) (Tyre-track Eel Zig-zag Fish Scorpion Alcock's Alligator Pipe Fish Alligator Rosen's Swamp Eel Swamp Rosen's

374 375 365 363 379 381 383 364 366 368 370 371 373 376 378 380 382 384 367 372 377 362 369

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‌ ‌

‌ Vernacular Name Vernacular Kal Matsyaṁ Kal Kaṭannalmatsyaṁ Kaṇṇan Veḷḷamukhan Kaṭannalmatsyaṁ Muḷḷmatsyaṁ Varayan Ōṟañc Pirivālan Ēden Paṟakkuṁ Puḷḷicciṟakan Gurnārṭ Gurnārṭ Paṟakkuṁ Perastya Gurnārṭ Paṟakkuṁ Nakṣatra Gurnārṭ Cetumpalmāṟan Ṟēābin Kaṭal Gurnārṭ Karimpuḷḷi Ṟēābin Kaṭal Kavacita Cappattalayan Mutala Cappattalayan Parukkan Jappān Cappattalayan Muḷḷan Cappattalayan Cappattalayan Varavālan Cappattalayan Muḻayan Glāsṁ Keāmēḻsaṇ Malabār Glāsṁ

Malayalam Name Malayalam IÂ aÕyw I®³ IS¶ÂaÕyw shÅapJ³ IS¶ÂaÕyw Hmd©v hcb³ apÅvaÕyw GsZ³ ]ncnhme³ ]pÅn¨ndI³ ]d¡pw KpÀ\mÀSv- s]uckvXy ]d¡pw KpÀ\mÀSv- \£{X ]d¡pw KpÀ\mÀSv- sNXp¼Âamd³ KpÀ\mÀSv ISÂ tdm_n³ Icnw]pÅn KpÀ\mÀSv IhNnX ISÂ tdm_n³ apXe N¸¯eb³ ]cp¡³ N¸¯eb³ P¸m³ N¸¯eb³ apÅ³ N¸¯eb³ hchme³ N¸¯eb³ apgb³ N¸¯eb³ sImtagvk¬ ¥mkv-aÕyw ae_mÀ ¥mkv-aÕyw Authority 1801) Bloch & Schneider, 1848) (Richardson, 1848 Richardson, (Cuvier, 1829) (Cuvier, Alcock, 1890 Alcock, (Linnaeus, 1758) 1829) (Cuvier, 1828 Cuvier, Kotthaus, 1970 Kotthaus, 1854) (Bleeker, 1887) (Nyström, 1914 & Thompson, Gilchrist 1907) (Lloyd, (Tilesius, 1812) (Tilesius, 1812) (Linnaeus, 1758) (Lacepède, 1802) (Boulenger, 1888) (Boulenger,

(Cuvier, 1829) (Cuvier, Bloch & Schneider, 1801 Bloch & Schneider, arabica arabica

Species Name carinatus Apistus leucogaster Richardsonichthys Choridactylus multibarbus Dactyloptena orientalis Dactyloptena Lepidotrigla longipinnis scaber Grammoplites rodericensis Kumococius Ambassis dussumieri Synagrops adeni Synagrops macracantha Dactyloptena peterseni Dactyloptena Lepidotrigla faurei adeni Satyrichthys Cociella crocodilus Inegocia japonica indicus Platycephalus Ambassis ambassis Pterygotrigla

Sorsogona tuberculata Synanceia verrucosa Synanceia English Name English scorpionfishes) (wasp Apistidae 85. Family Waspfish Ocellated (waspfishes) Tetrarogidae 86. Family Waspfish Whiteface (stonefishes) Synanceiidae 87. Family Stingfish Orangebanded (flying gurnards) Dactylopteridae 88. Family (searobins) Triglidae 89. Family searobins (armored Peristediidae 90. Family gurnards) or armored (flatheads) Platycephalidae 91. Family ORDER PERCIFORMES XXXV. glassfishes/ (asiatic Ambassidae 92. Family perchlets) Oriental Flying Gurnard Oriental Sea Robin Flathead Rough Flathead Spiny Perchlet¹ Malabar Glassy Aden Splitfin Spotwing Flying Gurnard Flying Gurnard Starry Gurnard Scalybreast Gurnard Blackspotted Robin Sea Armoured Flathead) (Spotted Flathead Crocodile Japanese Flathead Flathead Bartail Perchlet¹ Glassy Commerson's Tuberculated Flathead Tuberculated Stonefish

386 387 388 391 394 398 400 404 389 390 392 393 395 396 397 399 401 403 402 385

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Vernacular Name Vernacular Kaṣaṇṭi Glāsṁ Kaṣaṇṭi Nandan Nīṇṭamuḷḷan Nandan Arikuñeāṟi Nandan Ḍē Glāsṁ Ceṟunandan Āṟṟunandan Kāḷāñci Kalava Mayil Kalava Svarṇa Kalava Cēākkalēṟṟ Kalava Nīlavarayan Puḷḷippuli Kalava Paviḻakkalava Āṟupuḷḷi Kalava Cheṅkalava Kalava Ciṟakan Iruṇṭa Kalava Intyan Kalava Kūnan Kaṇikkalava Kalava Cāravālan Kalava Veḷḷapuḷḷi Mīśakkalava Kalava Taviṭṭupuḷḷi Kalava Muḷkaviḷan Puḷḷikkalava Kalava Aṟṟakkaṟuppan Maññakkalava

Malayalam Name Malayalam Ijïn ¥mkv-aÕyw \oïapÅ³ \µ³ AcnIpsRmdn \µ³ \µ³ tU ¥mkv-aÕyw sNdp\µ³ Bäp\µ³ Imfm©n abnÂ Ieh kzÀW Ieh tNm¡teäv Ieh \oehcb³ Ieh ]pÅn¸pen Ieh ]hng¡eh Bdp]pÅn Ieh sNwIeh Ccpï NndI³ Ieh C´y³ Ieh Iq\³ Ieh IWn¡eh Nmchme³ Ieh shÅ]pÅn Ieh aoi¡eh Xhn«p]pÅn Ieh apÄIhnf³ Ieh ]pÅn¡eh Aä¡dp¸³ Ieh aª¡eh Authority Bleeker, 1853 Bleeker, 1822 Hamilton, 1822) (Hamilton, 1828) (Valenciennes, 1812) (Shaw, 1775) (Forsskål, 1828) (Valenciennes, Bineesh & Akhilesh, 2013 1775) (Forsskål, (Bloch, 1790) 1842) & Schlegel, (Temminck (Lacepède, 1802) (Hamilton, 1822) (Hamilton, 1874) (Bleeker, 1870) (Day, (Bloch, 1790) 1801 Bloch & Schneider, (Bloch, 1790) (Lacepède, 1801) (Rüppell, 1830) 1801) (Forster, 1828) (Valenciennes, 1878) (Vaillant, 1861) (Castelnau, 1828) (Valenciennes, 1775) (Forsskål, (Valenciennes, 1828) (Valenciennes,

(Lacepède, 1802) Species Name Ambassis interrupta Chanda nama ranga Parambassis Cephalopholis aurantia Cephalopholis formosa Cephalopholis miniata Cephalopholis sonnerati maculicauda Chelidoperca areolatus Epinephelus caeruleopunctatus Epinephelus epistictus Epinephelus flavocaeruleus Epinephelus Ambassis nalua dayi Parambassis thomassi Parambassis calcarifer Lates Cephalopholis argus Cephalopholis boenack Cephalopholis leopardus Cephalopholis sexmaculata Cephalopholis urodeta altivelis Cromileptes bleekeri Epinephelus chabaudi Epinephelus diacanthus Epinephelus fasciatus Epinephelus Epinephelus chlorostigma Epinephelus

Ambassis gymnocephalus English Name English perches) (lates Latidae 93. Family groupers, basses, (sea Serranidae 94. Family basslets) fairy Long Spined Glass Perchlet Perchlet Glassy Elongate Fish Indian Glassy Cod) Golden Hind (Golden Rock Blue-Lined Hind (Blue-Lined Rockcod) Hind Coral Hind Tomato Indian Perchlet Grouper Areolate Cod Reef Spotted White Grouper Dotted Grouper Blue and Yellow Scalloped Perchlet Scalloped Perchlet¹ Glassy Day's Perchlet Glassy Ghats Western Seaperch) (Giant Barramundi Bluespotted Grouper, Hind (Peacock Peacock Grouper) Hind Chocolate Hind Leopard Hind Sixblotch Darkfin Hind Humpback Grouper Grouper Duskytail Grouper Moustache Grouper Spinycheek Grouper Blacktip Brownspotted Grouper Brownspotted Bald Glassy (Naked- Head Glass Perchelet)¹ Head (Naked- Bald Glassy

406 408 410 414 416 418 420 422 424 426 430 432 407 409 411 412 413 415 417 419 421 423 425 427 429 431 428 405

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jīni Paviḻakkalava ̄ Vernacular Name Vernacular Taviṭṭu Mārbiḷ Kalava Taviṭṭu Kalava Varayan Kalava Nīṇṭamuḷḷan Puḷḷikkalava Malabār Kalava Kalava Tēnkūṭ Kalava Chariññavarayan Pannikkalava Kalava Vaḷaññavarayan Sēāppmīn Āṟuvarayan Chandrakkalava Ṟānḍal Kalava Kalava Nār Kalava Ālkēākk Karin Paviḻakkalava Peāṭṭu Kalava Oṟṟavarayan Kalava Mārsiyā Kalava Balengar Kiḷi Tāṭimatsyaṁ Puḷḷippuli Tāṭimatsyaṁ Kīrimīn Naluvarayan Kīrimīn Chandrakkala KīrimīnCheṟucetumpal Kīrimīn Valiyachetumpal

Malayalam Name Malayalam Xhn«p amÀ_nÄ Ieh hcb³ Ieh \oïapÅ³ Ieh ]pÅn¡eh ae_mÀ Ieh tX³IqSv Ieh Ncnªhcb³ Ieh ]¶n¡eh hfªhcb³ Ieh Bdphcb³ tkm¸vao³ N{µ¡eh dm³UÂ Ieh \mcv Ieh BÂtIm¡v Ieh IcnwPo\n ]hng¡eh s]m«p ]hng¡eh Hähcb³ Ieh amÀknbm Ieh _use³KÀ Ieh Infn XmSnaÕyw ]pÅn¸pen XmSnaÕyw \oehcb³ Iocnao³ N{µ¡e Iocnao³ sNdpsNXp¼Â Iocnao³ henbsNXp¼Â Iocnao³ Authority (Bloch, 1790) Smith-Vaniz, Bineesh & Smith-Vaniz, 2012 Akhilesh, (Temminck & Schlegel, 1842) & Schlegel, (Temminck (Bloch, 1790) (Bloch, 1793) 1775) (Forskal, Bineesh & White, Akhilesh, 2012 Bineesh, Akhilesh, Gopalakrishnan 2014 & Jena, 1993 Randall & Hoover, 1829 Cuvier, (Kner, 1864) (Kner, 1801) Bloch & Schneider, 1868) (Day, 1824) & Gaimard, (Quoy 1863) (Guichenot, 2008 Randall & Heemstra, (Lacepede, 1801) 1955) (Kamohara, 1973) (Heemstra, Rüppell, 1830 1775) (Forsskal, 1829 Cuvier, (Thunberg, 1792) (Thunberg, (Bloch, 1790)

(Forsskål, 1775) (Forsskål, Species Name quadrilineatus Pelates Opistognathus pardus Opistognathus Epinephelus latifasciatus Epinephelus maculatus Epinephelus merra Epinephelus tauvina Epinephelus Liopropoma randalli alcocki Plectranthias marcia Pseudanthias puta Terapon Epinephelus longispinis Epinephelus malabaricus Epinephelus radiatus Epinephelus undulosus Epinephelus Liopropoma lunulatum filiferus Meganthias Plectropomus laevis fasciatus Pseudanthias Sacura boulengeri nigromarginatus Opisthognathus jarbua Terapon theraps Terapon Grammistes sexlineatus Grammistes Plectropomus maculatus

Epinephelus fuscoguttatus Epinephelus English Name English (jawfishes) Opistognathidae 95. Family or (grunters Terapontidae 96. Family tigerperches) Terapon Fourlined or (bigeyes Priacanthidae 97. Family catalufas) Leopard Jawfish Leopard Striped Grouper Rockcod Spotted Reef- (Wire-Netting Grouper Honeycomb Cod) Grouper Greasy Randall's Basslet Basslet Deep-reef Alcock’s Anthias Marcia's Terapon Small-Scaled Longspine Grouper Malabar Grouper Oblique-Banded Grouper Grouper Wavy-Lined Basslet Anthiine Filamentous Coralgrouper Blacksaddled Anthias One-stripe Anthias Boulenger's Jawfish Birdled Squeaking (Jarbua Terapon, Perch Crescent )¹ Perch Terapon Largescaled Sixstripe Soapfish Sixstripe Coralgrouper Spotted Brown-Marbled Grouper Brown-Marbled

454 453 434 436 438 440 444 446 450 456 435 437 439 441 443 445 447 449 451 452 455 457 442 448 433

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NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE IUCN Vernacular Name Vernacular Lās Kaṇṇan Kāḷakkaṇṇan Chandravālan Piṅkpuḷaḷi Kāḷakkaṇṇan Karddināḷmatsyaṁ Vaḷayavālan Cheṟuchetumpal Karddināḷmatsyaṁ Muttuciṟakan Karddināḷmatsyaṁ Puḷaḷicciṟakan Karddināḷmatsyaṁ Karddināḷmatsyaṁ Ēḻupaṭṭa Karddināḷmatsyaṁ Irubelṟṟ Karddināḷmatsyaṁ Irupaṭṭa Matsyaṁ Ruvarayankarddināḷ Varayan Ōṟañc Karddināḷmatsyaṁ Minnuṁ Karddināḷmatsyaṁ Karddināḷmatsyaṁ Varayan Vīti Karddināḷmatsyaṁ Ēḻuvarayan Karddināḷmatsyaṁ Arappaṭṭa Pūḻān Mantukālan Pūḻān Veḷḷi Taṟayēāṭumatsyaṁ Chāyakkēāl Parava Mēāta Sakkar Srāv Īrkkil Sakkar

Malayalam Name Malayalam ¥mkv- I®³ N{µhme³ Imf¡®³ ]n¦v]pffn Imf¡®³ hfbhme³ IÀ±n\mÄaÕyw sNdpsNXp¼Â IÀ±n\mÄaÕyw ap¯pNndI³ IÀ±n\mÄaÕyw ]pffn¨ndI³ IÀ±n\mÄaÕyw Ggp]« IÀ±n\mÄaÕyw Ccps_Âäv- IÀ±n\mÄaÕyw Ccp]« IÀ±n\mÄaÕyw Ccphcb³ IÀ±n\mÄ aÕyw Hmd©zcb³ IÀ±n\mÄaÕyw an¶pw IÀ±n\mÄaÕyw hoXnhcb³ IÀ±n\mÄaÕyw Ggphcb³ IÀ±n\mÄaÕyw Ac¸« IÀ±n\mÄaÕyw a´pIme³ ]qgm³ shÅn ]qgm³ Nmbt¡mÂ XdtbmSpaÕyw ]ch tamX {kmhv k¡À CuÀ¡nÂ k¡À Authority 1887) (Günther, 1801) (Bloch & Schneider, 1766) Linnaeus, 1758 Linnaeus, (Forskal, 1775) (Forskal, 1828) (Cuvier, 1903 Gilchrist, 1914) & Thompson, (Jordan 1828) (Cuvier, 1775) (Forsskål, Richardson, 1846 Richardson, (Lacepede, 1802) 1828 Cuvier, 1880) (Gunther, (Gon, 1986) 1849) (Cantor, 1790) (White, 1829) (Cuvier, 1849) (Bleeker, 1791) (Menzies, 1828) (Ehreberg, 1828) (Ehreberg, (Lacepede, 1801) Species Name Hoplolatilus fronticinctus lactarius Lactarius canadum Rachycentron naucrates Echeneis Priacanthus hamrur Priacanthus Apogon multitaeniatus Apogon queketti sialis Apogonichthyoides novemfasciatus Ostorhinchus Sillago sihama Priacanthus tayenus Priacanthus aureus Ostorhinchus Apogon poecilopterus Apogon septemstriatus pseudotaeniatus Apogonichthyoides Archamia fucata fasciatus Ostorhinchus thermalis Ostorhinchus Sillaginopodys chondropus lineatus Phtheirichthys Apogon taeniatus Archamia lineolata

Heteropriacanthus cruentatus Heteropriacanthus

Fish

English Name English (cardinalfishes) Apogonidae 98. Family Cardinal Twobelt Shimmering Cardinal Sillaginidae (smelt-whitings) 99. Family (tilefishes) Malacanthidae 100. Family Tilefish Pastel trevallies) (false Lactariidae 101. Family ) Jumper (Big-Jawed Trevally False (cobia) Rachycentridae 102. Family Cobia (King Fish) fish, (sucker Echeneidae 103. Family remoras) Sharksucker Live Moontail Bullseye (Crescent Tail Big Eye) Tail (Crescent Bullseye Moontail Fish Cardinal Smallscale Fish ) Fish (Signal Cardinal Spotfin Cardinal Fish Cardinal Twinbar Fish Cardinal Sevenstriped Sillago Silver Purple-Spotted Bigeye Purple-Spotted Tail Cardinal (Band Cardinalfish Ring-Tailed Fish) Fish Cardinal Pearly-Finned Fish Banded Cardinal Seven Fish Doublebar Cardinal Fish Cardinal Orangelined Fish Cardinal Broadbanded Fish Cardinal Half-barred Sillago Clubfoot Slender Suckerfish Glasseye

476 477 478 479 459 462 464 466 468 470 472 475 460 461 463 465 467 469 471 473 474 480 458

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̥ dumāṟiṭa Pāra dumāṟiṭa Vernacular Name Vernacular Veḷḷa Sakkar Veḷḷa Pāra Nūlvālan Pāra Nūlvālan Intyan Chem'mīn Pāra Pāra Kattivayaṟan Pāra Karimciṟakan Mattippāra Pāra Piḷavayaṟan Pāra Maññavālan Pāra Nīḷacciṟakan Pāra Mūkkan Tīrappāra Nīlappāra Maññapuḷḷi Pāra Mr Pāra Bampmūkkan Malabār Pāra Pāra Kampikkaviḷan Pāra Taviṭṭumutukan Kapaṭappāra Ppan Pāra Aṟṟakkaṟu Bhīman Pāra Pāra Nīlacciṟakan Cempan Pāra Vaṟṟappāra Ayalakkeāḻiyāḷa Keāḻiyāḷa Cheṟuchiṟakan Keāḻiyāḷa Intyan Kkāran Mārivil Ōṭṭa Malayalam Name Malayalam shÅ k¡À \qÂhme³ ]mc Cý³ \qÂhme³ ]mc sN½o³ ]mc I¯nhbd³ ]mc IcnwNndI³ ]mc a¯n¸mc ]nfhbd³ ]mc aªhme³ ]mc \of¨ndI³ ]mc aq¡³ ]mc Xoçmc \oe¸mc aª]pÅn ]mc arZpamdnS ]mc _¼vaq¡³ ]mc ae_mÀ ]mc I¼n¡hnf³ ]mc Xhn«papXpI³ ]mc I]S¸mc Aä¡dp¸³ ]mc òa³ ]mc \oe¨ndI³ ]mc sN¼³ ]mc hä¸mc Abes¡mgnbmf sNdpNndI³ sImgnbmf Cý³ sImgnbmf amcnhnÂ Hm« ¡mc³ 1775)

Authority (Bloch, 1787) (Forsskål, 1775) (Forsskål, 1839) (Swainson, (Rüppell, 1830) (Rüppell, 1830) (Bloch, 1793) 1833) (Cuvier, 1833) (Cuvier, 1833) (Cuvier, 1775) (Forsskål, 1833) (Cuvier, 1801) (Bloch & Schneider, 1830) ([Bennett], 1830) (Bennett, 1833 Cuvier, 1825 & Gaimard, Quoy 1851 Bleeker, 1825) & Gaimard, (Quoy (Rüppell, 1830) (Rüppell, 1830) 1775) (Forsskål, 1934) (Whitley, 1857 Bleeker, 1852 Bleeker, 1877 & MacLeay, Alleyne 1833) (Cuvier, (Forsskal (Bloch & Schneider, 1801) (Bloch & Schneider, (Temminck & Schlegel, 1850) & Schlegel, (Temminck Species Name Alectis ciliaris Alepes djedaba Alepes melanoptera Alepes russelli Decapterus Alectis indicus kleinii Alepes vari Alepes mate Atule Carangoides chrysophrys Carangoides ferdau Carangoides gymnostethus Carangoides malabaricus Carangoides praeustus heberi Caranx melampygus Caranx sexfasciatus Caranx macrosoma Decapterus Elagatis bipinnulata Carangoides armatus Carangoides coeruleopinnatus Carangoides fulvoguttatus Carangoides hedlandensis Carangoides plagiotaenia Carangoides talamparoides ignobilis Caranx papuensis Caranx macarellus Decapterus Atropus atropos Atropus Remora albescens Remora English Name English (jacks, Carangidae king fishes) 104. Family Trevally) (Indian Threadfin Pompano African Shrimp Scad Blackfin Scad Indian Scad Indian Threadfish Scad Razorbelly Herring Scad Scad Yellowtail Longnose Trevally Blue Trevally Trevally) (Nakedbreast Bludger Malabar Trevally Trevally Brown-Backed Trevally Blacktip Trevally Bluefin Jack Tella Shortfin Scad Runner Rainbow Longfin Trevally Longfin Trevally Coastal Trevally Yellowspotted Bumpnose Trevally Trevally Barcheek Trevally Impostor Jack) (Yellowfin Trevally Giant Trevally Brassy Scad Mackerel Cleftbelly Trevally Cleftbelly White Suckerfish White 482 484 486 508 483 485 487 489 491 493 495 497 499 501 503 505 507 509 490 492 494 496 498 500 502 504 506 488 481

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Vernacular Name Vernacular Svarṇa Pallillāppāra Svarṇa Vaṅkaṭa Matsyaṁ Payalaṟṟ Āvēāli Kaṟutta Mīn Pāla Rājñimatsyaṁ, Talāṅg Irupuḷḷi Rājñimatsyaṁ Rājñimatsyaṁ Sūciccetumpal Keāḻiyāḷa Kaṇṇan Keāḻiyāḷa Valiyakaṇṇan Pāra Karinvayaṟan Añcupuḷḷi Tālippāra Tālippāra Patimūkkan Tālippāra Intyan Nīḷappāḷa Pāra Pāra Veḷḷavayaṟan Pāra Paruttivāyan Ḍēāḷphin Peāmpānēā Matsyaṁ Ḍēāḷphin Matsyaṁ Nāṭan Pāra Ampaṭṭan Muḷḷankāra Varayan Īrkkil Muḷḷankāra Muḷḷankāra Chāṭṭacchiṟakan Muḷḷankāra Alaṅkāra Muḷḷankāra Tiḷakka Ceṟupallan Muḷḷankāra Muḷḷankāra Pallan Muḷḷankāra Svarṇavarayan Malayalam Name Malayalam kzÀW ]ÃnÃm¸mc h¦S ]beäv aÕyw Idp¯ Bthmen XemwKv cmÚnaÕyw, ]me ao³ Ccp]pÅn cmÚnaÕyw kqNns¨Xp¼Â cmÚnaÕyw I®³ sImgnbmf henbI®³ sImgnbmf Icnwhbd³ ]mc A©p]pÅn Xmen¸mc ]Xnaq¡³ Xmen¸mc C´y³ Xmen¸mc \of¸mf ]mc shÅhbd³ ]mc ]cp¯nhmb³ ]mc s]m¼mt\m tUmÄ^n³ aÕyw \mS³ tUmÄ^n³ aÕyw A¼«³ ]mc hcb³ apÅ³Imc CuÀ¡nÂ apÅ³Imc Nm«¨ndI³ apÅ³Imc Ae¦mc apÅ³Imc Xnf¡ apÅ³Imc sNdp]Ã³ apÅ³Imc ]Ã³ apÅ³Imc kzÀWhcb³ apÅ³Imc Authority 1801) (Bloch & Schneider, (Lacepède, 1803) (Cuvier, 1829) (Cuvier, (Linnaeus, 1758) (Bloch, 1795) 1775) (Forsskål, 1833) (Cuvier, (Rüppell, 1829) (Lacepède, 1801) 1833) (Cuvier, 1860) (Poey, 1758 Linnaeus, 1874) (Günther, 1835) (Valenciennes, 1917 & Starks, Jordan 1829) (Cuvier, (Linnaeus, 1758) 1801 Lacepède, 1832) (Cuvier, (Bloch, 1793) (Lacepède, 1801) 1832 Cuvier, 1801) (Forster, 1758 Linnaeus, 1860) (Günther, (Bloch, 1795)

(Forsskål, 1775) (Forsskål, Species Name maculata Mene Aurigequula fasciata Eubleekeria splendens Eubleekeria Megalaspis cordyla niger Parastromateus lysan Scomberoides Selar boops Seriolina nigrofasciata blochii Trachinotus Ulua mentalis Uraspis secunda Coryphaena equiselis elongatus Equulites lineolatus Equulites achlamys Gazza daura Karalla Naucrates doctor Naucrates commersonnianus Scomberoides tol Scomberoides Selar crumenophthalmus baillonii Trachinotus mookalee Trachinotus Uraspis helvola Coryphaena hippurus leuciscus Equulites minuta Gazza

Gnathanodon speciosus English Name English Coryphaenidae (dolphinfishes) 105. Family Menidae (moonfish) 106. Family Moonfish (slimys, Leiognathidae 107. Family slipmouths, ponyfishes) Striped Ponyfish Splendid Ponyfish Torpedo Scad Torpedo Black Pomfret Queenfish Double-Spotted Scad Oxeye Trevally Blackbanded Snubnose Pompano Trevally Longrakered Jack Cottonmouth Dolphinfish Pompano Slender Ponyfish Ponyfish Ornate Ponyfish Smalltoothed Ponyfish Goldstripe Pilotfish Paala Meen Queenfish, Talang Queenfish Needlescaled Scad Bigeye Dart) Dart (Baillon’s Small Spotted Indian Pompano Jack Whitemouth Common Dolphinfish Ponyfish Whipfin Toothpony Golden Toothless Trevally Golden Toothless

528 529 533 511 513 515 517 519 521 523 525 526 530 532 534 536 512 514 516 518 520 522 524 527 531 535 510

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cuvapp Chempalli cuvapp ̄ Vernacular Name Vernacular Ḍus'sumīr Muḷḷankāra Muḷḷankāra Berber Muḷḷankāra Cirakan Ōṟañc Muḷḷankāra Ceṟu Mūkkan Muḷḷankāra Nāṭan Irupuḷḷi Muḷḷankāra Muḷḷankāra Patimūkkan Muḷḷankāra Patimūkkan Valiya Ceṟupallan Chempalli Chempalli Āḻakkaṭal Chempalli Ṟṟāṅ Chempalli Kaṇṭal Chempalli Baṅgāḷ Irupuḷḷi Chempalli Chempalli Caturaṅga Chempalli Karimpuḷḷi Kaṭun Chempalli Karinvālan Chempalli Kūnan Chempalli Mēāsas Mañña Chempalli Chempalli Peruṅkaṇṇan Chempalli Intyan Malabār Chempalli Orupuḷḷi Chempalli Chempalli Añcuvarayan Chempalli Keāḻuppcuṇṭan Ṟas'sal Chempalli Chempalli Chakravartti

Malayalam Name Malayalam UpÊpaoÀ apÅ³Imc s_Às_À apÅ³Imc Hmd©vNndI³ apÅ³Imc sNdpaq¡³ apÅ³Imc \mS³ apÅ³Imc Ccp]pÅn apÅ³Imc ]Xnaq¡³ apÅ³Imc henb ]Xnaq¡³ apÅ³Imc sNdp]Ã³ sNw]Ãn Bg¡SÂ sN¼Ãn ämMv sN¼Ãn IïÂ sN¼Ãn _wKmÄ sN¼Ãn Ccp]pÅn sN¼Ãn NXpcwK sN¼Ãn Icnw]pÅn sN¼Ãn ISpwNph¸v sN¼Ãn Icnwhme³ sN¼Ãn Iq\³ sN¼Ãn tamkkv sN¼Ãn aª sN¼Ãn s]cpwI®³ sN¼Ãn C´y³ sN¼Ãn ae_mÀ sN¼Ãn Hcp]pÅn sN¼Ãn A©phcb³ sN¼Ãn sImgp¸vNpï³ sN¼Ãn dÊÂ sN¼Ãn N{IhÀ¯n sN¼Ãn

Authority (Valenciennes, 1835) (Valenciennes, 1775) (Forsskål, 1775) (Forsskål, 1775) (Forsskål, 1801) (Schneider, (Bloch, 1792) Bloch, 1790 1801) (Bloch & Schneider, (Bloch, 1790) 1849) (Bleeker, (Valenciennes, 1835) (Valenciennes, 1835) (Valenciennes, 1835) (Valenciennes, (Bloch, 1787) 1822) (Hamilton, (Lacepède, 1801) 1828 Cuvier, (Chan, 1970) (Bloch, 1790) 1828) (Cuvier, Bloch, 1790 1775) (Forsskål, 1775) (Forsskål, 1831) (Valenciennes, 1828) (Cuvier, 1828) (Cuvier, 1816) (Cuvier,

(Peters, 1869) (Peters, (Valenciennes, 1835) (Valenciennes, Species Name Leiognathus bindus Leiognathus equulus Lutjanus argentimaculatus Lutjanus bohar Lutjanus fulvus Lutjanus johnii Lutjanus lutjanus Lutjanus malabaricus Lutjanus quinquelineatus Lutjanus russellii Leiognathus berbis Leiognathus brevirostris Nuchequula blochii insidiator Secutor ruconius Secutor furcatus Aphareus carbunculus Etelis Lipocheilus carnolabrum Lutjanus bengalensis Lutjanus decussatus Lutjanus erythropterus Lutjanus gibbus Lutjanus kasmira Lutjanus madras Lutjanus monostigma Lutjanus rivulatus Lutjanus sebae

Lutjanus ehrenbergii Karalla dussumieri Karalla English Name English Lutjanidae (snappers) 108. Family Orangefin Ponyfish Ponyfish Orangefin Common Ponyfish ) Snapper Snapper (River Red Mangrove Snapper Red Two-Spot Banded Snapper) (Yellow Snapper Blacktail Moses Perch Snapper Bigeye Malabar Blood Snapper Snapper Five-Lined Russell's Snapper Berber Ponyfish Berber Shortnose Ponyfish Ponyfish Twoblotch Pugnose Ponyfish Ponyfish Deep Pugnose Jobfish Smalltooth Snapper Red Deep-Water Snapper Tang's Snapper Bengal Snapper Checkered Crimson Snapper Snapper Humpback Red Snapper Blue and Yellow Indian Snapper Onespot Snapper Blubberlip Snapper Snapper Emperor Blackspot Snapper Blackspot Dussumier's Ponyfish

539 541 548 550 554 556 558 560 562 564 538 540 542 543 544 545 546 547 549 551 553 555 557 559 561 563 565 552 537

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cuvapp Jēāb cuvapp ̄ ̥ ḍamuḷḷan Prāññil Vernacular Name Vernacular Taviṭṭuvarayan Chempalli Taviṭṭuvarayan Chempalli Veḷḷakkaṟupp Chempalli Pātirā Nīlachempalli Maññavālan Chempalli Piñcāleā Kaṭun Chempalli Svarṇavarayan Svarṇavarayan Tuppākkimatsyaṁ Phusilīr Varṇappuḷḷi Mūvālan Prāññil Pokka Prāññil Keāṭiyan Jīni Prāññil Dr Prāññil Nīḷakkiraṇa Īrkkil Prāññil Prāññil Nīḷacchiṟakan Mukkaṟamatsyaṁ Varṇṇa Veḷḷi Nīlakkaviḷ Mukkaṟamatsyaṁ Mukkaṟamatsyaṁ Veḷḷi Puḷḷi Mukkaṟamatsyaṁ Mukkaṟamatsyaṁ Paṭṭa Mukkaṟamatsyaṁ Veḷḷippuḷḷi Jīni Mukkaṟamatsyaṁ Mukkaṟamatsyaṁ Kēāḻi

Malayalam Name Malayalam Xhn«phcb³ sN¼Ãn shÅ¡dp¸v sN¼Ãn ]mXncm sN¼Ãn aªhme³ \oesN¼Ãn ]n©msem sN¼Ãn ISpwNph¸v tPm_v-sN¼Ãn kzÀWhcb³ sN¼Ãn kzÀWhcb³ Xp¸m¡naÕyw hÀW¸pÅn ^pkneoÀ aqhme³ s]m¡ {]mªnÂ sImSnb³ {]mªnÂ Po\n {]mªnÂ ZrUapÅ³ {]mªnÂ \of¡ncW {]mªnÂ CuÀ¡nÂ {]mªnÂ \of¨ndI³ {]mªnÂ hÀ® ap¡daÕyw \oe¡hnÄ shÅn ap¡daÕyw shÅn ap¡daÕyw ]pÅn ap¡daÕyw ]« ap¡daÕyw shÅn¸pÅn ap¡daÕyw Po\n ap¡daÕyw tImgn ap¡daÕyw

Authority 1830) (Cuvier, (Bleeker, 1850) (Bleeker, (Forsskål, 1775) (Forsskål, 1931 Fowler, 1869) (Bleeker, 1830) (Valenciennes, 1871) (Day, 1830) (Valenciennes, (Bloch, 1790) (Bloch, 1791) 1829 Cuvier, 1830 Cuvier, (Lacepède, 1801) 1854 Bleeker, 1830 Cuvier, 1849) (Cantor, 1792) (Thunberg, 1833) (Valenciennes, 1775) (Forsskål, (Lacepède, 1801) 1801) (Bloch & Schneider, (Bloch, 1790) 1867) (Playfair, (Bloch, 1793) (Quoy & Gaimard, 1824) & Gaimard, (Quoy Species Name chrysozona Pterocaesio Pinjalo pinjalo Macolor niger Macolor macularis Macolor xanthura Paracaesio filamentosus Pristipomoides multidens Pristipomoides balteatus Dipterygonotus surinamensis Lobotes erythrourusGerres filamentosus Gerres limbatus Gerres longirostris Gerres macracanthus Gerres oblongus Gerres longimanus Pentaprion Diagramma pictum argyreus Pomadasys argenteus Pomadasys commersonnii Pomadasys furcatus Pomadasys hasta Pomadasys multimaculatus Pomadasys maculatus Pomadasys Lutjanus vitta )

English Name English Caesionidae (fusiliers) 109. Family Goldband Fusilier Lobotidae (tripletails) 110. Family (mojarras) Gerreidae 111. Family Haemulidae (grunts) 112. Family (Blotched-grunt Saddle Grunt Pinjalo Snapper Black and White Snapper Black and White Snapper Midnight Snapper Blue Yellowtail Crimson Jobfish Goldbanded Jobfish Mottled Fusilier Tripletail) (Atlantic Tripletail Deep-Bodied Mojarra Mojarra Whipfin Saddleback Silver-Biddy (Longtail Silver-Biddy Strongspine Silverbiddy) Mojarra Long-Rayed Slender Silverbiddy Longfin Mojarra Sweetlip Painted Grunt Bluecheek Silver Grunt Silver Grunter) (Smallspotted Grunter Spotted Banded Grunter Grunt Spotted Silver Cock Grunter Brownstripe Red-Snapper Brownstripe

573 570 567 568 569 571 572 574 575 576 577 578 579 580 581 582 583 584 585 586 587 588 589 590 566

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‌ Mukk Vernacular Name Vernacular Olīv Mukkaṟamatsyaṁ Veḷḷivarayan aṟamatsyaṁ Karimmukkaṟamatsyaṁ Pāṭṭumukkaṟamatsyaṁ Intyansamudra Mukkaṟamatsyaṁ Puḻa Arimīn Arimīn Iruvarayan Arimīn Patimūkkan Chakravarttimatsyaṁ Nakṣatra Viralaṭayāḷa Chakravarttimatsyaṁ Pannimukha Chakravarttimatsyaṁ Chakravarttimatsyaṁ Kaṭiññāṇ Cheṟupallan Chakravarttimatsyaṁ Chakravarttimatsyaṁ Kuḻal Minnal Chakravarttimatsyaṁ Chakravarttimatsyaṁ Alaṅkāra Maññavarayan Chakravarttimatsyaṁ Chem'mūkkan Chakravarttimatsyaṁ Karimpuḷḷi Chakravarttimatsyaṁ Īrkkil Chakravarttimatsyaṁ Maññaccuṇṭan Chakravarttimatsyaṁ Peruṅkaṇṇan Patimūkkan Chakravarttimatsyaṁ Kiḷimīn Blīker Jappān Kiḷimīn Malayalam Name Malayalam Heohv- ap¡daÕyw shÅnhcb³ ap¡daÕyw Icnw ap¡daÕyw ]m«pap¡daÕyw C´y³kap{Z ap¡daÕyw ]pg Acnao³ Ccphcb³ Acnao³ ]Xnaq¡³ Acnao³ \£{X N{IhÀ¯naÕyw hnceSbmf N{IhÀ¯naÕyw ]¶napJ N{IhÀ¯naÕyw ISnªm¬ N{IhÀ¯naÕyw sNdp]Ã³ N{IhÀ¯naÕyw IpgÂ N{IhÀ¯naÕyw an¶Â N{IhÀ¯naÕyw Ae¦mc N{IhÀ¯naÕyw aªhcb³ N{IhÀ¯naÕyw sN½q¡³ N{IhÀ¯naÕyw Icnw]pÅn N{IhÀ¯naÕyw CuÀ¡nÂ N{IhÀ¯naÕyw aª¨pï³ N{IhÀ¯naÕyw ]Xnaq¡³ s]cpwI®³ N{IhÀ¯naÕyw »osIÀ Infnao³ P¸m³ Infnao³ Authority 1830) (Valenciennes, (Linnaeus, 1758) 1801) (Forster, 1830 Valenciennes, (Cuvier, 1830) (Cuvier, (Linnaeus, 1758) 1775) (Forsskål, 1775) (Forsskål, 1801) (Forster, (Lacepède, 1802) 1830 Valenciennes, 1775) (Forsskål, 1775) (Forsskål, 1830 Valenciennes, 1870 Klunzinger, (Bloch, 1791) (Forsskål, 1775) (Forsskål, 1775) (Forsskål, 1775) (Forsskål, (Lacepède, 1802) 1830 Valenciennes, 1830 Valenciennes, 1775) (Forsskål,

(Day, 1875) (Day, Species Name bipunctatus Nemipterus Plectorhinchus diagramus Plectorhinchus miniatus Lethrinus reticulatus Lethrinus Plectorhinchus nigrus Plectorhinchus vittatus Plectorhinchus berda Acanthopagrus Rhabdosargus sarba miniata Lethrinella lentjan Lethrinus microdon Lethrinus nebulosus Lethrinus ramak Lethrinus semicinctus Lethrinus xanthochilus Lethrinus japonicus Nemipterus Plectorhinchus schotaf Plectorhinchus bifasciatus Acanthopagrus harak Lethrinus mahsenoides Lethrinus ornatus Lethrinus variegatus Lethrinus grandoculis Monotaxis

Pomadasys olivaceus Pomadasys English Name English Sparidae (porgies) 113. Family or (emperors Lethrinidae 114. Family scavengers) breams, (threadfin Nemipteridae Family 115. breams) whiptail (Bleeker's Bream Threadfin Delagoa Threadfin-Bream) Silver Banded Sweetlip Banded Silver Emperor Trumpet Emperor Redsnout Black Sweet-Lip Sweetlip oriental Indian Ocean Bream) (River Picnic Seabream Stumpnose (Goldlined Seabream) Natal Bream Pigface Starry Emperor) (Pink Ear Bream Pig-Face Emperor Smalltooth Spangled Emperor Bream Pigface Yellow-Banded Emperor Black Blotch Emperor Yellowlip Bream Japanese Threadfin Minstrel Sweetlip Minstrel Seabream Twobar Emperor Thumbprint Bridled Pigface-Bream Emperor Ornate Slender Emperor Bream Humpnose Big-Eye Olive Grunt Olive

613 592 604 608 593 595 596 598 599 601 603 605 607 609 611 614 594 597 600 602 606 610 612 591

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chiṟakan chiṟakan ̄ Vernacular Name Vernacular Chennūlan Kiḷimīn Kiḷimīn Irucāṭṭa Kiḷimīn Kuḻicchiṟakan Ṟānḍal Kiḷimīn Īrkkil Kiḷimīn Minusa Oṟṟakkaṇṇaṭamatsyaṁ Oṟṟakkaṇṇaṭamatsyaṁ Kuḷḷan Chen Oṟṟakkaṇṇaṭamatsyaṁ Kuḷḷan Ṟēās Oṟṟakkaṇṇaṭamatsyaṁ Veḷḷakkaviḷan Oṟṟakkaṇṇaṭamatsyaṁ Nārumatsyaṁ Nāluviral Nārumatsyaṁ Intyan Karimpuḷḷi Aṟēbyan Nārumatsyaṁ Nārumatsyaṁ Varayan Nārumatsyaṁ Āṟuviral Nārumatsyaṁ Paṟudīsa Nārumatsyaṁ Karimpuḷḷi Kēāra Baṅgāḷ Tāṭikkēāra Blēācch Kēāra Chuṟṟikakkēāra Mūkkan Kēāra Belāṅṅēr Kēāra Kraṭṭ Sin Kēāra Valiyamēāntakkēāra Kēāra Katala Kēāra Karinvara Kēāra Cheṟu Kaṭuvappallan Malayalam Name Malayalam sN¶qe³ Infnao³ CcpNm« Infnao³ Ipgn¨ndI³ Infnao³ dm³UÂ Infnao³ CuÀ¡nÂ Infnao³ an\pk Hä¡®SaÕyw IpÅ³ Hä¡®SaÕyw sNwNndI³ Hä¡®SaÕyw tdmkv IpÅ³ Hä¡®SaÕyw shÅ¡hnf³ Hä¡®SaÕyw \mephncÂ \mcpaÕyw C´y³ \mcpaÕyw Atd_y³ Icn¼pÅn \mcpaÕyw hcb³ \mcpaÕyw BdphncÂ \mcpaÕyw ]dpZok \mcpaÕyw Icnw]pÅn \mcpaÕyw _wKmÄ tImc XmSnt¡mc t»m¨v tImc aq¡³ NpänIt¡mc s_emt§À tImc {Iu«v tImc kn³ tImc henbtam´t¡mc IXe tImc Icnwhc tImc sNdp ISph¸Ã³ tImc Authority (Valenciennes, 1830) (Valenciennes, 1830) (Cuvier, 1974 Trewavas, (Bleeker, 1853) (Bleeker, Russell, 1986 1981) (Rao & Rao, 1981) (Rao & Rao, (Bloch, 1792) 1804) (Shaw, 1926) (Hora, 1831) (Valenciennes, 1801) (Bloch & Schneider, 1855) (Bleeker, 1851) (Bleeker, 1830) (Cuvier, (Mohan, 1976) 1801) (Bloch & Schneider, (Bleeker, 1856) (Bleeker, Russell & Golani, 1993 1909) & Richardson, (Jordan 1804) (Shaw, 1830 Valenciennes, 1758 Linnaeus, 1830) (Cuvier, (Bloch, 1793) 1830) (Cuvier, 1830) (Cuvier,

(Bleeker, 1853) (Bleeker, Species Name Nemipterus peronii Nemipterus Johnius dussumieri cuvieri Otolithes Nemipterus nematophorus Nemipterus randalli Nemipterus aspinosa Parascolopsis boesemani Parascolopsis vosmeri Scolopsis tetradactylum Eleutheronema mullani Polydactylus sexfilis Polydactylus sextarius Polynemus Johnius amblycephalus Johnius borneensis Johnius carutta Johnius macrorhynus Nibea maculata Nemipterus zysron Nemipterus baranesi Parascolopsis eriomma Parascolopsis indicum Leptomelanosoma plebeius Polydactylus paradiseus Polynemus Daysciaena albida Johnius aneus Johnius belangerii axillaris Kathala

Nemipterus mesoprion Nemipterus English Name English (threadfins) Polynemidae 116. Family Notchedfin Threadfin Bream Threadfin Notchedfin Sin Croaker Croaker Lesser Tigertooth Doublewhip Threadfin Bream Threadfin Doublewhip Bream Randall's Threadfin Monocle Bream Smooth Dwarf Monocle Bream Dwarf Redfin Flash (Silver Monocle Bream Whitecheek Spine Cheek) Salmon ) (White Threadfin Fourfinger Threadfin Blackspot Arabian Threadfin Sixfinger Fish Black Spot Thread Croaker Bearded Sharpnose Croaker Hammer Croaker Karut Big-Snout Croaker Fish) (Black Banded Jew Croaker Blotched Slender Threadfin Bream Slender Threadfin Monocle Bream Dwarf Monocle Bream Dwarf Rosy Fish Indian Thread Striped Threadfin Threadfin Paradise Corvina Bengal Bloch's Croaker Croaker Belanger's Croaker Kathala Mauvelip Threadfin Bream (Redfilament (Redfilament Bream Threadfin Mauvelip Threadfin-Bream)

617 638 642 616 618 620 622 624 625 627 629 631 633 635 637 639 641 619 621 623 626 628 630 632 634 636 640 615

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Vernacular Name Vernacular Kaṭuvappallan Kēāra Kaṭuvappallan Kēāra Peruntala Kēāra Puḷḷi Peruntala Āṭumīn Maññavarayan Āṭumīn Svarṇavarayan Āṭumīn Iraṭṭavarayan Jīni Āṭumīn Āṭumīn Intyan Āṭumīn Nīḷasparśani Āṭumīn Veḷḷavarayan Āṭumīn Iruvarṇṇa Āṭumīn Svarṇappaṭṭa Āṭumīn Kāvivarayan Āṭumīn Nūlvarayan Āṭumīn Maññappaṭṭa Malabār Svīppar Svīppar Aṟṟakkaṟuppan Svīppar Sarayu Puḷḷi Villāḷimīn Nīla Kākkuṟāṭṭi Chempan Kākkuṟāṭṭi Matsyaṁ Arivāḷ Varayan Matsyaṁ Puḷḷi Arivāḷ Malayalam Name Malayalam ISph¸Ã³ tImc s]cpwXe tImc ]pÅn s]cpwXe tImc aªhcb³ BSpao³ kzÀWhcb³ BSpao³ Cc«hcb³ BSpao³ Po\n BSpao³ C´y³ BSpao³ \ofkv]Ài\n BSpao³ shÅhcb³ BSpao³ CcphÀ® BSpao³ kzÀW¸« BSpao³ Imhnhcb³ BSpao³ \qÂhcb³ BSpao³ aª¸« BSpao³ ae_mÀ kzo¸À Aä¡dp¸³ kzo¸À kcbp kzo¸À ]pÅn hnÃmfnao³ \oe Im¡pdm«n sN¼³ Im¡pdm«n hcb³ AcnhmÄ aÕyw ]pÅn AcnhmÄ aÕyw Authority Cuvier in Cuvier & Valenciennes 1831 1822) (Hamilton, 1775) (Forsskål, (Lacepède, 1801) (Lacepède, 1801) 1868) (Day, (Cuvier, 1830) (Cuvier, (Lacepède, 1801) (Lacepède, 1801) (Lacepède, 1802) 1855) (Bleeker, 1829 Cuvier, 1825) & Gaimard, (Quoy 1801) (Bloch & Schneider, (Bleeker, 1850) (Bleeker, 1874) Günther, 1855) (Bleeker, 2014 Randall & Bineesh, (Linnaeus, 1758) 1803) (Shaw, (Forskal, 1775) (Forskal, 1829) (Cuvier, (Bloch & Schneider, 1801) (Bloch & Schneider,

Species Name malabarica Pempheris chatareus Toxotes cinerascens Kyphosus Parupeneus cyclostomus Parupeneus macronemus Parupeneus guttatus Upeneus Pterotolithus maculatus Pterotolithus flavolineatus Mulloidichthys bifasciatus Parupeneus pleurotaenia Parupeneus moluccensis Upeneus taeniopterus Upeneus vaigiensis Kyphosus Drepane longimana Pennahia macrophthamlus Pennahia somoensis Mulloidichthys sundaicus Upeneus sarayu Pempheris Drepane punctata indicus Parupeneus vittatus Upeneus mangula Pempheris Otolithes ruber Otolithes )

English Name English Mullidae (goatfishes) 117. Family Indian Goatfish Goatfish Yellowstriped/Banded (sweepers) Pempheridae 118. Family Malabar Sweeper (Molucean Sweeper Sweeper Black-Edged (archerfishes) Toxotidae 119. Family Archerfish) (Largescale Archerfish Spotted (sea chubs) Kyphosidae 120. Family Blue ) Seachub (Ashen Drummer batfish, (spotted Drepaneidae 121. Family sickle fish) Monodactylidae (moonyfishes 122. Family or fingerfishes) Goldsaddle Goatfish Longbarbel Goatfish Goatfish Two-Tone Blotched Tiger-Toothed Croaker Tiger-Toothed Blotched Goatfish Yellowstripe Doublebar Goatfish Goatfish White-Lined Goldband Goatfish Goatfish Finstripe rudderfish) Chub (Lowfinned Brassy Concertina FishDrepane) (Bandede Bighead Pennah Croaker Bighead Pennah Slender Goldband Goatfish Goatfish Ochre-Banded Sweeper Sarayu Sicklefish Spotted Tiger-Toothed Croaker Tiger-Toothed

658 661 662 649 651 653 645 646 648 650 652 654 656 659 663 664 644 647 655 657 660 665 643

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8065 Fishes of Kerala Bijukumar & Raghavan

CITES

WPA

WG WG END

LC LC LC LC LC LC LC LC LC LC LC LC LC LC NE NE NE NE NE NE NE NE IUCN

tta Mālākhamatsyaṁ tta ̥ K ‌

Vernacular Name Vernacular Veḷḷi Chandramatsyaṁ Veḷḷi Citraśalabhamatsyaṁ Nūlvālan Chenvālan Citraśalabhamatsyaṁ Alasan Intyan Citraśalabhamatsyaṁ Chariññavarayan Citraśalabhamatsyaṁ Citraśalabhamatsyaṁ Ṟakkūṇ Kaṟuppmutukan Citraśalabhamatsyaṁ Citraśalabhamatsyaṁ Alakṣya Alasan Citraśalabhamatsyaṁ Maññattalayan Citraśalabhamatsyaṁ Paviḻamatsyaṁ Penneṟṟ Bānar Matsyaṁ Āṟumuḷḷan Citraśalabha Matsyaṁ Mālākhamatsyaṁ Maññavālan Iruḷan Mālākhamatsyaṁ Mālākhamatsyaṁ Vaḷaya Mālākhamatsyaṁ Chakravartti Ard'dhavr Pāychiṟak Mīn Kariyila avacattalayan Mīn Marayēānt Vālkkaṇṇan Cempalli Āttu Malayalam Name Malayalam shÅn N{µaÕyw \qÂhme³ Nn{XieàÕyw sNwhme³ Nn{XieàÕyw Cý³ Aek³ Nn{XieàÕyw Ncnªhcb³ Nn{XieàÕyw d¡q¬ Nn{XieàÕyw Idp¸vapXpI³ Nn{XieàÕyw Ae£y Nn{XieàÕyw Aek³ Nn{XieàÕyw aª¯eb³ Nn{XieàÕyw s]s¶äv ]hngaÕyw _m\À aÕyw BdpapÅ³ Nn{Xie` aÕyw aªhme³ amemJaÕyw Ccpf³ amemJaÕyw hfb amemJaÕyw N{IhÀ¯n amemJaÕyw AÀ²hr¯ amemJaÕyw ]mbvNndIv- IhN¯eb³ Icnbne ao³ hmÂ¡®³ actbm´v ao³ Bäp sN¼Ãn Authority Bloch & Schneider, 1801 Bloch & Schneider, 1867) (Playfair, (Forsskal, 1775) (Forsskal, 1801 Bloch & Schneider, (Linnaeus, 1758) 1831) (Cuvier, 1844 & Schlegel, Temminck 1822) (Hamilton, Ali Britz, & Philip, 2012 1849 Jerdon, Cuvier, 1829 Cuvier, (Lacepède, 1802) 1829) (Cuvier, (Bennett, 1833) (Bennett, 1833) (Bennett, (Bloch, 1787) (Bloch, 1787) (Forskal, 1775) (Forskal, (Bloch, 1787) (Linnaeus, 1758) 1831) (Cuvier, (Linnaeus, 1758)

Species Name Chaetodon meyeri Chaetodon multispinis Centropyge Chaetodon fasciatus Chaetodon melanotus Chaetodon Heniochus acuminatus ocellatus Parachaetodon typus Histiopterus Nandus nandus Dario urops marginata Pristolepis Chaetodon decussatus Chaetodon lunula Chaetodon Heniochus varius Chaetodon auriga Chaetodon collare Chaetodon vagabundus Chaetodon xanthocephalus Chaetodon xanthurus Apolemichthys annularis Pomacanthus imperator Pomacanthus semicirculatus Pomacanthus Monodactylus argenteus )

)

English Name English (butterflyfishes) Chaetodontidae 123. Family Butterflyfish Threadfin Fish Fish Butterfly Butterflyfish Yellow-Head (angelfishes) Pomacanthidae 124. Family Fish Angel (Ringed Blue Ring Angelfish Angelfish Emperor Fish (Blue Angel Angelfish Semicircle (armorheads) Pentacerotidae 125. Family fishes) Nandidae (leaf 126. Family Badidae (dario) 127. Family (catopra) Pristolepididae 128. Family Scrawled Butterflyfish Scrawled (Dusky Cherub) Dusky Angelfish Redtail Butterflyfish (Pakistani Butterfly (Pakistani Butterflyfish Redtail Banded (Eight Butterflyfish Diagonal Fish) Butterfly Blackback Butterflyfish Fish (Black Banded Butterfly Vagabond Fish Coral Pennet Butterflyfish Sixspine Angel (Yellow-Brown Angelfish Yellowtail Fish) Sailfin Armourhead fish Leaf Gangetic Dario³⁶ Ghats Western Common Catpora Indian Vagabond Butterflyfish Indian Vagabond Butterfly (Halfmoon Butterflyfish Raccoon Fish) Banner Fish Silver Moony (Silver Batfish) (Silver Moony Silver

667 673 680 682 668 670 672 674 676 678 679 681 683 684 685 686 687 669 671 677 675 666

8066 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

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KL END Exotic * LC LC LC NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NT IUCN

Vernacular Name Vernacular Ōṟañc Vālan Āṟṟucempalli Vālan Ōṟañc Nāṭamīn Karimpuḷḷi Kaṇamp Valiyacetumpal Kaṇamp Svarṇapuḷḷi Kaṇamp Pacchamutukan Kaṇamp Ṟṟēṭ Kaṇamp Chaturavālan Kaṇamp Nīḷakkayyan Nīlapuḷḷi Kaṇamp Kaṇamp Chappattalayan Kaṇamp Nīlavālan Karimīn Tilāppiya Meāsāmbikk Paḷḷatti Paṭattaṁ Varayan Paṭattaṁ Katrikavālan Paṭattaṁ Karimpuḷḷi Inḍēā-pesaphik Paṭattaṁ Paṭattaṁ Maññavālan Paṭattaṁ Nīḷavālan Paṭattaṁ Veḷḷappuḷḷi Nīla Paṭattaṁ Śud'dhajala Paṭattaṁ Puṣpa Mārivilmatsyaṁ Iruḷan Mārivilmatsyaṁ Malayalam Name Malayalam Hmd©v hme³ BäpsN¼Ãn Icnw]pÅn \mSao³ henbsNXp¼Â IW¼v kzÀW¸pÅn IW¼v ]¨apXpI³ IW¼v täSv IW¼v NXpchme³ IW¼v \of¡¿³ IW¼v \oe¸pÅn IW¼v N¸¯eb³ IW¼v \oehme³ IW¼v Icnao³ samkmw_n¡v Xnem¸nb ]Å¯n hcb³ ]S¯w I{XnIhme³ ]S¯w Icnw]pÅn ]S¯w C³tUm-s]k^nIv ]S¯w aªhme³ ]S¯w \ofhme³ ]S¯w shÅ¸pÅn ]S¯w \oe ]S¯w ip²Pe ]S¯w ]pjv] amcnhnÂaÕyw Ccpf³ amcnhnÂaÕyw Authority 1835) (Valenciennes, 1846) (Smith, (Bloch, 1790) 1830) (Cuvier, (Bloch, 1791) (Valenciennes, 1836) (Valenciennes, 1825) & Gaimard, (Quoy (Forsskål, 1775) (Forsskål, 1853) (Bleeker, 1775) (Forsskål, 1833) (Bennett, 1825) & Gaimard, (Quoy 1856) (Bleeker, (Hamilton, 1822) (Hamilton, 1775) (Forsskål, 1836) (Valenciennes, 1758 Linnaeus, 1852) (Peters, (Lacepède, 1801) 1825) & Gaimard, (Quoy 1882) (Macleay, 1825 & Gaimard, Quoy (Ruppell, 1835) (Bloch, 1795) Britz, Kumar & Baby, 2012 & Baby, Britz, Kumar

Species Name limbata Acanthocepola Chelon macrolepis suratensis Etroplus septemfasciatus Abudefduf Cheilinus chlorourus Chelon subviridis vaigiensis Liza Moolgarda seheli buchanani Valamugil sordidus Abudefduf xanthurus Aplomichthys Plectroglyphidodon lacrymatus taeniurus Pomacentrus Chelon parsia tade Liza Moolgarda cunnesius Mugil cephalus Oreochromis mossambicus sexfasciatus Abudefduf vaigiensis Abudefduf filamentosus Neopomacentrus caeruleus Pomacentrus Halichoeres marginatus Halichoeres Pseudetroplus maculatus Pseudetroplus Pristolepis rubripinnis Pristolepis

English Name English Cepolidae (bandfishes) 129. Family Bandfish Blackspot Mugilidae (mullets) 130. Family Mullet Largescale Cichlidae (pearlspot) 131. Family Spot Pearl (damselfishes) Pomacentridae 132. Family Banded Sergeant wrasses) fish, Labridae (rainbow 133. Family Wrasse Floral Dusky Wrasse Greenback Mullet Greenback Mullet Squaretail Bluespot Mullet Mullet Bluetail Sergeant Blackspot Angelfish) (Yellowtail Angelfish Brown Yellow Devil) (Jewel Devil Whitespotted Damsel Freshwater Goldspot Mullet Mullet Tade Mullet Longarm Mullet Flathead Tilapia* Mozambique Sergeant Scissortail Sergeant Indo-Pacific Demoiselle (Long-Lobed Damsel) Brown Caerulean Damsel (Blue Devil) Orange Chromide Orange Red Finned Catopra⁸ Finned Red 689 690 699 702 711 692 694 696 698 704 706 708 710 691 693 695 697 700 703 705 707 709 712 701 688

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8067 Fishes of Kerala Bijukumar & Raghavan

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WPA

END

LC LC LC LC LC LC LC LC LC LC LC LC NE NE NE NE NE NE NE NE NE NE NE DD IUCN

Vernacular Name Vernacular Kumiḷacchiṟakan Kumiḷacchiṟakan Mārivilmatsyaṁ Mārivilmatsyaṁ Puḷavan Mārivilmatsyaṁ Kampiccuṇṭan Irupuḷḷi Mārivilmatsyaṁ Mārivilmatsyaṁ Katti Mārivilmatsyaṁ Mayil Mārivilmatsyaṁ Añcuviral Mārivilmatsyaṁ Nīlavarayan Mārivilmatsyaṁ Chandra Tattamatsyaṁ Mūkkan Tattamatsyaṁ Nīlakkampi Tattamatsyaṁ Nāṭan Iruḷan Tattamatsyaṁ Mānattunēākki Mūkkan Mānattunēākki Maṇalpercc Kēāmāḷi Mucchiṟakan Nāṭan Blenni Kirkk Mimikri Blenni Kallarippan Nīlavarayan Blenni Paviḻavakkan Blenni Viravarayan Kallarippan Varayan Kallarippan Karinvara

Malayalam Name Malayalam Ipanf¨ndI³ amcnhnÂaÕyw ]pfh³ amcnhnÂaÕyw I¼n¨pï³ amcnhnÂaÕyw Ccp]pÅn amcnhnÂaÕyw I¯n amcnhnÂaÕyw abnÂ amcnhnÂaÕyw A©phncÂ amcnhnÂaÕyw \oehcb³ amcnhnÂaÕyw N{µ amcnhnÂaÕyw aq¡³ X¯aÕyw \oe¡¼n X¯aÕyw \mS³ X¯aÕyw Ccpf³ X¯aÕyw aq¡³ am\¯pt\m¡n am\¯pt\m¡n tImamfn aWÂs]À¨v \mS³ ap¨ndI³ InÀ¡v s»¶n anan{In s»¶n \oehcb³ IÃcn¸³ ]hngh¡³ s»¶n hnchcb³ s»¶n hcb³ IÃcn¸³ Icnwhc IÃcn¸³ 1903

Authority 1775) (Forsskål, 1843) & Schlegel, (Temminck 1909) (Weber, 1868) (Günther, (Bloch, 1792) 1840) (Valenciennes, (Linnaeus, 1758) 1829 Cuvier, 1836) (Valenciennes, 1836) (Valenciennes, 1836) (Valenciennes, (Bennett, 1832) (Bennett, (Rüppell, 1829) 1840) (Valenciennes, 1775) (Forsskal, 1840 Valenciennes, 1801) (Bloch & Schneider, 1836) (Valenciennes, 1836) (Valenciennes, Fowler, (Valenciennes, 1839) (Valenciennes, (Linnaeus, 1758) Forsskål, 1775 Forsskål, (Bloch & Schneider, 1801) (Bloch & Schneider,

Species Name harid Hipposcarus pulchella Parapercis fasciatus Enneapterygius Alticus kirkii Hemigymnus fasciatus cyanifrons Iniistius pentadactylus Iniistius gattatus Uranoscopus Blenniella periophthalmus vermiculatus Entomacrodus lineatus Istiblennius Halichoeres scapularis Halichoeres bimaculatus Iniistius pavo Iniistius ghobban Scarus russelii Scarus lebeck Ichthyscopus striatus Entomacrodus dussumieri Istiblennius Labroides dimidiatus Thalassoma lunare tractus Aspidontus

Scarus psittacus Scarus Halichoeres nigrescens Halichoeres

English Name English Wrasse Cleaner Bluestreak Moon Wrasse (parrotfishes) Scaridae 134. Family (Indian Ocean Candelamoa Parrotfish Longnose Parrotfish) (stargazers) Uranoscopidae 135. Family Pinguipedidae (sandperches) 136. Family Harlequin Sandperch blennies) (triplefin Tripterygiidae 137. Family Banded Triplefin blennies) Blenniidae (combtooth 138. Family Blenny Kirk's Barred Thicklip Barred Razorfish Wrasse Fivefinger Skygazer Blue-Dashed Rockskipper Blenny Vermiculated blenny) (Black-lined Lined Rockskipper Zigzag Wrasse Zigzag Razorfish Two-spot Wrasse Peacock (Blue-Barred Parrotfish Blue-Barred Parrotfish) Parrotfish Eclipse Longnosed Stargazer Mimic Blenny Blenny Reefmargin (Dussumier's Rockskipper Streaky Rockskipper) Common Parrotfish Bubblefin Wrasse Bubblefin

722 728 729 730 715 717 719 721 727 732 734 736 714 716 718 720 723 725 726 731 733 735 724 713

8068 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

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KL END

LC LC LC LC LC LC NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE DD IUCN Vernacular Name Vernacular Puṣpa Blenni Blenni Muṭivālan Vyāḷipūḻān Intyan Vyāḷipūḻān Āṟṟu Vyāḷipūḻān Jappān Nīḷavālan Maṇal Vyāḷipūḻān Vyāḷipūḻān Astra Iruḷan Pūḻān Ñeāṟipūḷān Kariṁ Puḻa Pūḷān Silēāṇ Pūḷān Pūḻān Ṭāṅk Cheru Pūḻān Maliññīl Pūḻān Pūḻān Kūḻāli Pūḻān Vālkaṇṇan Pūḻān Kuruṭan Pūḻān Keāmpan Vāvalmatsyaṁ Tūmpa Vāvalmatsyaṁ Gēāḷa Iruḷan Vāvalmatsyaṁ Vāvalmatsyaṁ Nīlacchiṟakan Puḷḷi Nacchār Karaṭ Veḷḷappuḷḷi

Malayalam Name Malayalam ]pjv] s»¶n apSnhme³ s»¶n C´y³ hymfn]qgm³ Bäp hymfn]qgm³ P¸m³ \ofhme³ hymfn]qgm³ aWÂ hymfn]qgm³ Akv{X hymfn]qgm³ Ccpf³ ]qgm³ Icnw sRmdn]qfm³ ]pg ]qfm³ kntem¬ ]qfm³ Sm¦v ]qgm³ sNdp ]qfm³ aenªoÂ ]qgm³ Iqgmen ]qgm³ hmÂI®³ ]qgm³ IpcpS³ ]qgm³ sIm¼³ ]qgm³ Xq¼ hmhÂaÕyw tKmf hmhÂaÕyw Ccpf³ hmhÂaÕyw \oe¨ndI³ hmhÂaÕyw ]pÅn \¨mÀ shÅ¸pÅn IcSv Authority 1890 Alcock, 1801) (Forster, (Rüppell, 1830) (Linnaeus, 1766) 1797) (Park, Swainson, 1839 Swainson, Houttuyn, 1782 1770 Pallas, 1989 Pethiyagoda, & Kottelat & John, 1983 Geevarghese 1837) (Valenciennes, 1801) (Bloch & Schneider, 1775) (Forsskal, Day, 1876 Day, 1919 Regan, 1877) (Day, 1822) (Hamilton, 1822) (Hamilton, 1853) (Bleeker, 1837) (Valenciennes, (Bloch, 1787) (Linnaeus, 1758) 1775) (Forsskal, Rüppell, 1830 Species Name carebares Callionymus Eleotris fusca fuscus Bathygobius argus Scatophagus Siganus canaliculatus Xiphasia setifer japonicus Callionymus sagitta Callionymus deraniyagalai Schismatogobius Glossogobius minutus tentacularis Oxyurichthys vagina Trypauchen orbicularis Platax Callionymus fluviatilis Callionymus marleyi Callionymus griseus Sicyopterus Glossogobius giuris rubicundus Odontamblyopus polynema Parachaeturichthys criniger Yongeichthys orbis Ephippus pinnatus Platax Platax teira Platax

Petroscirtus mitratus Petroscirtus )

English Name English (dragonets) Callionymidae 139. Family Dragonet Indian Deepwater Eleotridae (sleepers) 140. Family Dusky Sleeper¹ Gobiidae (gobies) 141. Family Frillfin Brown Ephippidae (spadefishes, 142. Family and scats) batfishes Batfish (Tiera Longfin Batfish (scats) Scatophagidae 143. Family Butterfish) (Spotted Scat Spotted (rabbitfishes) Siganidae 144. Family Spinefoot White-Spotted Hairtail Blenny (Hairtail Snakeblenny) (Hairtail Blenny Hairtail Dragonet Japanese Longtail Dragonet Arrow Goby Redneck Goby¹ Lake Veli Goby Tentacle Goby Burrowing Orbicular Batfish River Dragonet River Sand Dragonet Goby Clown Goby¹ Tank Rubicundus Eelgoby Goby Taileyed Horn Goby (Spadefish) Orbfish Round Batfish) ( Dusky Batfish Floral Blenny (White Spotted Blenny) Spotted (White Blenny Floral

739 744 745 759 760 738 741 743 747 749 751 753 756 758 740 742 746 748 750 752 754 755 757 737

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8069 Fishes of Kerala Bijukumar & Raghavan

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END

LC LC LC LC LC LC LC NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE IUCN

Vernacular Name Vernacular Murivarayan Karaṭ Murivarayan Karaṭ Svarṇavarayan Karaṭ Kēāṟiya Ṣū Karaṭ Karaṭ Viravarayan Karaṭ Iruvara Pālamīn Varayan Ōṟañch Pālamīn Peāṭinīla Nīḷan Pālamīn Pālamīn Nīlavarayan Pālamīn Taviṭṭu Pālamīn Varayan Puḷḷi Pālamīn Pālamīn Aḻakiya Keāṭiyan Śīlāv Peruṁ Śīlāv Maññavarayan Śīlāv Peruṅkaṇṇan Śīlāv Varayan Bālaṁ Ayalappāmp, Chākk Ayalappāmp Ayalappāmp Ṟeḍi Ayalappāmp Rājakīya Eṇṇa Ayalappāmp Ayalappāmp Kaṟuppan

Malayalam Name Malayalam apcnhcb³ IcSv kzÀWhcb³ IcSv tImdnb IcSv jq IcSv hnchcb³ IcSv Ccphc IcSv Hmd©v hcb³ ]meao³ s]mSn\oe ]meao³ \of³ ]meao³ \oehcb³ ]meao³ Xhn«p ]meao³ hcb³ ]meao³ ]pÅn ]meao³ AgInb ]meao³ sImSnb³ s]cpw ioemhv aªhcb³ ioemhv s]cpwI®³ ioemhv hcb³ ioemhv Abe¸m¼v, _mew Nm¡v Abe¸m¼v sduUn Abe¸m¼v cmPIob Abe¸m¼v F® Abe¸m¼v Idp¸³ Abe¸m¼v

Authority 1752) (Walbaum, Fowler, 1929 Fowler, (Valenciennes, 1835) (Valenciennes, 1835) (Valenciennes, 1833) (Bennett, 1829) (Cuvier, 1825) & Gaimard, (Quoy (Rüppell, 1829) 1829 Cuvier, Bonde, 1924) & von (Gilchrist 1856) (Bleeker, (Linnaeus, 1758) 1835) (Valenciennes, (Linnaeus, 1758) 1828) (Bennett, 1884 Klunzinger, 1829 Cuvier, 1829 Cuvier, 1832) (Cuvier, 1833 Cocco, (Valenciennes, 1835) (Valenciennes, (Valenciennes, 1835) (Valenciennes, 1775) (Forsskal, (Linnaeus, 1758) (Linnaeus, 1766) Species Name barracuda Sphyraena Thyrsitoides marleyi Thyrsitoides Siganus lineatus Siganus virgatus leucosternon Acanthurus mata Acanthurus striatus Ctenochaetus Naso elegans forsteri Sphyraena Neoepinnula orientalis prometheoides Rexea Siganus spinus Siganus vermiculatus lineatus Acanthurus strigosus Ctenochaetus chrysotaenia Sphyraena jello Sphyraena serpens Gempylus prometheus Promethichthys pretiosus Ruvettus Siganus sutor dussumieri Acanthurus nigrofuscus Acanthurus cornutus Zanclus

Siganus javus

)

) )

English Name English Fish Surgeon Fish (moorish idol) Zanclidae 145. Family Moorish Idol (barracudas) Sphyraenidae 146. Family Barracuda Great mackerels) (snake Gempylidae 147. Family (cutlassfishes) Trichiuridae 148. Family Black Snoek Golden-Lined Spinefoot Rabbit Spotted (White Spinefoot Shoemaker Spinefoot Double-Barred Surgeonfish Powderblue Fish) Fish (Bluelined Surgeon Lined Surgeon Surgeonfish Striated Unicornfish Elegant Barracuda Bigeye Sackfish Escolar Royal Scribbled Rabbitfish Spine-Foot Vermiculated Banded (Orange Surgeonfish Eyestripe Surgeonfish Elongate Tailed Surgeon (White Surgeonfish Brown Surgeonfish Spotted Barracuda Yellowstripe (Banded Barracuda) Pickhandle Barracuda Mackerel Snake Escolar Roudi Oilfish Streaked Spinefoot Streaked

775 776 785 762 764 766 768 770 772 774 778 781 783 763 765 767 769 771 773 777 779 780 782 784 761

8070 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

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LC LC LC LC LC LC LC LC LC LC LC NT NE NE NE NE NE NE NE NE NE DD DD DD IUCN

Vernacular Name Vernacular Nīḷappallan Vāḷa Vāḷa Ceṟutalayan Vāḷa Savāḷa Vāḷa Muttu Vāḷa Peruntalayan Oriyamīn Vāhū, Elicchūra Buḷḷeṟṟ Chūra, Ayalacchūra Uruḷan Chūra Chūra Nāypallan Skipjākk Chūra Ayala Varicchūra Nem'mīn Cheṟuvarayan Rājā Nem'mīn Nem'mīn Keāṟiyan Nem'mīn Varayan Cūra Pūvan Chūra Nīlacchiṟakan Kutiramīn Vāḷmīn, Pāymīn Intyan Mārlin Kariṁ Rēāmamatsyaṁ Intyan Oḻukkumīn Intyan

Malayalam Name Malayalam \of¸Ã³ hmf sNdpXeb³ hmf khmf hmf ap¯p hmf s]cpwXeb³ hmf hmlq, Hcnbao³ _psÅäv Nqc, Fen¨qc Abe¨qc Dcpf³ Nqc \mbv]Ã³ Nqc kvIn]vPm¡v Nqc Abe hcn¨qc sNdphcb³ s\½o³ cmPm s\½o³ sImdnb³ s\½o³ hcb³ s\½o³ ]qh³ Nqc \oe¨ndI³ Nqc hmÄao³, IpXncao³ Cý³ ]mbvao³ Icnw amÀen³ Cý³ tcmaaÕyw Cý³ Hgp¡pao³ Authority 1832) (Cuvier, 1890) (Alcock, 1894) (Waite, (Cuvier, 1829) (Cuvier, 1758 Linnaeus, (Lacepède, 1800) (Rüppell 1836) 1816) Cuvier, (Lacepède, 1800) 1788) (Bonnaterre, 1832) (Cuvier, (Gray, 1831) (Gray, 1884 Klunzinger, 1810) (Risso, 1849) (Cantor (Linnaeus, 1758) 1844) & Schlegel, (Temminck 1801) (Bloch & Schneider, 1829) (Cuvier, 1851 Bleeker, 1758 Linnaeus, 1792) (Shaw, (Kishinouye, 1915) (Kishinouye,

(Bleeker, 1860) (Bleeker, Species Name solandri Acanthocybium cyanea Psenopsis whiteleggii Cubiceps Lepturacanthus savala Lepturacanthus lepturus Trichiurus Auxis thazard unicolor Gymnosarda kanagurta Rastrelliger commerson Scomberomorus Thunnus albacares indica Istiompax Eupleurogrammus muticus Eupleurogrammus auriga Trichiurus Auxis rochei affinis Euthynnus pelamis Katsuwonus Sarda orientalis guttatus Scomberomorus lineolatus Scomberomorus Thunnus tonggol Xiphias gladius platypterus Istiophorus Scomberomorus koreanus Scomberomorus

Eupleurogrammus glossodon Eupleurogrammus English Name English tunas, (mackerels, Scombridae 149. Family bonitos) Wahoo Xiphiidae (swordfish) 150. Family (billfishes) Istiophoridae 151. Family (medusafishes) Centrolophidae 152. Family Indian Ruff (driftfishes) Nomeidae 153. Family Driftfish (Indian Driftfish) Shadow Savalai Hairtail (Small-headed Ribbonfish) Hairtail Savalai Hairtail Largehead Tuna) (Frigate Tuna Frigate Tuna Dogtooth ) (Indian Mackerel Mackerel Gillat Rake (King Spanish Mackerel Narrow-Barred Seer ) Fin Tuna Yellow Black Marlin Smallhead Hairtail Smallhead Hairtail Hairtail Pearly Tuna Bullet Tuna) (Mackerel Kawakawa (Skiy Jack ) Skipjack Tuna ) Bonito (Oriental Striped Bonito Spanish (Spotted King Mackerel Indo-Pacific ) Mackerel Seer Streaked Tuna) (Longtail Tuna Longtail Swordfish Sailfish (Indian Ocean Sail Fish) Indo-Pacific Korean Seerfish Seerfish Korean Longtooth Hairtail Longtooth

791 808 809 788 790 793 795 797 799 803 807 787 789 792 794 796 798 800 802 804 805 806 801 786

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̥ ḍamūkkan Māntaḷ ḍamūkkan Vernacular Name Vernacular Intyan Ariyēāmma Intyan Āvēāli Veḷḷa Chaina Āvēāli Karippiṭi Kariṅkaṇṇi Kariṅkaṇa Ḍēyuṭe Pulivāka Vaṭṭēān Cēṟumīn Puḷḷi Varāl Varāl Matsyaṁ Panni Palli Āyiraṁ Māntaḷ Chāra Ḍisk Māntaḷ Puḷḷi Māntaḷ Māntaḷ Pelikkan Dr Māntaḷ Valiyachetumpal Muppuḷḷi Māntaḷ Māntaḷ Kākki

Malayalam Name Malayalam C´y³ Acntbm½ shÅ Bthmen ssN\ Bthmen Icn¸nSn Icn¦®n tUbpsS Icn¦W ]penhmI ht«m³ tNdpao³ ]pÅn hcmÂ hcmÂ ]¶n aÕyw Bbncw ]Ãn Nmc am´Ä UnkvIv am´Ä ]pÅn am´Ä s]en¡³ am´Ä ZrUaq¡³ am´Ä henbsNXp¼Â am´Ä ap¸pÅn am´Ä Im¡n am´Ä Authority 1788) (Euphrasen, 1865) (Day, 1860) (Günther, (Köhler, 1908) (Köhler, 1822) (Hamilton, (Bloch, 1793) 1846) & Schlegel, (Temminck 1894 Alcock, 1846) & Schlegel, (Temminck 1931 Norman, (Euphrasen, 1788) (Euphrasen, (Bloch, 1792) 1831) (Cuvier, 1822) (Hamilton, (Bloch, 1793) (Bloch & Schneider 1801) 1865) (Bleeker, (Rüppell, 1830) 1890) (Alcock, 1865) (Bleeker,

(Day, 1871) (Day, Species Name argenteus Pampus Channa diplogramma rubescens Antigonia Pseudosphromenus dayi Channa marulius Channa striata Bothus myriaster lugubris Chascanopsetta grandisquama Engyprosopon natalensis Laeops Pampus chinensis Pampus Anabas testudineus Pseudosphromenus cupanus Channa gachua Channa punctata erumei Psettodes Arnoglossus tapeinosoma Bothus pantherinus Crossorhombus valderostratus polyophthalmus Grammatobothus

Ariomma indica English Name English (butterfishes) Stromateidae 155. Family Pomfret Silver (climbing perch) Anabantidae 156. Family fish) (paradise Osphronemidae 157. Family fishes) Channidae (snakehead 158. Family Snakehead Tiger (boarfishes) Caproidae 159. Family Boarfish Indo-Pacific XXXVI. ORDER PLEURONECTIFORMES (psettodids) Psettodidae 160. Family flounders) Bothidae (lefteye 161. Family Day's Paradise Fish Paradise Day's Snakehead Giant Striped Snakehead Flounder (Disc Flounder) Oval Indo-Pacific Flounder Pelican Flounder Largescale Khaki Flounder Chinese Silver Pomfret Silver Chinese Climbing Perch Fish Paradise Tailed Spike Snakehead Dwarf Snakehead Spotted Turbot) Indian Halibut (Indian Spiny Flounder Drab Flounder) Flounder (Panther Leopard Flounder Strongsnout Flounder Threespot 154. Family Ariommatidae (ariommatids) Ariommatidae 154. Family Indian Ariomma

811 816 821 815 818 820 824 826 828 830 812 813 814 817 819 822 823 825 827 829 810

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NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE IUCN tta Māntaḷ tta ̥ Vernacular Name Vernacular Karimpuḷḷi Māntaḷ Karimpuḷḷi Māntaḷ Valiyapallan Māntaḷ Kaṇṇan Māntaḷ Peākka Māntaḷ Jāvan Ceṟupallan Māntaḷ Muppuḷḷi Māntaḷ Māntaḷ Uccippūvan Māntaḷ Oṟṟakkeāmpan Māntaḷ Māntaḷ Pāl Māntaḷ Ard'dhavr Puḷḷi Māntaḷ Māntaḷ Kaṇṇan Nīḷan Māntaḷ Māntaḷ Muṭṭa Māntaḷ Kāpp Māntaḷ Keāmēḻsaṇ Māntaḷ Kēraḷa Māntaḷ Kvāga Māntaḷ Sībra Intyan Nākkmāntaḷ Valiyacetumpal Nākkmāntaḷ Nāluvari Nākkmāntaḷ Chūṇṭa Nākkmāntaḷ Parukkattalayan Nākkmāntaḷ Kāraṟṟ

Malayalam Name Malayalam Icn¼pÅn am´Ä henb]Ã³ am´Ä I®³ am´Ä s]m¡ am´Ä Pmh³ am´Ä sNdp]Ã³ am´Ä ap¸pÅn am´Ä D¨n¸qh³ am´Ä Häs¡m¼³ am´Ä am´Ä ]mÂ am´Ä am´Ä AÀ²hr¯ am´Ä ]pÅn am´Ä I®³ am´Ä \of³ am´Ä ap« am´Ä Im¸v am´Ä sImtagvk¬ am´Ä tIcf am´Ä IzmK am´Ä C´y³ ko{_ am´Ä henbsNXp¼Â \m¡vam´Ä \mephcn \m¡vam´Ä Nqï \m¡vam´Ä ]cp¡¯eb³ \m¡vam´Ä Imcäv \m¡vam´Ä Authority (Bloch & Schneider 1801) (Hamilton, 1822) (Hamilton, 1912 Ogilby, 1904 Gilchrist, 1913 Weber, 1894) (Alcock, 1801) (Bloch & Schneider, 1877 Day, 1858 Kaup, 1976 Joglekar, (Jenkins, 1910) 1889 Alcock, 1873 Day, Regan, 1905 Regan, 1853) (Bleeker, 1801) (Bloch & Schneider, 1831 Gray, 1858) (Kaup, 1896) (Steindachner, 1934 Fowler, 1889) (Alcock, 1846 Richardson, (Lacepède, 1802) (Lacepède, 1802) 1877 Day, 1858) (Kaup, von Bonde, 1922 Bonde, von Species Name Cynoglossus arel Pseudorhombus arsius Pseudorhombus elevatus Pseudorhombus natalensis cyaneu Aseraggodes umbratilis Aseraggodes Brachirus orientalis Solea elongata albomaculata Synaptura keralensis Zebrias synapturoides Zebrias Cynoglossus carpenteri Cynoglossus dubius Pseudorhombus dupliciocellatus Pseudorhombus javanicus Pseudorhombus triocellatus Samaris cristatus Aesopia cornuta kobensis Aseraggodes Brachirus annularis oculus Heteromycteris Solea ovata commersonnii Synaptura Cynoglossus bilineatus Cynoglossus dispar Zebrias quagga Zebrias

Laeops nigromaculatus Laeops )

English Name English flounders) Samaridae (crested 162. Family Soleidae (soles) 163. Family Sole Sole (Zebra Zebra Fringefin Cynoglossidae (tonguefishes) 164. Family Tonguesole Largescale Large Toothed Flounder Toothed Large Deep Flounder Flounder Smalltooth Blue Sole Sole Sole Oriental Sole Elongate Sole Kaup's Sole Kerala Sole Indian Zebra Tonguesole Hooked Tonguesole Carrot Ocellated Flounder Ocellated Flounder Javanese Spot Flounder Three Flounder Righteye Cockatoo Sole) Sole (Horned Unicorn Milk Solefish Annular Sole Sole Eyed Sole Ovate Sole Commerson's Tonguesole Fourlined Toungesole Roundhead Blackspotted Flounder Blackspotted

853 832 834 836 840 842 844 846 848 850 852 855 857 833 835 837 838 839 841 843 845 847 849 851 854 856 831

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LC NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE NE IUCN ̥ ga Phayalmatsyaṁ ga Vernacular Name Vernacular Tēāḷeppuḷḷi Nākkmāntaḷ Tēāḷeppuḷḷi Malabār Nākkmāntaḷ Puḷḷi Nākkmāntaḷ Nākkmāntaḷ Baṅgāḷ Chempuḷḷi Nākkmāntaḷ Nākkmāntaḷ Iruvarayan Kuṟṟimatsyaṁ Keāmpumūkkan Kuṟṟimatsyaṁ Īrcchavāḷmuḷḷan Nīlamuḷḷan Mukkālimatsyaṁ Cheṟumūkkan Mukkālimatsyaṁ Mukkālimatsyaṁ Veḷḷi Klātti Nakṣatra Chēāppupallan Klātti Klātti Chuṇṭan Klātti Maññavakkan Klātti Veḷḷavarayan Mukhammūṭi Klātti Klāttimatsyaṁ Varayan Phayalmatsyaṁ Oṟṟakkeāmpan Phayalmatsyaṁ Tēnkūṭ Phayalmatsyaṁ Āppuvālan Phayalmatsyaṁ Muṭicchiṟakan Phayalmatsyaṁ Nūlvarayan Kāṇṭāmr

Malayalam Name Malayalam tXmsf¸pÅn \m¡vam´Ä ae_mÀ \m¡vam´Ä ]pÅn \m¡vam´Ä _wKmÄ \m¡vam´Ä sN¼pÅn \m¡vam´Ä Ccphcb³ \m¡vam´Ä sIm¼paq¡³ IpänaÕyw CuÀ¨hmÄapÅ³ IpänaÕyw \oeapÅ³ ap¡menaÕyw sNdpaq¡³ ap¡menaÕyw shÅn ap¡menaÕyw \£{X ¢m¯n tNm¸p]Ã³ ¢m¯n Npï³ ¢m¯n aªh¡³ ¢m¯n shÅhcb³ ¢m¯n apJwaqSn ¢m¯n hcb³ ¢m¯naÕyw Häs¡m¼³ ^bÂaÕyw tX³IqSv ^bÂaÕyw B¸phme³ ^bÂaÕyw apSn¨ndI³ ^bÂaÕyw \qÂhcb³ ^bÂaÕyw ImïmarK ^bÂaÕyw Authority 1849) (Cantor, 1801) (Bloch & Schneider, (Linnaeus, 1758) Norman, 1928 Norman, 1877 Day, (Bloch, 1787) 1934 Fowler, 1801) (Bloch & Schneider, (Linnaeus, 1758) 1855) (Peters, (Rüppell, 1829) (Richardson, 1846) (Richardson, 1939 Norman, 1934 Fowler, (Bloch, 1786) (Rüppell, 1836) (Rüppell, 1829) 1804) (Latreille, 1850) & Schlegel, (Temminck 1850) & Schlegel, (Temminck 1852) (Bleeker, 1854) (Hollard, (Ruppell, 1837) (Bleeker, 1851) (Bleeker, Species Name strigilifer Pseudotriacanthus stellaris Abalistes monocerus Aluterus Cynoglossus macrostomus Cynoglossus semifasciatus bilineata Paraplagusia retrospinis Paratriacanthodes longirostris Oxymonacanthus aculeatus Rhinecanthus frenatus Paramonacanthus pusillus Paramonacanthus Cynoglossus puncticeps Cynoglossus zanzibarensis Macrorhamphosodes platycheilus biaculeatus Triacanthus Odonus niger flavimarginatus Pseudobalistes fraenatum Sufflamen oblongus Paramonacanthus nasicornis Pseudalutarius Triacanthus nieuhofii Triacanthus lineopunctatus Xanthichthys pardalis Cantherhines

Cynoglossus lida

English Name English XXXVII. ORDER TETRAODONTIFORMES (spikefishes) Triacanthodidae 165. Family (triplespines) Triacanthidae 166. Family Fish Long Spined Tripod Tripodfish Silver (trigger fishes) Balistidae 167. Family Triggerfish Starry Striped Triggerfish (filefishes) Monacanthidae 168. Family Leatherjacket Unicorn File Fish Honeycomb Malabar Tonguesole Toungesole Bengal (Doublelined Tonguesole Fingerlip Tonguesole) Spikefish Sawspine Longnose Filefish (Blackbar Triggerfish White-banded Triggerfish) Filefish Wedgetail Filefish Faintstripe Speckled Toungesole Speckled Toungesole Tonguefish Redspotted Spikefish Trumpetsnout Tripodfish Short-nosed Toothed File (Red Triggerfish Redtoothed Fish) Triggerfish Yellowmargin Triggerfish Masked Filefish Hair-finned Leatherjacket Rhinoceros Shoulderspot Tonguesole Shoulderspot

866 869 876 859 861 863 865 868 871 873 875 878 880 860 862 864 867 870 872 874 877 879 881 858

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Vernacular Name Vernacular Nīlakkeāmpan Phayalmatsyaṁ Nīlakkeāmpan Mañña Peṭṭimatsyaṁ Peṭṭimatsyaṁ Trikēāṇa Peṭṭimatsyaṁ Kūnan Āttuṇṭa Kaṭalmākri Veḷḷappuḷḷi Śud'dha Kaṭalmākri Puli Kaṭalmākri Kaṭalmākri Karimpuḷḷi Kaṭalmākri Jālikā Kaṭalmākri Nakṣatra Kaṭalmākri Mūkkan Kaṭalmākri Kirīṭa Kaṭalmākri Pālpuḷḷi Kaṭalmākri Valakaṭiyan Kaṭalmākri Chandravālan Kaṭalmākri Veḷḷikkaviḷan Kiḷicchuṇṭan Muḷḷanpannimatsyaṁ Puḷḷi Muḷḷanpannimatsyaṁ Puḷḷicchiṟakan Muḷḷanpannimatsyaṁ Nīḷamuḷḷan Muḷḷanpannimatsyaṁ Nāluvarayan Muḷḷanpannimatsyaṁ Sūryamatsyaṁ Peruṁ Īrkkil Sūryamatsyaṁ

Malayalam Name Malayalam \oes¡m¼³ ^bÂaÕyw aª s]«naÕyw {XntImW s]«naÕyw Iq\³ s]«naÕyw Bäpï shÅ¸pÅn ISÂam{In ip² ISÂam{In ]pen ISÂam{In Icn¼pÅn ISÂam{In PmenIm ISÂam{In \£{X ISÂam{In aq¡³ ISÂam{In IncoS ISÂam{In ]mÂ]pÅn ISÂam{In heISnb³ ISÂam{In N{µhme³ ISÂam{In shÅn¡hnf³ ISÂam{In Infn¨pï³ apÅ³]¶naÕyw ]pÅn apÅ³]¶naÕyw ]pÅn¨ndI³ apÅ³]¶naÕyw \ofapÅ³ apÅ³]¶naÕyw \mephcb³ apÅ³]¶naÕyw s]cpw kqcyaÕyw CuÀ¡nÂ kqcyaÕyw

Authority 1941) & Nair, (Hora (Bloch, 1785) 1758) Linnaeus, (Linnaeus, 1758) (Linnaeus, 1758) (Bloch ,1785) (Linnaeus, 1758) 1801) (Bloch & Schneider, 1878) (Day, 1801) (Bloch & Schneider, 1801) (Bloch & Schneider, 1798) (Anonymous, 1854) (Bleeker, 1875) & Sauvage, (Vaillant 1822) (Hamilton, 1850) & Schlegel, (Temminck 1801) (Bloch & Schneider, 1818) (Cuvier, 1854) (Bianconi, 1776) (Pennant, (Linnaeus, 1758) 1789) (Gmelin, (Linnaeus, 1758) (Linnaeus, 1758)

Species Name travancoricus Carinotetraodon orbicularis Cyclichthys Mola mola Lactoria cornuta Lactoria cubicus Ostracion concatenatus Tetrosomus gibbosus Tetrosomus immaculatusArothron leopardus Arothron nigropunctatus Arothron reticularis Arothron stellatus Arothron bennetti Canthigaster coronata Canthigaster Chelonodon patoca inermis Lagocephalus lunaris Lagocephalus jaculiferus Tragulichthys Lophodiodon calori laevis Ranzania hispidusArothron sceleratus Lagocephalus Diodon holocanthus Diodon hystrix ) )

)

English Name English fish) cow fish, (box Ostraciidae 169. Family fish ) (puffer Tetraodontidae 170. Family Fish Malabar Puffer Blaasop fish) (porcupine Diodontidae 171. Family Burrfish Birdbeak Fish Porcupine Porcupinefish Molidae (molas or ocean 172. Family sunfishes) Sun Fish) (Giant Sunfish Ocean Silver-cheeked Toadfish (Silverstripe (Silverstripe Toadfish Silver-cheeked Longhorn Cowfish Fish) Box (Ocellated Boxfish Yellow Boxfish Triangular Humpback Turretfish (White-spotted Puffer White-Spotted Blassop) Blossop) Edged (Black Puffer Immaculate (Bebo) Banded Leopardblowfish Puffer Blackspotted Fish Blow Reticulated Fish) Blow (Staring Puffer Stellate Puffer Sharpnose Bennett's Puffer Crowned Pufferfish) (Gangetic Puffer Milkspotted Fish ) Blow Smooth Blaasop (Smooth Backed Puffer Lunartail (Blotched Long-spine Porcupinefish (Spotted Porcupinefish Spot-Fin Longspined Porcupinefish Porcupinefish Four-bar Sunfish) (Truncated Slender Sunfish

886 899 904 898 882 883 884 885 887 888 889 890 891 892 893 894 895 896 897 900 901 902 903 905

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1 Known only from the Ashambu Hills 2 Known only from the Nilgiri Biosphere Reserve 3 Known only from the Periyar Watershed including the diverted waters into Tamil Nadu 4 Known only from the Santhampara Hills; Previously in the genus Horalabiosa 5 Known only from the Periyar River at Pooyamkutty 6 Known only from the south of the Palakkad Gap 7 Known only from the north of the Palakkad Gap 8 Known only from the Silent Valley National Park and Chinnar Wildlife Sanctuary 9 Known only from the Periyar Tiger Reserve 10 Possibly an introduced population 11 Known only from the rivers in Kasargod and Kannur districts 12 Known only from the Kabini River 13 Known only from the confluent rivers of the Vembanad Lake 14 In Kerala, its distribution is restricted to the rivers in Wayanad 15 Known only from the Kunthi River in Mannarkad 16 Known only from the Periyar, Chalakudy and Bharathapuzha Rivers 17 Known only from the Chinnar Wildlife Sanctuary 18 Known only from the Silent Valley National Park 19 Known only from the Silent Valley National Park and adjoining landscapes (Bhavani and Kunthipuzha) 20 Known only from the Anamalai Hills 21 Known only from the Santhampara Hills 22 In Kerala, it is known only from Wayanad District; Previously in the genus Longischistura 23 In Kerala, it is currently known only from the east flowing Kabini River 24 Known only from the Periyar and Chalakudy rivers 25 Known only from Thrissur District 26 Known only from the Chaliyar River 27 Known only from its type locality in the Manimala River 28 Known only from the Aralam Wildlife Sanctuary 29 Known only from Kottayam District 30 The parent taxa is now considered as Tylosurus acus acus (Lacepède, 1803) and the sub species in western Indian ocean, including Indian coastal waters is represented by Tylosurus acus melanotus (Bleeker, 1850) (Bailly, 2015) 31 Known only from its type locality in Kottayam; Not collected since its description * Exotic species

8076 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

Image 1. Sphyrna lewini Image 2. Torpedo sinuspersici

Image 3. Rhina ancylostoma

Image 4. Pastinachus sephen

Image 5. Gymnothorax reticularis

Image 6. Neoharriota pinnata

Image 7. Dussumieria acuta

Image 8. Plotosus lineatus

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8077 Fishes of Kerala Bijukumar & Raghavan

Image 10. Pterygotrigla arabica

Image 9. Antennarius nummifer

Image 12. Epinephelus tauvina

Image 11. Pterois miles

Image 13. Parupeneus indicus

Image 14. Trachinotus blochii

Image 16. Lutjanus argentimaculatus

Image 15. Pseudanthias marcia

8078 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 Fishes of Kerala Bijukumar & Raghavan

Image 17. Pomacanthus semicirculatus Image 18. Thalassoma lunare

Image 20. Brachirus annularis

Image 19. Odonus niger

Image 22. Garra periyarensis Image 21. Crossocheilus periyarensis

Image 24. Lepidopygopsis typus Image 23. Hypselobarbus periyarensis

Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080 8079 Fishes of Kerala Bijukumar & Raghavan

Image 25. Nemacheilus menonii Image 26. Mesonoemacheilus periyarensis

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Threatened Taxa

8080 Journal of Threatened Taxa | www.threatenedtaxa.org | 17 November 2015 | 7(13): 8036–8080