INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 261

The genus Scelolyperu8 Crotch in North America (Coleoptera: Chrysomelidae: Galerucinae)

Shawn M. Clark

Plant Industries Division West Virginia Department of Agriculture 1900 Kanawha Blvd., East Charleston, West Virginia 25305-0191

Abstract: A key and diagnoses are provided for the North American species of &elolyperus Crotch. Descriptions are gIven for SCelolyperus pasadenae. new speCies from Califorma and SCelolyperus tetone1l8~. new speCIes from Wyonnng. Luperus morriaoni Jacobyis treatedasajuniorsynonymofLuperusvaripes LeConte. Luperodes nigroviresce1l8Fallistreated as ajunioP syn~ofLupe1'us nig1'9Gyaneus LeC_te, and Scelolype1'us chautauquus Wileox is treated as ajunioP syllElDYftl of&elolyperusliriophilus Wilcox, new synonymies. Luperuslecontii Crotchis transferredto &elolyperus, new combination. Distributional andbiological data are provided for each species.

Introduction mques sunilar to those described by Smith (1979) In 1874, Croteh proposed the monotypie genus and Reid (1992). Genus Scelolyperus Crotch Scelolyperu8 Crotch, 1874:79 (Type: Scelolyperu8 tejoni. ,, , Arnett, 1962:912, 934; Wilcox, 1965:18, 19, 126.

cox prOVI e t e CrItIc mterpretatIOn t at IS Brancsik, 1899, by monotypy); Ogloblin, 1936:273; thebasisoftheeUffent understandingofthegroup. WIlcox, 1973:461 ( Scelolyperu8 Crotch, 18'14). Since his admirable treatment, two additional spe- ties were described by Hatch (1971). My investiga­ Diagnosis. Scelolyperus belongs to the section tions bave discovered two additional new species, Scelidites, a group ofclosely related genera in the as well as several other taxonomIC changes dIS­ subtrIbe Luperma of the trIbe Lupermi. This sec­ cussed in the treatment below. tion is usually recognizable by the presence of a In an effort to reduce the length of this paper, rectangular lobe at the apex of the male abdomen. detailed descIiptions have not been included, ex­ In NUl th AmeIica, rudy three otheI gIUUpS of Girl­ cept for new species. Likewise, data from material erucinae have similar lobes The first of these, examined (at least from localities in the United Phyllobrotica (sectIOn PhyllobrotIcItes) IS unlike States where specimens are often abundant) are Scelidites in having an extremely narrow epipleu usuallyreducedtocountyleveldistrIbutIOn, months ron. The second, section Monoleptites, is unlike eolleeted, and hosts mentioned. Previously pub Seelidites inha-v"ing a selerotized operculum to the lishedlocalityrecords are usuallymentionedonlyif aedeagal orifice and usually in having a very long they are consideled authentic and if they extend basitarsus on the hind leg. The third, Pteleon (sec­ distributions beyond those indicated by material tion Exosomites wbich may well be synonymous that I have personally exammed. Most ofthe mfor­ WIth Scelidites), IS represented m AmerIca by spe­ mation in this paper has been condensed from cies with rather shOl't antennae that reaeh only portions of an unpublished Ph.D. diSsertation per­ slightly beyond the humerus, antennomeres 2, 3, manently presetved at the Ohio State University and 4 being about equal in length. (Clark, 1987), anddetailedinformationistherefore Scelolyperus has a basal bead on the pronotum not entirely inaccessible to future researchers. but lacks obvious antennal or elytralmodifications Ibroregiven a greatdeal ofempbasis totbeform in male The elytra lack a conspicuous transverse of the aedeagI (a prelimmary study mdicated that unpressIOn atthe basalthIrd. The rectangularlobe the spermatheeae were not very diagnostic). at the apex ofthe male abdomen is less than halfas Aedeagal preparations were made following tech- long as broad. The longitudinal interantennal car­ illa is well developed, and the antenna! fossae are 262 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI separated by no more than twice the diameter of Ke" to New "'/odd Species ofScelolypel us each fossa. The eyes ofmost species are somewhat distantfromtheoralfossa, thegenaebeingaboutas 1. Pronotum pale 2 long as the terminal palpomere olthemaxj]]a This - Pronotum black, dark brown, metallic green, metallic combmatlOnofcharacterswilldistmgulshthISfrom blue, or metallic purple 16 other New World genera of Scelidites. All known species of Scelolyperus have entirely dark (either 2(1).Length 4.7 7.0 mm 3 Length 3.0 4.7 mm 9 black: or metallic) ely tra. Additional discussion and a key to genera will be provided in forthcoming 3(2). Aedeagus pointed er reunded at apex, asymmetl'i papersin whichtwonew generaare describedfrom cal (figs. 4c-d); hind tibia of male straight..... 4 Mexico Alldeagus broadlytruncate at apex, symmetrical (figs Non-metallic speCIes of ScelOlyperus from the 3b, 4l-m); hmd tlbla of male curved or stralght westemU.S.A arefrequentlyeonfusedwithPseudo 5 luperus longulus (LeConte), but that species has genae that aTe only about half as long as the 4(3).Tibiae of male with terminal spurs; length 2.9-4 7 termin rti 1 h m xilla 1 . M alIi mm; British Columbia, California, and Arizona ...... r uaus one - Tibiae of male lacking terminal spurs; length 5.9-7.0 mm; Califomia {lavicollis (LeConte)

5(3).Posterior tibia of male strongly curved, with a often confused with Lysathia ludoviciana (Fall), a prominent tooth on inner side at basal 1/3 (fig. :fleabeetle tllathas cffinpalatively widely sepalated 3d); terminal ahdominal tergite not produeed; front coxae terminal spurs present on middle tibiae, absent Comments. In spIte ofthe difficulty m charac­ from front and hind tihiae; length 5.0 mm; California tejonicus Crotch teriZin~ ~s genus, the ~P~Cies th;} are ~clu~ Posterior tibia without tooth 6

6(5).'I'ermmaI abdommal terglte of male strongly pro­ This is the only genus of SceIidites that has duced (fig 3g); bind tihia of male arcuate; representativesin theOldWorld(Asia) as wellas in length 4.5-6.0 mm; Cahforma . the Western Hemisphere Having examined Asian ...... megalurus Wilcox speCImens of S. altaicus (MannerheIm), I concur Terminal abdominal tergite ofmale not produced ... 1 with. Wilcox (1965) that this species does belong in ScelOlyperus. Only one other OldWorld species, S. ,((6).Hind tibiae with apicalspurs, innerside orhind tibia ormale evenly pubescent, withoutglabrous area is in sericeus (Jacobson), eunently classified the ...... 8 genus (Wilcox, 1973), but I have not examined Hind tibiae ofmale lacking apical spurs, inner side of specimens. Perhaps several ofthe Old World spe­ hindtibia ofmost males with a broad, glabrous, cies now included in Luperus or similar genera margined ehannel; elytra dark hiue, hiuish properly belong m ScelQlyperus 81so, butI have not green, or purple; pronotum black; apical and fully investigated this possibility. My treatment of hasal margins ofpronetum may he dark brown (variation in color indicates that specimens may occur with pale pronotuDl); length 5 5-60 da andthe USA, andin nearby areas ofnorthern mm; California curvipes Wilcox Mexico. 8(7). Hind tibia arcuate in male (fig. 3e); ventral side of The f~Uo~g key is a modification of ~at apex of aedeagus with two long longitudinal provided y W"Cox (1965). Most species are est carinae and two short longitudinal carinae (fig. recognized by male characters, and the key will 4j); length 5.3-5.9 mm; Cahfornia .. thereforenotenableidentificationofmanyfemales. . loripes Horn However, females ofafew species are recognizable Hind tibia ofmata nearly straight, ventral side ofapex and may be identified by consulting the diagnoses of aedeagus with 4 long longitudinal carinae of the vanous speCIes. (fig. 4k), length 5. 0 mm, California .. . ratulus Wilcox INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 263

9(2).Elytralpunetation moderato to rather ooarse; elytral 17(16). Elytra distinetly metallie blue, green, or purple . punctures separatedby a distance equalto twice 18 their diameter er less 10 Elytra blaek 01' dark bl'Own, without distinct blue, - Elytralpuncturesfine, separatedbya distance equalto green, or purple luster 26 three or four times their diameter .. 12 18(17). Hind tibia of male distinctly curved 19 10(9) Male with hind tibiae straight and last dorsal Hind tibia of male straight or nearlY so 21 abdominal segment normal 11 Male With hmd tibiae strongly curved and With termi­ 19(18). Pronotum distmctly green or blue, metaIhc; gla­ nal abdominal tergite greatly produced (figs. 3f- brous inner area ofhindtibia ofmale reduced or g); elytrausuallygreen, rarelyblue-green;length absent, not carinate or explanate at margins; 4.5-6.0 mm; California megalurus Wilcox apical portion ofaedeagus not widened; apex of aedeagus pointed or emarginate 20 ll(lO).Aedeagus nearly straight in lateral view, with - Pronotum without distinct blue, green, or purple Ius- sides gradually narrowing to apex in dorsal ter, hind tibia ormale usually with a ~ery broad, \tiew, with tip lateral (fig. 4d), elytra blue or shallow, glabrous ehannel on inner side, mar- green; length 2.9-4.7 mm; Arizona, California, gins of channel usually distinct, explanate on and possibly British Columbia .. ventl'aI1l2; apical pol'tion ofaedeagus widened; ...... torquatus (LoConte) apex of aedeagus truncate, not strongly emar­ Aedoagus distinctly sinuate in lateralview, somewhat ginate or pointed (fig. 3b); length 5.56.0 mm; spatulate in dorsaI view, with tip nearly central California curvipes Wilcox (fig 4£): elytra dark metallic blue, a little longer and more nearly parallel than in S. torquatus 20(19).Hind tibia of male strongly curved; aedeagus (hg an; length 34-42 mm; CalifOrnia emarginate at apex (fig 4e); length 41/-61/ mm; ...... phoxus Wilcox British Columbia to Montana to Wyoming to California schwarzii Horn 12(9). Each elytron with a strong, lateral, submarginal - Hind tibia of male weakly curved; aedeagus pointed carina; aedeagus symmetrical, with distal pOI'- at apex; length 4.4-5. '7 mm; Oregon . tion gradually narrowed to acute tip; length 3.3- ...... wilcoxi Hatch 4. '7 mm, California, Oregon ...... carinatus Wilcox 21(18). All femora and tibiae entirely dark 22 Elytra lacking distinct carinae 13 At least some legs with femora and tibiae at least partly pale 24 13(12). Aedeagus slender, nearly symmetrieal, not spat ulate at apex (figs. 2, 4h) 14 22(21).Aedeagus with acute tip at apex (figs. 1, 4b); Aodeagus broader, distinetly asymmetrieal or spatu terminal spurs pl'Osent on all tibiae of male ... late at apex (figs. 4a, 4g) 15 ...... 23 - Aedeagus With rounded apex (fig. 5c); all tibiae of 14(13).Aodeagus strongly sinuate in lateral view (fig male lacking terminal spurs; length. 6 6 7 0 4h): length 3.4-3.9 mm; Califorma .. mm; Cahforma graptoderoides (Crotch) phenacus Wilcox Aedeagus only slightly slDuate, nearly straight lD 23(22). Aedeagus nearly straight in lateral View; distal lateral view (fig. 2); length 3.0-3.7 mm; Wyo- 1/4 of aedeagus with an acute, very well-devel- ming tetonensis, new species oped, ventrolateral carina; length 5.3-6.7 mm; California smaragdinus (LeConte) 15(13).Aodeagus spatulate, without acute or rectangu­ - Aedeagus sinuate in lateral view; ventrolateral cari­ lar tip (fig. 4a); length 3.2-4.2 mm; California nae on distal portion ofaedeagus either weak or ...... transitus (Horn) absent (fig. 1); length 5.5-7.1 mm; California . Aedeagus strongly sinuate in lateral view, not spat- ...... pu8tJi.letwe, {lew species ulate in dorsal view; acute or rectangular tip of aedeagus far to the right (fig. 4g), length 3.0 4.7 24(21). Pronotum with distinet metallie luster, aedea mm; California, Idaho laticeps (Hom) gus straight or weakly sinuate in lateral view; elytl'al punetures larger : 25 16(1).Western species found west of the Great Plains - Pronotum lacking metallic luster; aedeagus, in later­ (100 0 VI'OSt longitude) . 17 9.1 view, appeariiig bent near dist91 third; ely-

- Eastern species found east of the Great Plains (100 0 tral punctures very fine, sometimes obscured west longitude) 28 byalutaceousmicroscu1pture; length3 8-5 8mm; British Columbia to South Dakota to New Mex- iCO to Califorma leContn (CrotCh) 264 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI

25(24). Distal pOItion ofaedeagus distinctly asymmetri­ Diagnosis. In females of this species, each cal (figs. 5b, 5d); length 3.4-5.3 mm; British elytron has a well-developed, acute, submarginal Columbia to .Alberta to Ne''''' Mexico to CaliCoI' carina. Also, the aedeagus inlateralview is sinuate nia varipes (LeConte) in the basal 1/2 and straight in the distal 1/2; in Distal portion of aedeagus symmetrieal; length 4.4 dorsal VIew, It IS not spatulate m the distal 1/2. 5.7 mm; Oregon wilcoxi Hatch These characters distinguish this species (either 26(17) Middle and hind tibiae largely or entirely dark sex)from allother species ofScelolyperus occurring brown; surface between elytraI punctures usu­ in the eastern half of Nor th America. Specimens ally alutaceous 27 measure 4.0-5.4 mm long. All tibiae testaceous; surface between elytral punc­ Type locality. "Mt. Mitchell, North Carolina, tures not alutaceous; apex ofaedeagus strongly 6,000£t" produced, asymmetrical (fig. 5h); 3. 7-4.9 mm, Distribution. West Vrrgmla to North Caroli­ Washington to California hatchi Wilcox na and Tennessee. I have examined specimens

27(26).Basal 3/4 of aedeag uS neally straight in lateIal view; apex of aedeagus only slightly asymmet- rical, length 3.5 mm, Idaho lemhi Hatch Virginia Beyond material that I have seen, Wilcox - Aedeagus, including basal 3/4, sinuate in lateral view; (1965) reported thIS species from Tennessee. apex of aedeagus distinetly asymmetrieal in Comments. Specimens have been collected dorsal view (figs. 5e-g); length 3.0-4.3 mm; from May to August. One of them was taken on British Columbia to Alberta to New Mexieo .. H~llebtJru8viridis L. (Ranuneulaeeae) and another ...... nigrocyaneus (LeConte) on Phlox subulata L. (polemoniaceae). I have ex- amilled themaleholotype(NMNH), tenother males, 28(16). Elytra offemale without distinct, acute, submar- and 24 females ginal carina . . 29 - Elytra of female with an acute, submarginal carina; Scelolyperus carinatus Wilcox aedeagus with basal % sinuate and distal 112 straight in lateral view, lacking spatulate dis­ (Figure 3i) tal portion in dorsal view (fig. 5i), length 4.0-5.4 mm; West Virginia to North Carolina to Ten- Scelolyperus carinatus Wilcox, 1965:128, 131, 144. nessee bimalginatus (Blake) Diagnosis. In this species, each elytron has a 29(28). Apical 1/2 of aede.agus not, or little, wider than prominent, acutecarinaextendingposteriorlyfrom basal half(figs. 5j-l); hind tibia usually with an the humerus. This, in combination with the pale apieal, lamellate lobe in male 30 pronotum, will distinguish S. Cal inatas from all other species of the genus as well as of the entire half ( Ig. 5a); ind ti ia 0 male wit out apica section Scelidites. Specimens measure 3.3-4.7 mm ~mellll:e lobe; l;gti ~8 ~? mm; Illinois to lOng. ennsy vania to art. aroima . cyanelluB (LeConte) Typelocality."HumboldtCo., California, 13.6, Bair's Reb., Redwd. Crk." 30(29) AedeagllS sinllate in lateral view (fig 5j); femQra Distribution. California and Oregon. I have usually dark; length 3.8-5.2 mm; New Brun- seen specimens from the following counties. Cali- swick to Tennessee to Georgia...... fornia' Humboldt Co, Madera Co Oregon' Lane ...... meracus (Say) Co. Aedeagus straight in lateral view (figs. ok-I); front Comments. Specimens have been cOllected in and middle femora usually testaceous; length Ma; and June. I have examined the female holo- 3.0-4.'1 mm; Missouri to Maille to Georgia ...... liriophilus Wilcox Scelolyperus bimarginatus (Blake) Scelolyperus curvipes Wilcox (Figure 5i) (Figures 3b, 3c)

Lupefodes bimarginata Blake, 1928.183. Scelol:yperus curvipes Wilcox, 1965:128. 131, 132. 141. Scelolyperus bimarginatus: Wilcox, 1965:128, 133, 153. Diagnosis The strongly curved bind tibia of the male, together with the pronotum that is dark INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 265 but lacks metallic luster, will distinguish this from 8eelolYPeFUS flalJieoUis: Horn, 1893:103, 104; Wileox, other species of Scelolyperus. Specimens measure 1965:128, 130, 137. 5.5-6.0 mm long. Diagnosis. The pronotum of this species is Type locality Boulder Creek, Fresno Co, pale, the apex of the aedeagus is pointed and CaIiforma. asymmetrical, and the overall size is quite large Distribution. California. I have seen speci (59-70 mm long) These characters distinguish mens from Fresno and Mariposa Counties, both in this from other speCIes of Scelolyperus. Califomia. Type locality. Tejon, CaliIOinia. Comments. Specimens have been collected in Distribution. California. I haveexaminedspec- May. Somehavebeen associatedwithArtemisiasp. imens from Kern, Monterey, and San Diego Coun­ (Asteraceae) and others with flowers of Madia ties in California. All of the localities that were elegans Don (Asteraceae). I have exammed the erroneously listed by Wilcox (1965) as being in male holotype (CASC), 25 other males, and 15 Arizona are actually in California females. Comments. Speclffiens have been conected ill April and May. Some were found on Erysimum Scelolyperus cyanellus (LeConte) argillosum (Greene) Rydb. (Brassicaceae). Thepre- (Figure 5a) ceding treatment is based on the female holotype (MCZC), on 40 other females, and on 17 males. LuperU8 cyanellu8 LeConte, 1865.209. Luperus (Luperus) cyanellus: Weise, 1924:118. Scelolyperus graptoderoides (Crotch) Luperodes cyanellus: Horn, 1893:116; Wilcox, 1954:434. (Figure 5c) &elolyperus cyanellu8. Vlileox, 1965.128, 133, 151; (not Horn, 1895:251). Luperus graptoderoides Crotch, 1874:80. Scelolyperu8 Rraptoderoides. Horn, 1893:105; Wilcox. Diagnosis. The distal 1/2 of the aedeagus is 1965:128, 132, 149. about 2X as wide as the basal 1/2. This chalacter will distinguish S. cyanellusfrom the other species Diagnosis. In this species, the prothorax and of Scelolyperus found m the eastern half of North legs are entirely dark, the hilld tIbIa of the male IS America. Specimens measure 3.8-5.0 mm long. nearlystraight, andallofthe tibiaeofthemalelack Type locality. "Western States; Michigan, Il­ terminal spurs. These characters together diStin­ linois." guish this from other species of Scelolyperus that Distribution. Illinois toPennsylvaniatoNorth occur in the western halfofNorth America. Speci­ Carolina I have seen specimens from thefollowing mens measure 6.6-7.0 mm long. counties Kentucky' Menifee Co Ohio' Clinton Co, Type locality Specimens in the twe series Delaware Co., Franklin Co., Pike Co., Warren Co. came from "Santa Barbara" and"Santa Buenevim­ North Carolina: Buncombe Co. Pennsylvania: AI tura" in California. Ho'...,ever, itis notclear'...,hieh of legheny Co., Lawrence Co., Washington Co. West these is the actual type locality. Virginia. Pocahontas Co. Dishibution. California. I have seen speci- Earlier records, such as those ofFattig (1948) mens from Ventura County, California. Also, part fOr Georgia, may refer to species other tnan S of the type series is ftom Santa Barbara County, eyanellus. California, and Horn (1893) recorded this species Comments. I have seen specimens collectedin from Los Arigeles County, California. May, June, and September and ha'\l e personally Comments. Specimens have been collected in associated this species with Phlox paniculata L. May. Theprecedingtreatmentis basedonthe male (polemoniaceae) holotype (MCZC), on two other males, and on one The preceding treatment is based on the male female. holotype (MCZC), 30 other males, 49 females, and one specimen of unknoJNn sex. Scelolyperus hatchi Wilcox (Figures 3h, Oh) ScelolJ'Perus flol'icollis (LeConte) (FIgure 4C) Scelolyperus hatchi Wilcox, 1965:128, 133, 157; Hatch, 1971:205. Phyllobrotica flavicollis LeConte, 1859b:81. Luperu8 flfWic6llis: LeConte, 1861>:209. 266 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI

Diagnosis. The elytIaofthis species aleblack, Scelolypel U8 lecontii (ClUtch), new without metallic luster, the elytralinterspaces are combination not alutaceous, and allofthe tibiae are testaceous. Tbese characters will together distinguish this LuperUB rulipeB LeConte, 1859a:27; LeConte, 1865:209; from all other speCIeS of Scelolyperus. Specunens Crotch, 18'73.54 ( Luperu8 Lecontii Crotch, 18'73), measure 3.7 4.9 mm long. (not Scopoli, 1763:73; not Goeze, 1777:322; not Fab- Type locality. Boyer, Oregon. rieius, 1787.89, not Gyllenhal, 1813.513, not Ratze­ burg, 1837.254). Distribution. WashingtontoCalifolnia. I have LuperUB Lecontii Crotch, 1873:54 (replacement name seen specimens from the following counties. Cali- for LupernB rufipes LeConte, 18a9a). fornia: Del Norte Co., Humboldt Co., Mendocino LuperUB lecontii: Jacoby, 1888:596. Co Oregon' Coos Co , Tillamook Co , Yamhill Co Luperod88 LeGontii: IWl'l'l, 1893:115. Washmgton King Co. PBeudoluperuB leconth: Wilcox, 1965:101, 102, 107: Beyond speeimens that I have seen, \Vileox Hatch, 1971'203 1965) re orted this s ecies from Skamania Coun· LuperUB LeconteJ: Crotch, 1880:98. LuperUB (LuperuB) Lecontei: Weise, 1924:119. u ero con eJ aBC J JB a cox, 1965:101 (=Zecontii Crotch, 1873). Luperu8 pallipe8 Beller & Hatch, 1932.113; Wilcox, 1965:101 (=Zecontii Crotch, 1873). Scelolyperus laticeps (Horn) (Figure 4g) Diagnosis The hind tibiae of the male are LuperodeB laticepB Horn, 1893:114. LuperuB (Luperui) latll:epB' weise, 1924:119. Scelolyperuslaticep8: Wilcox, 1965:128, 132, 141; Hatch, 1971'203 westernhalfofNorthAmerica. Specimensmeasure Diagnosis. The pronotum of thIS speCIes is 3.8-5.8 lInn long. pale, the elytral punetures are separated by a Tr,ree )octjtiesitfeeru;e~lIfipes·~?taFe, distance three to four times the diameter of a punctule, and the distalpoltion ottlIe aedeagus is distinctly asymmetrical, with the acute or rectan­ ton, Washington. gular t.ip located far 10 the right This combination Distribution. Blitish Columbia to South Da- of characters distinguishes S. laticeps from other kota to New Mexico to California. I have examined species ofScelolyperus. Specimens measure 3.0-4.7 mmlong. Type locality. "California, region unknown." Ittooet;iIlay~; errltt; Nmee, TInce: Distribution California, Oregon, and Idaho ton; Shingle Cr. Road, Keremeos; Tel'l'aee. I have seen specimensfrom the following counties: I have also examined specimens from the fol­ CalifOrnia: Alameda Co., Contra Costa Co., Lake lowing counties in the United States. Arizona. Co., Mendocino Co., Santa Clara Co., Siskiyou Co., Apache Co, Cochise Co, Coconino Co, Gila Co Trinity Co. Idaho: Boise Co., Canyon Co., Twin Califorma: Del Norte Co., Modoc Co., Napa Co., Falls Co. Oregon. Bakel Co., Malheul Co. Beyond ~~:: g:.·'2J~~:;0~~:a~~: ~'.: ~~:i~:; specimens thatI have seen, Wilcox (1965) recorded g::: this species ft()lTl San Francisco, California [San Clear Creek Co., EIPaso Co., Garfield Co., Larimer Francisco County]. Co., Mesa Co., Mineral Co., Ouray Co., Routt Co., Comments. I have examined the male lecto­ San Miguel Co. Idaho. Bannock Co., Bonneville type (designatedby Wilcox, 1965; MCZC), 67 other males, and 75 females. Specimens have been col­ on ana: ana mo., owe CO: ew eXlCO: lectedfrom April to July. Some werefoundonRibes Colfax Co., San Miguel Co., Santa Fe Co. [I have sp. (Saxifragaceae) and others on Solanum sp. also seen specimens from the Jemez Mts. (Rio (SOlanaceae) Aniba 01 Sandoval Co.»). Oregon. Jackson Co.! Josephine Co., Klamath Co., Lincoln Co., Umatilla Co., Wallowa Co., Wasco Co. South Dakota: INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 267

Lawrence Co. Utah: Cache Co., Grand Co., Salt i\meriea. However, for confident identification Lake Co.. San Juan Co., Utah Co., Wasatch Co., aedeagal examination is necessary. The aedeagus Weber Co. Washington. Kittitas Co., Masrol Co., is about as bIOad in the basal ~ as in the distal ~, Skamania Co , Walla Walla Co , Yakima Co Wyo. and it is nearly straight in lateral view (figs 5k·l) mmg: Albany Co., Fremont Co., SherIdan Co. SpecImens measure 3.0-4.7 mm long. Comments. Apocynum androsaemi{9lium L. Type localities. Scelolyperus liriophilus: (Apocynaceae) and Psoralea physodes Doug!. (Fa- Haines Falls, New York. ScelOlyperus chautauq- baceae) have been repozted as hosts. Asclepias ltUs: Ashland Co., Ohio. speciosa Torr. (Asclepiadaceae) has also been re- Distribution. Missouri to Arkansas to Geor- ported as a possible host, but, as pointed out by gia to Maine. I have seen specimens from: the Wilcox (1965), this is probably not correct Speci- following counties Alabama' Shelby Co Georgia' mens have been collected from April to August. Dade Co., Jackson Co., Omon Co. IllinOIS: Johnson I haveexaminedtheholotypeofLuperus rufipes Co., Pope Co., Pulaski Co., Putnam Co. Kentucky: (sex unknown, MCZC), the female holotype ofLu- Elliot Co., Rowan Co. Maine: [I have seen a speci- pel as pallipes (NMNII), the male holotype of Lu· men from "Wissataquoik R." which is in Penobscot perodes lecontei asclepiadis (NMNH), 279 other and Piscataquis Counties). Missouri: Barry Co., males, 303 other females, and two specimens of Gasconade Co., Jefferson Co., Taney Co. New Jer- unknown sex. sey' [The label included no specific locality] New York: Albany Co., Greene Co., UlSter Co. North Scelolyperus lemhi Hatch Carolina: ..ANery Co., Jackson Co., Macon Co. Ohio: Scelolyperu8 lemhi Hatch, 1971:205. Athens Co., Delaware Co., Franklin Co., Greene 00., Hocking 00., Lake 00., 'linton 00., 'Nanen 00. Diagnosis. In this species, the middle tibia, P nns Ivania'All h n Co. But! r . Nor h m - n 0., es more an o. es irginia: ar our dark brown, without metallic luster. Within Sce­ go ,~e~te C~' C~a~ CoGxe~nbrie~C~,H:;dy lolyperus, only S. nigroeyaneus is similar in these o.,arrlSOll 0.; ae son CO:; anaw a 0:; m­ characters. However, unlike S. nigrocyaneus, the era! Co., Monongalia Co., Pendleton Co., Poeahon apicalportionofthe aedeagus isonly weakly asym­ tas Co., Preston Co., Rale~? Co., Rando~h Co., metrical, and the basal three-fourths ofthe aedea· Roane Co., Taylo:r Co., Tuc el Co., Wilt 0 . gus are nearly straight in lateral view. The single ~y~~mater~l::hatl~~~een !~~1965) specimen examined is 3.5 mm lOng Type locality. "8 mi. e Tendoy, Ida., Lemhi Logan County in Arkansas, and Hamilton County Co." inTennessee. Riley andEnns (1979, 1982) reported Dish ibution. This species is known only flOm it from Washington Oounty in Arkansas andfrom the type locality. Benton and Marion Counties in Missouri. Comments This species has heen collected in July. ItisverysimilartoS. nigrocyaneusbutdiffers in the form Of the aedeagus. The preceding treat- us . an . va. Iaceae ,an arptnus ment is based on examination ofthe male holotype earoliniana \Jlalt. (Betulaceae). Also, Riley and (CASC). I havenot discover edany other specimens of this species. leave of Coti us obovatus RaE. ( lacmdiaceae) n S h lea tri olia L. Sta h leaceae . Be ond (Figures 3a, 5k, 51) s, ave examIne specimens a were co ec . ed while :weding on Juglans nigra L. (Juglandace- Scelolyperu8 liriophilu8 Wiloox, 1965:128, 133, 155; ae) andoncultivatedPhlox(polemoniaceae). I have RIley & Enns, 1979:70; RIley & Enns, 1982:34. also seen specimens thatwere collectedon Sambu &elolyperus chaUtaUQUU8 Wiloox, 1965:128, 133. 156, cus (Caprifoliaceae). Quercus (Fagaceae), andSalix New synonymy. (Salicaceae), but feeding was not specifically indio cated In my personal field work, I have observed Diagnosis. In most specimens of this species, the beetles on a WIde vanety ofplants, but usuany the legs are largely or ent.irely pale This will often without evidence offeeding. Specimens have been diStmgulSh S. ltrwphdus from the other speCIes of collected from April to August. Seelolyperus oecurring in the easternhalfofNorth 268 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI

Scelolypel us iiI iophilus and S. chautauquus Diagnosis. In the male of this species, dIe were originally described in the same publication terminal abdominal tergite is strongly, narrowly and were distinguishedby differences in the shape produced. This character is unique in the genus ofthe aedeagus andin overall size However, these and in the entire section Scelidites. Specimens characters are somewhat vanable and not neces­ measure 4.5-6.0 mm long. sarily cOlTelated with each other. I have seen sev Type locality. "Kaweah, Tulare Co., Califor eral specimens that are quite intermediate in the nia, 1,000 ft." form of the aedeagus. I cruiclude that the two taxa DistIibution. This species is known rudy f10m are synonymous Tulare County, California As both names were proposed m the same Comments. SpecImens have been collected publication, either couldbe chosen for this species. during May and July. Some were taken from the I am follOWIng tradition m selecting S. liriophilus, flowers ofMadia elegans Don (ASteraceae). I have the name whose formal description appeared first examined the male holotype (CASC), ten other in the publication. males, and 38 females. I have been unable to locate dte holotype of S. chautauquus Contrary to the indication in the Scelolyperu8 meraCU8 (Say) origInal descrIption, thIS speCImen IS apparently (Figure oj) not in the personal collection of John Wilcox. The precedirig treatment is based on the male Galleraca meraca Say, 1826.299. holotype of S. liriophilu8 (NMNH), on two male Luperu8 meraca: LeConte, 1865:209. LuperU8 (Luperu8) meraca: Weise, 1924:120. paratypes of S. chautauquus (OSUC). on one fe- Lupelodes meiUca. Hom, 1893.115, Wilcox, 1954.433. male paratype of S. chautauquus (OSUe), on 212 Scelolyperu8 meracu8: Wilcox, 1965:129, 134, 154. other males, on 465 other females, and on one specImen of unknown sex. Diagnosis. The aedeagus of this species is about as broad in the basal 112 as in the distall/2; Scelolyperus loripes Horn in lateral view, the basal 1/2 is straight and the (Figures 3e, 4j) apIcal 1/2 IS sInuate. These characters distmgUIsh othe~~cies Scelolyperus loripes Horn, HI93'104; Fall, 1901:156; tisfrom ofScelolyperus occurringin Wilcox, 1965:128, 131, 136.

Diagnosis. The pronotum of this species is Type locality. United States. pale, the terminal abdominal tergite ofthe male is Distribution New Brunswick to Georgia to not unusually pIOduced, and each hind tibia of dte male is distinctly arcuate but lacks a prominent o owmg loca ty In ana a~ ew runSWIC: tootb on the inner side at the basal third This Queens Co., 2 mi. N. ofAnnis Lk. combination ofcharacters distinguishes S. loripes I have also examined specimens from the fol­ from otherspecies ofSeelolyperus. Specimens mea lowing counties in dte United States. Gemgia. sure 5.3-5.9 mm long. Rabun Co. Maine: [I have seen specimens from Mt. Typelocality. "OccuISin Califomia,plObably Bigelow whiCh is in Franklin and Somerset Coun- northern." ties]. New Hampshire: Coos Co., Grafton Co. North Distribution. California. I have seen speci­ CaroIiria: Buncombe Co., Haywood Co., Jackson mens from Madera County, CaIiIOrnia AdditiOnal- Co., Macon Co., Swain Go., Yancey Co. Ohio: Ash ly, Fall (1901) reported this species from Tulare tabula Co., Ross Co. Pennsylvania: Allegheny Co., County, California. Columbia Co , Westmoreland Co Tennessee: Sevi- Comments. Thepreceding treatment is based er Co. Vermont: Bennington Co. Virginia: Page Co. on the male lectotype (no. 3800, MCZC; designated West VrrgmIa: Grant Co. by Wilcox, 1965), on four other males, and on three Beyondmaterialthat I haveseen, Wileox (1965) females. The biology ofthis species is unknown. recorded this species from Essex and Rensselaer Counties in New Yllrk and {10m Kansas. Early Scelolyperu8 megaluru8Wilcox recordspublishedbefore aedeagalexaminationwas (Figures 3£; 3g, 4m) deemed critical are of doubtful reliability Scelolyperu8 megaluru8 Wilcox, 1965:128,130, 131,134. Comments. Wilcox (1965) reportedthatBetu- lapopul~7oliaMarsh.(Betulaceae) andHamamelis INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 269 uilginiana L. (llamamelidaceae) ale hosts of this scens was oliginally desclibed flOm specimens col­ species Specimens have been collected from May to lected on the flowers of Juncus (Juncaceae). September. The characters thathave been used to separate The type specimen ofthis species has probably S nigroc)'anfius from S nigrovirfiscfins are minor, been destroyed. I have examined 73 males and 130 insignificant, and somewhat variable. I am corifi­ females. dent that the two 8pecieB are 8ynonymoU8. I have examined the male holotype of Luperus 8celolyperus nigrocyaneus (LeConte) nigrocyaneus (MCZC), the female holotype of Lu­ (Figures 5e, 5f, 5g) perodes nigrovirescens (MCZC), 138 other males, 147 other females, and one speCImen of unknown Luperus nigrocyaneus LeConte. 1879:517; Horn, sex. 1893.106 ( Luperus longulus LeConte, 1857). Luperus (Luperus) nigrocyaneus: Weise, 1924:121. Scelolyperuspasadenae Clark, new species 8celolypel us niglOcyaneus. Vlilcox, 1965.129, 133, 158. (FIgure 1) LupelOdes niglOoitescens Fall, 1910.152, New Synony- my. Scelolypel us nigt OUil escens. Wilcox, 1965:129, 133, 159; Hatch, 1971:205.

Diagnosis. In this species, the prothorax, the ventrolateral carmae m the dIstal fourth of the elytra, mIddle tibiae, and hind tibiae are all dark aedeagus are absent or very 'lleakly developed (fig. and lack met3llic luster Furthermore, the apex of 1). From the other species ofthe genus, S. pasade- the aedeagus is conspIcuously asymmetrIcal, and, nne diffelsin that the pronotum and elytIa ale dalk in lateral view, the aedeagus is sinuate along its and distinctly metallic, the legs are entirely dark, entire length. This combination of characters dis­ the hind tIbIa of the male IS nearly straIght, and tinguishes this from othel species of 8celol ¥pel as. terminal spurs are present on all tibiae ofthe male Specimens measure 3.0-4.3 mm long. Description. Form elongate; prothorax nar­ Type locahtIes. Luperus nLgrocyaneus: Veta lowel than ely tIa. Colol black 01 dark blown, with Pass, Colorado. Luperodes nigrovirescens: Floris blue, green, orpurple luster. Length ofmale 5.5-6.8 sant, Colorado. mm; length of female 5.5-7.1 mm. DistIibutiOIl. BIitish Columbia to Mbelta to He3d met311ic blue or green Vertex we3kly New Mexico to California. I have examined speci­ alutaceous, WIth a broad, mesal ImpreSSIOn adJa­ mens from the fulJowing localities in Canada Al- cent to frontal tubercles; punctures and pubescence berta: Banff; Belly Riv British Colllmbi3' Fernie; largely absent. lnterocular sulcus deep. Interocu- Lizard Creek, Fernie. lar distance equal to 0.6 times maximum width of r have also examined 8pecimens from following ::~ ;~:;:~Y~:e:::~~~s~~:;:~:i:~et:::tl::~:~: counties in the United States. California: Alameda Co, Inyo Co, Mono Co Colorado. Boulder Co., dinal interantennal carina by distinct but rather Chaffee Co., Delta Co., Denver Co., Jefferson Co., shallow sulci, delimited laterally by broad impres­ Mesa Co., Ouray Co., Routt Co , Teller (~o [I h3ve sion. Antenna!fossae sepalatedby a distance slightly also examined specimens from Veta Pass on the greater than width of antennomere I. Interanten- border of Costilla and Huerfano Counties]. Mon­ nal carma well-developed, forming a longitudinal tana. Flathead Co., Gallatin Co., Glacier Co., Mad- ang'llate ridge Genal length subequ31 to width of ison Co., Powell Co. Nebraska: [No specific locality antennomere II. Antennae slender, extendmg to "'3S iildlc3ted} New Mexico' [I have seen a speci- near middle of elytra, black or dark brovm, with men from the Sandia Mts. which are in Bernalillo basal few antennomeres paler brown; antenno- and Sandoval Counties]. Utah: SanJuan Co., Utah Co. \Vyoming. Mbany Co., Palk Co. Beyond specimens that I have seen, Wilcox (1965) reported this species from Larimer, Park and Rio Blanco Counties in Colorado. wide as elytra across humeri; sides arcuate in Comments. Specimens have been collected dOl sal vie w. Latelal beads well-developed, basal from June to September. One was found on Rubus bead small. Surface finely punctate, often aluta- deliciosus Torr. (Rosaceae). Luperodes nigrovire- ceousnearmargins, polishedelsewhere. Color black 270 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI

OI dmkbrown, withmore orlessdistinctgleenorblue NMNH). Santa Cruz Co.. Laurel, June 1893 (1 metallic luster male, ] female, IJMRM); Laurel, 4 .June (2 males, Elytra 1.8 times as long as broad, WIth SIdes asue). nearly parallel. Punctures distinct, separated by a Comments. This species is similar to S. sma distance slightly greater than diameter of a punc­ ragdinus and has been confused with it in the past. ture. Interspaces alutaceous. Color black or dark The specific name, pasadenae, is a leminder that brown, with distinct, metallic blue, green, orpurple many of the specimens in the type series are from luster. near Pasadena, California. Ventral surface brown or black, usually with metallic luster. Prothorax largely glabrous; front 8celolypet us phenacus Wilcox coxae contiguous or nearly so; front coxal cavities (Figure 4h) open behind. Mesothorax alutaceous. Metathorax pubescent. Legs black or dark brown, usually with Scelolyperus phenaclIs Wilcox, 1965'129, 131, 145 weak metallic luster on femora; terminal spurs present on all tibIae of male and of female; male Diagnosis. The pronotum of this species is with basitarsi, especially of front and middle legs, pale, and the elytra lack obvious carinae. Further- dilated; tarsal claws appendiculate. Abdomen pu­ more, the aedeagus is velY slendel and nearly bescent; terminal segment of male impressed dis- s~mmetJ:ic~l;,n ~or~al ~w, an: it is distinctly tally, with a broad, truncate, very short lobe at. apex; termmal segment of female narrowly round­ ed' at apex. Scelolyperus. Specimens measure 3.4-3.9 mm long. Aedeagus (dorsal view) narrowed near middle, Typc loeality. Chester, California. " , Distribution. California. I have seen s eci- central, asymmetrical, angulate tip; sinuate in lat­ pine an eral view Distal fourth lacking strong ventrolater- nia al carinae, sometimes with weak carinae. Basal Comments. ThIS speCIes has been collected spurs absent. OrifIce lackmg sclerotlzed covermg. during July. 80me specimens have been found on Distribution. California. Ribes inerme Rydb. (Saxifragaceae). I have exam- Material examined. Holotype: San Gabriel ined the male holotype (OSUC), eight othel males, Mts., near Pasadena, CalifoIIlia (male, NMNH). and two females Paratypes (all from California): Los Angeles Co.: SCelOly~grUSphoxus Wilcox Devils Can, 12 May 1941, c: Henne (1 male, 'Figures 3j, 4t) AJGC); E. Fork San Gabriel Can., 21 Apr. 1956, R. L. Westcott (2 males, 1 female, AJGC); Glendale, Scelolyperus phoxus Wilcox, 1965:129, 131, 143. March 1932 (l male, OSUO), Glendale, May 1932 (1 male, astra); Hidden Springs, 16 mi. S. Palmdale, Diagnosis. The pronotum of this species is 27 Apr 19811, N J Smith, A ;J Gilbert, collected pale, the hind tibiae of the male are straight or from Leptodactylon californicum (8 males, 4 fe- nearl so and the overall len h is 3.4 to 4.2 mm. , , ur ermore, e ae e us i s u u MCZC); Mt. 'Nilson, 17 June 1939, G. P. Mackenzie ate in lateral view In dorsal view, the apical por- (2 females, VCRV); Mt. Wilson, 7 May 1947, G. P. tion of the aedeagus is distinctly asymmetrical Mackenzie (1 male, 1 female; SEMC); Mt. 'VVilson, although the acute tip is nearly central. These 10 May 1947, G. P. Mackenzie (1 male, VCRV); Mt. characters distinguish this from other species of Wilson, 4 J'lne 1947, GP Mackenzle (1 male, 1 Scelolypel us. female, SEMC); Pasadena, 14 May 1906 (1 male, Type locality Green River Camp, Lower San- CNCI); Pasadena, April (1 male, UMMZ); Pasade­ ta Ana Canyon, California. na, 2 July, Dr. A. Fenyes (3 males, lOCM); Pasade- Distribution. California. I haveexaminedspec- na (1 male, 5 females, NMNH); Pasadena, 3000 Ft., imens from the type locality and from Los Angeles III June 1917 (I female, CNCT); Pasadena, Sister and Riverside Counties in California. Elsie Mt., 10 June 1917 (1 male, 1 female, CNCI); Comments. I have been unable to locate the San Gabriel Mts., near Pasadena (2 males, NMNH, holotype ofthis species. Contrary to the statement 1 male SMCI); [no specific locality], April (1 male, ofWilcox (1965), it is apparentJy not in the Califor- NMNH); [no specific locality], June (2 females, INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 271 nia Academy ofSciences. I have examined six male Latah Co., Lemhi Co., Lewis Co., Minidoka Co., Nez paratypes,twofemale paratypes, sevenothermales, Perce Co, Shoshone Co, Valley Co Montana- Flat- and five other females. One senes was collected head Co., Lake Co., Mmeral Co., Ravalli Co. Ore­ from Adenostoma ,fasciculatum Hook. & Arn. (Ro- gon: Baker Co., Clatsop Co., Coos Co., Hood River saceae). Specimens have been taken from April to Co., Lincoln Co., Morrow Co., Multnomah Co., June. Umatilla Co., Union Co., Wallowa Co., 'Nasca Co. Washington: Asotin Co., Chelan Co., Columbia Co., Seelelyperus ratulus Wilcox King Co., Kittitas Co., Klickitat Co., Lewis Co., (Figures 4i, 4k) Okanogan Co, Pierce Co, Spokane Co, Stevens Co., Walla Walla Co., Whitman Co., Yakima Co. Scelolyperus ratulus Wilcox, 1965:129, 131, 136. The record for \~lyoming by ',Jlilcox (1965) prob ably resulted from a misprinting of Washington. Mso, the IecoId for Baja Califonria by Weise (1924) and the apex of the aedeagus is broadly truncate. ommen s. I have exammed speCimens that These characters will together distinguish this were collected on "Rosa montana" (Rosaceae), and one other was doing damage to roses. Also, I have seenspecimenscollected onRubus pat vi{lOl us Nutl. (Rosaceae) and on goatweed (Hypericum perfora­ tum L., Hypericaceae), but it was unclear whether California, but no specific locality was indicated. or not the beetles were feeding Hatch (1971) re- Connnellts. I have examined only the male ported this species from Berberis (Berberidaceae) holot e CZC and the female allot e CZC . e io ogy 0 is species is un own. collected from March to September. As noted by Wilcox (1965), many of the sped- Scelolype, us schwa,zii HOlU (Figure 4e) mens identified as Luperus variipes by Beller & Hatch (1932) probably belong to thiS speclCS. Scelolj'PeTUs Schwarzz, Horn, 1893·105 I have examined the male holotype (MCZC), Scelolyperus schwarzi: Wilcox, 1965:129, 132, 146. 531 other males, and 619 females. Scdolyperus schwarZ/I" Fall, 19111.156; Hatch, 19'11·2114 Luperus variipes: Beller & Hatch, 1932:114 (misidenti­ Scelolyperus smaragdinus (LeConte) fication) (Figure 4b)

Diagnosis. The dorsal surface of thiS speclCS is Luperus smmagdinus LeConte, 1859c.286, LeCoI1te, entirely metallic, the hind tibia of the male is 1865:209. strongly curved, and the aedeagus is symmetrical Lapel us (Lapel as) smwagdinas. '.Veise, 1924.122. Luperodes smaragdinus: Horn, 1893:116. and emarginate at the apex. These characters en- Scelolypel us Slital agdi7tus. \Vileox, 1965.129, 132, 148. able recognition ofthis species. Specimensmeasure Diagnosis. The ~ronotum and elytra of this 4 0-6 II mm long species are dark and metallic. The legsare entirely Type locality. Hood River Valley, Oregon. dark also Furthermore, the aedeagus is nearly Distribution. British Columbia to Montana to straight in lateral view, and the apical portion has CalifOInia. I have examined specimens from the well developed, ventrolateral carinae. The apex of following localities in Canada. British Columbia: the aedea~us is.pointed. and .distinctly asymmet~i­ Creighton V, TJlmhy; Crest.on, 1900 ft; Kamloops; cal in dorsal View. ThIS sUlte of characteIs will Kustkanook; Lavington; Mara; McCulloch Rd., distinguish this from other species ofthe genus and Kelowna; Robson; Salmon Arm; Tnmty ValIey; ofthe entire section Scelidites. Specimens measure VeInon, Westbank; 'Nhitemans Cr., Vernon. 5.3-6.7 mm long. I have also examined specimens from the fol­ Type locality. Punta de los Reyes, California. lowing counties in the United States. Califomia. Distribution. California. I hll'v'e examinedspec Modoc Co., Monterey Co., Nevada Co., Placer Co., imens from the following counties in the U.S.A. Shasta Co., SiskijrOll (:0, 'I'rimty Co Idaho· Ada California. Alameda Co., Colusa Co., Kings Co., Co., Adams Co., Boise Co., Bonner Co., Canyon Co., Lake Co , Marin Co , Monterey Co , San Luis Obis- Clearwater Co., Gem Co., Idaho Co., Kootenai Co., po Co., Sonoma Co. 272 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI

Beyond material that f have seen, LeConte confluent with orbits. Distance between antennal (1865) reported this species from Mendocino Coun- fossae slightly greater than width of antennomere ty, Cahforma, and WIlcox (1965) reported It from 1. Longitudmal mterantennal carma narrow and Los Angeles and Nevada Counties in California. 'Nell developed in male, poorly developed infemale. However, these records may be based on specimens Genal length subequal to maximum diameter of of s. pasadenae or perhaps other species. distal article of maxillary palp. Antennae slender, Comments. The preceding treatment is based extending to near middle of elytra, testaceous; on the male holotype (MCZC), 31 other males, and antennomere I nearly twice as long as II; III slight­ 14 femaleil ~d'lltil have been collected from March ly longer than II, slightly shorter than IV to May. Some were taken fi-om mustard (BrassI­ Pronotum 1.5 tImes as WIde as long, 0.8 tImes as caceae) and others from the flowers of Ceanothu8 -.vide as elytra across humeri, widest near anterior sp. (Rhamnaceae). third; sides strongly sinuate in dorsal view. Lateral beads well-developed; basal bead small. Surface ScelolyperustejonicusCrotch impunctate, lackingpubescence, shining, not aluta- (Figures 3d, 41) ceous. Color yellow. Elytra 1.7 1.9 times as long as broad, parallel Scelolyperus tejonicus Crotch, HI74:79; Horn, 1893:104; sided or slightly broader near posterior fourth. Wilcox, 1965:129, 130, 134. Punctures separated on average by a distance sub- equal to three times diameter ofa puncture. Inter- Diagnosis The hind tibia ofthe male is strong- spaces alutaceous. Color dark brown with weak ly curved with a prominent tooth on the inner side blue hlilter at the basal third. Such tibiae are not found in any Ventral areas of prothorax testaceous, largely other species ofScelolyperus or ofthe entire section glabrous; front coxae nearly contiguous; front coxal Scelidites. The single specimen examined mea­ cavities open behind. Metathorax brown, pubes­ surp~ 5 0 ~m l~g cent. Femora brown at base, testaceous at apex. Tibiae testaceous; terminal spurs present on all tIbIae of male and female. Frontand mIddle basitar­ rence of this species only in Kern County, Califor- si of male slightly, rather inconspicuously dilated. nia. 'Neise (1924) listed it nom Oregon and Lower Tarsal claws appendiculate. Abdomen brown, pu­ California, but the basis for these records is un- bescent, terminal segment of male with a broad, known to me. truncate, very short lobe at apex; terminal segment Co~nts The ~~ingtreatment ~s:ised of female rounded at apex on the maeholotype, type no. 5070 . ave Aedeagus (fig. 2) slender and nearly symmetri- seen no other specimens. The biology oHms species cal in dorsal view, nearly straight in lateral VIew. is unknown. Basal spurs absent. Orifice lacking sclerotized cov ering. Distribution Wyoming Scelolyperus tetonensis Clark, new species Material examined. Male holotype and fe­ (Figure 2) maIe allotype: 23 mi. sOllth Jackson, Teton Co, Vlyoming, VII 4 53, w. F. Barr Collector (CASC). Diagnosis. The elytra of this species lack dis­ Paratypes: Lincoln Co., Wyo., July 201985, FredA. tinct carinae Also, the aedeagus is slender and Lawson(l male, NMNH, 2 males, 9 females, OSUe), symmetricalin dorsalview, and itis nearly straight Sublette Co., Wyo., 24 July 1980, FredA. Lawson (1 in lateral view (fig. 2). These characters distingUish male, NMNH; 3 females, aSTIe; I male, I female, this from other species of Scelolyperus. SMCI). Description. Form elongate; prothorax nar­ SCelolyperu~orqu~~s rower than elytra. Length of male 3.2-3.7 mm; (LeConte) length of female 3.0-3.3 mm. Head dark brown, shmmg; alutaceous micros- culpture weak or absent. Vertex nearly glabrous. Luperus torquatus LeConte, 1884:28. Luperus (Luperus) torquatus: Weise, 1924:123. Interocular sulcus deep in male, shallow in female. Luperodes torquatus: Horn, 1893:115. lnterocular distance equal to 0.6 times maximum Scelolyperus torquatus: Wilcox, 1965·129, 130, 131, 138 width of head across eyes. Frontal tubercles nearly INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 273

Diagnosis. Inthis species, the pIonotumispale. Lupel us (Lupel us) tl ansitus. Vleise, 1924.123. Also, the aedeagus is nearly straightinlateralview, Scelolyperus transitus: Wilcox, 1965:130, 132, 140. and the apex IS dlstmctly asymmetncal WIth the tIp lateral Theile characterlldillting1lillh thill from other Diagnosis. The pronotum of this species is species of Scelolyperus. SpeCImens measure 2.9-4.7 pale, and the hind tibiae of the male are nearly mmlong. straight Also, the apical portion ofthe aedeagus is Type locality. Specimens in the type series asymmetncal, spatulate, and bent upward, but It were collected at San Mateo and Mariposa, Califor­ lacks an acute or angulate tip (fig. 4a). Specimens nia measure 3.2-4.2 mm long. These characters enable DIstrIbutIOn. Cahforma, ArIzona, Baja Cali­ lecognition of this species. fornia Norte, and possibly British Columbia. I have Type locality Santa Barbara, California seen male specimens from La Rumorosa, Baja Cal­ DistrIbutIOn. Cahforma. I have exammed ifUInia NOI te, Mexico. I have also seen male spec- specimens from Kings, Monterey, and Santa Bar imens labeled from Victoria, British Columbia, bara Counties in California. Beyond material that Canada, but these may be labeled m error. I have seen, Vhlcox (1965) IepoIted this species I have also examined llpecimenll from the fol- from Butte, Contra Costa, Los Angeles, Marin, and lowing counties in the United States. Arizona: San Diego Counties in California, and specimens in Yavapai Co. [this IecoId is based on a female ~e ty~e ll~ri~s ~Bre ~rom San Mateo and Sillkiyon specimen and requires confirmation]. California: ountIes In a forma. Alameda Co., Butte Co., Calaveras Co., Colusa Co., Colllftlents. I have examined specimens taken Contra Costa Co, Fresno Co, Imperial Co, Kern on Ribes divaricatum Dougl. (Saxifragaceae), and Co., Kings Co., Lake Co., Los Arigeles Co., Madera Wilcox (1985) recorded specimens from Ceanothus Co., Marin Co., Mariposa Co., Monterey Co., Napa ClIneatlls (Hock) Nutt (Rbamnaceae) Specimens Co., Orange Co., Riverside Co., San Benito Co., San have been collected from AprIl to June. Bernardino Co., San Diego Co., San Luis Obispo The precedingtreatment isbased onthe female Co., San Mateo Co., Santa Barbara Co., Santa lectotype (no. 3805, MCZC; designated by Wilcox, Clara Co., Santa Cruz Co., Solano Co., Sonoma Co., 1965), 15 otheI females, and 30 males. Tehama Co., Tulare Co., Ventura Co. Scelolyperus varipes (LeConte) Comments. I have examined specimens col­ (Figures 5b, 5d) lected from Sambucus glauca Nutt. (CapIifohace- ae), Arctostapylos sp. (Ericaceae), Quercus sp. (Fa­ gaceae), "tree poppy Dendmmecon rigida?" (Papav- Luperodes varipes. Hom, 1893.110, 118. naceae), Ceanot us mtegernmus 00. n. Scelolyperus varipes: Wilcox, 1965:130, 132, 150; Hatch, (Rhamnaceae), and Califumia slippery elm [Frem- 1971.204. ontodendron californicum (Torr.) Cov.] (Sterculi­ Luperus morrisoni Jacoby, 1888:595, New Synonymy. aceae) However, it is unclear whether or not any of Lupcllts (Calomicllts) :JOII isoni. Weise, 1924.115. these are the normal host plant. I have also seen Luperodes Morrisoni: Horn, 1893:110, 117. specimens taken from the flowerll of ()onvolvulus Scelolyperus rnorrisoni: Wilcox, 1965:129, 132, 151. Luperus concavus Beller & Hatch, 1932:114; Wilcox, (Convolvulaeeae). ,ALdults have been collected from 1965:130 ( Lupcrus varipcs LeConte, 1857). February to July. Vv'ilcox (1985) noted that thele is some vaIiabil- Diagnosis. The dOL sal suzmce ofthis species is ity among specimens that have been assigned to entirel metallic the front tibiae are ale the hind i Iae 0 e ma e are near y s raig , an e is a than one species is involved. portion of the aedeagns is distinctly asymmetrical I have examined the female holotype (MCZC), This combination of characters distinguishes this 435 otheI females, 248 males, and ten specimens of from all other species of Seelolyperus occurring in unknown sex. the western half of North America. Specimens Scelolyperus transitus (Horn) measure 3.4-5.3 mID long. (Figme 4a) "!~ localities ~llperzL: ~="~n ~an-

Luperodes transuus Horn, 1893:113. 274 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI this locality is likely inconecL, the type pIObably theleaves.Also, specimenshave lesscommonly been comes from Arizona) Luperus concavus· "Mt Rain- collected on Haplopappus lineariloliusHe (Asterace- Ier, WashIngton, Greenwater RIver." ae), LLbocedrus decurrens Torr.(Cupressaceae),Arc­ Distribution. British Columbia to Montana to tostaphylos (Ericaceae), Syringa(Oleaceae), pine (Pi New Mexico to California. I have examined speci­ nus sp., Pinaceae), Prunus sp. (Rosaceae), Ptelea mens from the following localities in Canada. AI- crenulata Greene (Rutaceae), and Salix(Salicaceae). berta: Waterton. British Columbia: Creston; En- Itis notclear whetherthese are true hosts or purely denby; Fernie; Howser, Selkirk Mts.; Kaslo; Mad­ incidental records. A small series offive specimens den L, Oliver; Midday Val, Merritt; Mt Revel- wall collected"eatingripelltrawberriellinfield"AdlUts stoke, 5400'; Salmon Arm; Summit Creek, Creston; have beencollectedfi-om March toAugust. Trail; Vlestbank; \Vyndel. &elolyperus uaripes'" ariesslightlyinsize, color, I have also examined specimens from the fol­ punctation, andform ofthe aedeagus. For instance, lowing counties in the United States. Arizona. some specimens from California are larger than Cochise Co , Coconino Co , Graham Co , Maricopa normal and have violaceous rather than the usual Co., Mohave Co., NavaJO Co., "Y. avapal Co. [I have blue or greenelytra. Nevertheless, I believe that an also seen specimens from the Santa Catalina Mts., specimens I have seen belong to a single species. which are in Pima and Santa Cruz Counties]. Furthermore, specimensformerly calledS. morriso­ California. Alameda Co., Butte Co., Calaveras Co., ni fit well within the variability of rather typical Contra Costa Co., ElDorado Co., Fresno Co., Hum- populations. boldt Co., Kern Co., Kings Co., Lake Co., Lassen As pomted out by Wilcox (1965), most of the Co., Los ...A..ngeles Co., Madera Co., Marin Co., Mari llpecimenll identified all Lup€rus variip€s by Beller posa Co., Mendocino Co., Monterey Co., Napa Co., and Hatch (1932) are probably Scelolyperus Nevada Co., Placer Co., Plumas Co., Riverside Co., schwarzii. San Bernardino Co., San Diego Co., San Francisco During this study, I have examined several Co, San Mateo Co, Santa Barbara Co, Santa syntypes of Luperus morrisoni (BMNH, MCZC). Clara Co., Santa Cruz Co., Shasta Co., Siskiyou One of the specimens from the British Museum Co., Sonoma Co., Tehama Co., TrImty Co., TUlare (Natural HIstOry) bears Jacoby's "type" label and IS Co., Tuolumne Co., Ventura Co. Colorado: Boulder hereby designated as lectotype. The original type Co., Denver Co., Douglas Co., Grand Co., Jefferson series included specimens ofEusattodera intermix­ Co., Larimer Co. [I have also seen a specimen flOm tus (Fall) and E. piILi Schaeffer, as well as other Veta Pass which is on the border of Costilla and specimens that are conspecific with the lectotype. Hllerfano COllntlell] Idaho· BOIse Co, Bonneville My concept ohms species is based on the male Co., Custer Co., Franklin Co., Idaho Co., Kootenai holotype of Luperus varipes (MCZC), the male Co., Latah Co., Lewis Co., Shoshone Co., Valley Co. lectotype of Luperus morrisoni (BMNH), the fe­ Montana. Flathead Co., Missoula Co., Ravalli Co., male holotype of Lupel us cOILcavus (NMNII), 533 Sanders Co. Nevada: Elko Co., Washoe Co. New other males, 783 other females, and six specimens Mexico· Grant Co, San Miguel Co ()regoO" Baker of unknown sex Co., Benton Co., Coos Co., Curry Co., Douglas Co., Grant Co., Hood River Co., Jackson Co., Josephine Scelolypet us wilcoxi HatclI Co., Klamath Co., Lake Co., Lane Co., Lincoln Co., Multnomah Co., Umatilla Co., Union Co., Wasco Scelolyperus wilcoxi Hatch, 1971.204. Co., Vvashington Co., Yllmhill Co. Utah. Salt Lake Diagnosis. In this species, the pronotum and Co., Sanpete Co., Utah Co., Wasatch Co. Washing­ dis~in~ ~e tibi~e ton: Chelan Co" Jefferllon Co., King Co, Kittitas elytia are me:;llic, ani fr:t Co., Pierce Co., Spokane Co., Walla Walla Co., Whitman Co., Yakima Co. Wyoming: Carbon Co. Conullents. Many of the beetles I have exam- characters will t9;etherdisti:uish this from other species ofSceloly erus. Sped ens measure 4.4-5.7 ined were collected from various species of Cean­ mm~ong othu8 (Rhamnaceae) CWtnothus J:n tegerdwusHook & Arn., C. sanguineus Pursh, C. thyrsiflorus Eschsch., and G. velutinus Doug!. have specifically 18." been mentioned. However, it is unclear whether the beetleswereinthe flowers oractuallyfeeding on INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 275

DistIibution. Olegon. I have examined speci­ [VlSUC], Charles C. Coffman, 'vVest\1:1 giniaDepalt­ mens from Benton, Jefferson, Josephine, and Kla- mentofAgriculture [WVDA]. math Countles m Oregon. I acknowledge Laura Torres-Miner, who pre­ Comments. The treatment of this species is pared aedeagal illustrations for the two new spe- based on the male holotype (NMNH), on 14 other des. Also, I extend special thanks to Charles A. males, and on eight females. Specimens have been Triplehorn who advised throughout much of this collected from May to July. Some have been taken study. from Phacelia (Hydrophyllaceae). I gratefully thank John A. Wilcox. He kindly granted permission to reproduce i1lustrations from Acknowledgments hIS 1965 pubhcatIOn. Even more Importantly, he I express my appreciation to the following indi- generously shared his abundant expertise of the vlduals and mstltutIOns that have loaned speci­ Galerucinae. mens (the acronyms following the entries have Finally, I acknowledge Edwald G. Riley and C. been referred to throughout this paper; my person- L Staines, Jr. Their reviews ofan earlier version of al collection has been referred to by the acronym the manuscript SIgnifICantly Improved thIS paper. SMCI): ArthurJ. Gilbert, Clovis, California [AJGCl; Sharon L. Shute, The Natural History Museum References (London) [BMNH]; Richard W Baumann and Arnett, R. H, 1962. The Beetles ofthe United States (A Stephen L. Wood, Brigham Young Umverslty Manual for Identification), xiii + 1112 pp. Washing­ [BYUC]; David H. Kavanaugh, California Acade ton, D.C., Catholic University of America Press. m of Sciences CASC; Robert S. Anderson, Cana- Beller, S., and M. H. Hatch. 1932. Coleoptera of i , ington Publications in Biology 1: 65-144. ~a~ad~n Na~oDAl ~ogp~tion of ~n:c~ ~CNC~]; Blake, D. II. 1928. A new species ofLupelodes fiom Mt. Oris. Ron ratief; a oradoS amverSlY Mitchell, North Carolina. Bulletin of the Brooklyn [CSUC]; Cornell University [CUIC]; Edward G. Entomological Society 13: 183 184. Riley, College Station, Texas [EGRC]; John B. Brancsik, C. 1899. Aliquot Coleoptera nova Russiae Ke thley, FieldMuseumofNatmalHistory [FMNII], asiaticae. Jahl'esheft Natlll'Wisse des Vel'eins ZIl Robert E. Woodruff, Florida State Collection of Trencsen 21: 97-105. Arthropods [FSCA]; Robert Davidson and John E. Clark, So M. 1987. .'>. Revision of the Section Scelidites in theWesternHemisphere (Coleoptera: Chrysomel­ Musell~ ~itural ~istor~ Ra',vlins, Carnegie of idae). Unpublished Ph.D. dissertation, Department [ICCM]; Kathryn C. McGI'en, inois atura of Entomology, The Ohio State University, 397 pp. HiStOly Sm vey [INIIS], John A. Vlilcox, Lancaster, Crotch, G R 1873 Materials for the study of the Ohio [JAWC]; Joseph F. Cavey, Eldersburg, Mary- Phytophaga ofthe United States Proceedings ofthe land [JFCC]; Michael A. lvie, Bozeman, Montana Academy of Natural SClences ofPhIladelphIa 25: 19­ [MAle]; Alfred F Newton, Jr and Scott Shaw, 83 Museum of ComparatIve Zoology [MCZC]; M. A. Crotch, G. R. 1874. DescnptlOns of new speCles of Ivie, Montana State University [MTEC]; Ed'Nard Coleoptera from the Pacific Coast of the United U. Balsbaugh, Jr., North Dakota State University States. Transactions of the American Entomologi­ [NDSU]; David G. FUlth and Richald E. '.Vhite, cal Societv 5: 73-80. Crotch, G. R. 1880. Check List of the Coleoptera of urn NMNH . Charles chusetts. , ertH. Turnbow, Jr., FortRucker, Alabama [ HT California, with notes on habits and distribution Richard S. Millel, Bozeman, Montana [RSMC]; and descriptions of new species. Occasional Papers James Pakaluk, University of Kansas [SEMC]; of the California Academy of Sciences 8: 1-282. Horace R. Burke and Edward G. Riley, Texas A. & Fall, H. C. 1910. Miscellaneous notes and descriptions M University [TAMU]; Sydney G Cannings, IIni- of North American Coleoptera. Transactions of the versity of BritIsh ColumbIa [OBCZ]; Saul I. From­ American Entomological Society 36: 89 197. Fattig, P. W, 1948. The Chrysomelidae or leafbeetles of meI', Unrlersity ofCalifornia, Riverside [UCR'A; F. Georgia Emory I lniversity Museum Bulletin 6' 1. Merickel, University of Idaho [DICM]; Mark F. 47. O'BIien, UnivelsiLy ofMiclrigan [UMMZ1; Robelt L. Fabricius,.1 «: 1787 Mantissa JDsectoTuID, vol 1, 348 Blinn, University of Missouri-Columbia [UMRM]; pp. Hafniae. RIchard S. Zack, Washmgton State University 276 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI

Goeze, J. A. E. 17'7'7. Entumulugische BeitIage zu des als schadlich udeI nutzlich bekannt gewuldenen In- Ritter Linne 12. Ausgabe des Natursystems, vol. 1, sekten ..., vol. 1, Die Kafer, 202 pp. Berlin. 736 pp. Leipzig. Reid, C. 1992. Making full use of intemal abdominal Gyllenhal, L. 1813. Insecta Svecica descripta. Classis characters. Chrysomela 27:2. 1. Coleoptera sive eleuterata 1: 1 760. Riley, E. G., and IN. R. Enns. 1979. An annotated Hatch, M. H. 1971. The beetles of the Pacific North­ checklist ofMissouri leafbeetles (Coleoptera: Chry­ 'Nest. Part V: Ripiceroidea, Sternoxi, Phytophaga, somelidae). Transactions of the Missouri Academy Rhynchophora, and Lamellicornia. University of of Sciences 13: 53 83. Washington PubhcatIons m BIology 16: 1-650. Riley, E. G., and W. R. Enns. 1982. Supplement to an Henshaw, S 1895. List of the Coleoptera of America, annotated checklist of Missouri leaf beetles (Co- North of MeXICO, 161 pp. PhIladelphIa. leoptera: Chrysomehdae): new state records and Horn, G H. 1893 The Galerucini of boreal America host plant associations Entomological News 93' 32- TransactIOns of the Amencan EntomologIcal SOCI­ 36. ety 20' 57-136 Say, T ]826 Descriptions of new species of coleopter- Horn, G. H. 1895. Coleoptera of Baja Califorma (sup­ ous mseets mhabltmg the Umted States. Journal of plement I) Proceedings of the California Academy the Academy of Natural Sciences of Philadelphia of Sciences (Series 2) 5: 225-259. 5(2): 237-284, 293-304. Jacoby, M. 1888. Biologia Centrali-Americana, Insec- Schaeffer, C. F. A. 1932 Notes on some Galerucinae ta, Coleoptera, Galerucidae, vol. 6, pt. 1, pp. 585­ with descriptions of new species (Col., Chrysomel­ 625. idae). Canadian Entomologist 64. 236-239. LeConte, J. L. 1857. Report upon insects collected on Scopoli, J. A. 1763. Entomologia Carniolica exhibens the sUlvey. (Repotts ofexplotations and sUlveys fot insecta Camioliae indigena et distIibuta in oIdines, a railroad route from the Mississippi River to the genera, species, varietates, methodo Linneana, 420 Pacific Ocean.) 72 pp. Washington. pp. TIattner. LeConte, J. L. 1859a. The Coleoptera of Kansas and Selander, R. B., and P. Vaurie. 1962. A gazetteer to eastern Ne", Mexico. Smithsonian Contributions to accompany the "Insecta" volumes of the "Biologia Knowledge 2: 1·58. Centrali-Americana." American Museum Novitates LeConte, J. L. 1859h. Catalogue of the Coleoptera of 2099: 170. Fort Tejon, California. Proceedings of the Academy Smith, E. H. 1979. Techniques for the dissection and of Natural Sciences of Philadelphia 11: 69-90. mounting of the male (aedeagus) and female (sper­ LeConte, J. L. 1859c. Additions to the coleopterous matheca) genitalia of the Chrysomelidae (Co- fauna of northern California and Oregon. Proceed­ leoptera). Coleoptensts BulIetlll 33: 93-103. ings olthe Academy ofNat'lraI Sciences ofPhiladel- Weise, J 1924 Coleopterorum catalogus, Chrysome1- phla 11: 281-292. Idae: Galeruclllae, pars '/8, pp. 1-225. [Vol. XXV.] LeConte,.J L ]865 On tbe species of Galeruca and Wilcox, J A 1954 Leaf beetles of Obio (Cbrysomel- allIed genera mhabltlllg North Amenca. Proceed­ Idae: Coleoptera). OhIO BIOlogIcal Survey Bulletm ings oHbe Academy ofNatural Sciences ofPbiladel- 8(3)' 353-5116 phia 17: 204-222. Wilcox, J. A. 1965. A synopsis of the North American LeConte, J. L. 1879 The Coleoptera of the alpine Galerucinae (Coleoptera: Chrysomelidae). Bulletin Rocky Mountain regions. Part II. Bulletin of the ofthe New York State Museum and Science Service United States Geological and Geographical Survey 400: i-iv, 1-226. of the Territories 5: 499-520. Wilcox, J. A. 1973. Coleopterorum Catalogus Supple­ LeConte. J. L. 1884. Short studies of North American menta, Clnysomelidae. GaieI ucinae (edilio segun- Coleoptera (No.2). Transactions of the American do), pars 78, fasc. 3, pp. 433-664. Entomological Society 12. 1-32. \Vilcox, J. A. 1979. Leaf Beetle Host Plants in NOlth­ Ogloblin, D. A. 1936. Fauna de l'URSS. Insectes, Co- eastern North America, 30 pp. Kinderhook, New leopteres, Chrysomelidae, Galerueinae, vol. 26, No. York. 1, pp. i-xiv, 1-455. Ratzeourg, J. T. C. 1837. Die FOl'stinsekten oderl'.bhil dung und Beschreibung der in den Nachbarstaaten INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 277

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Figure 2. &elolyperus tetonensis new species, median lobe ofaedeagus, dorsal, lateral, and ventral views. 278 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI

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Figure 3.8eelelypernsSJlIl. a) 8. liFiephilullWiloox; b) 8. euroipeII'1lileox, dorsalantllateralVHlWS efaedeagus; e) 8. eurtJipeil Wiloox; d) S. teJonlCus crotch, posterior femur and tibia oflnllIe; e) S. 'oripell Horn, posterior femur and tibia ofDlllIe; t) S. megaiurull Wilcox, posterior tibia and tarsus ofmale' lr) S. megaluTus Wilcox, apical segments ofmale abdomen, ventrolateral view; h) S, haremWilcox; i) S. carinatus Wilcox, female; j) S. phoxus Wilcox (all figures from Wilcox, 1965). . INSECTA MUNDI, Vol. 10, Nos. 1-4, March - December, 1996 279 .... ~ ,..... ""'" -= I ' / I\ ?~ V- /'\1 ... ,...... aT \ l'r, ('" It: H ~ ':to \ l' III , ,.. ' L-- p"\I ~ :1' lH (,., U i ~ ... V , ~ ~ ~ \ , - v ~ u --" .1 .- 11 \ " ~"' 1\\- , ~ I - ,...... , (Y ,- ''\ Y\ ~ :- ~ ~ ~ I.... , ~7 1\ il '" T ...i 1 II \ 1 ~I I I II-. p( _:J11. \ ) I / J /IA I f n I h If J 'iiii/T \.-IV \J , \II( J tJ -. - () -- ~ "'"'- 1"'7 1:( " ,"• J . ,/\ ) «( /( f\ l\ i r/-'~ ,- A .. I 1/11 !J~~ I J : ":.' f.. ~1 ~I Ii; /\ J ~8 " 'ifJI ~ ,ii' 7"" 'Y ~ r. ~ ;';,: l-i I/ ..." - I 1'1) \\ II v £ ) ;/ i ( }., , t&... ..,~ (/W\ ...... • •••

Figure 4. Aedeagi of&elolyperus spp. a) S. transitus (Horn), dorsal andlateral views; b) S. smaragdinus (LeConte), dorsal view; e) 8. flavieoUis (LeCente), del'sal andlateral'1iews; d) S. terquatus (LeConte), del'sal andlateral '1ieY's; e} S. sGhwa,.zi Hern, dorsal and Iater8Iviews; f) S. p1lOrii8 Wilcox, dors8I and Iater8IViews; g) S. latlCeps (Horn), dors8I andIater8I Views; h) S. phimacus Wilcox, dors8I andlateralviews; i) S. ratulusWilcox, dorsalandlateralviews; j) S. loripesHorn, ventralview ofapex; k) S. ratulusWilcox, ventralview of apex; 1) S. tejonicus Crotch, dorsal view; m) S. megalurus Wilcox, dorsal and ventral views (all figures from Wilcox, 1965). 280 Vol. 10, Nos. 1-4, March-December, 1996, INSECTA MUNDI

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Figure e.•\edeagi of&elelypem8 SJlfl. a) 8. cYGne""" (LeCante), deIosal andlatel'al views; b) 8. IIGnpea (LeCante), del'sal view; e) S.graptoderoJiJea (Crotch), dorsalView; d) S. van-pea (LeConte), dorsalandlateralview; e-g) S. nigrocyaneua(LeConte),dorsalandlateral views; h) S. hatcNWilcox, dorsal and lateral views; i) S. bimarginatu8 (Blake), dorsal and lateral views; j) S meracu8 (Say), dorsal and lateral views; k.l) S. liriophilua Wilcox, dorsal andlateralviews (all figures from Wilcox; 1965).