WTU Herbarium Specimen Label Data Generated from the WTU Herbarium Database September 29, 2021 at 6:37 pm http://biology.burke.washington.edu/herbarium/collections/search.php

Specimen records: 1251 Images: 334

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Label Query: Genus = "" Rosaceae Rubus parviflorus Nutt. Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, WHATCOM COUNTY: Ross Lake National Recreation Area: Little Ross Lake National Recreation Area: About 1/4 mile down from Beaver Creek trail. Along trail on east-facing Ross Dam from Happy Flats Trailhead. slope, within 0.5 miles of trailhead (before Elev. 1640 ft. switchbacks that take trail northwest and finally UTM Zone 10, 641674E, 5399158N; NAD 83, Source: west). Georeferenced, Georef'd by WTU Staff Elev. 1589-2037 ft. Douglas-fir overstory. Along tight switchbacksof trail; dry, rocky soil. 48° 55.014' N, 121° 4.642' W; UTM Zone 10, Phenology: Flowers. Origin: Introduced. 640860.997E, 5419780.74N; Source: Field notes, UTM calculated from Lat-Lon. Michael Brondi 02-1 12 Jul 2002 Furthest point west of lake. Douglas fir forest; along edge of trail. In ; meter tall ; Herbarium: NOCA, NPS accession 632, catalog 22204 common in area. Phenology: . Origin: Native. Rosaceae Sarah Gage 7266 13 Jul 2002 Rubus idaeus L. ssp. strigosus (Michx.) Focke WTU-360226, WTU-V-045610, NPS accession 632, catalog 22495 U.S.A., WASHINGTON, WHATCOM COUNTY: Ross Lake National Recreation Area: Mountain Powerplant Overlook Rosaceae off of State Route 20. Elev. 984-1640 ft. Rubus leucodermis Douglas ex Torr. & A. Gray 48° 42' 58.69646" N, 121° 4' 32.46619" W; UTM Zone 10, 641548E, 5397487N; Source: Calc. from UTM, UTM from field U.S.A., WASHINGTON, WHATCOM COUNTY: notes. Ross Lake National Recreation Area: Little Beaver Creek trail. Within South side of highway 20 above milepost 133; dried moss-encrusted 0.5 miles of trailhead (before switchbacks that take trail northwest ledges overhanging highway, talus slope, and rocky outcrop with and finally west). Douglas fir and Acer circinatum. Sparse, occasional; less than 15 Elev. 2035 ft. inches on talus slope; flowers white. Elevation estimated from map. 48° 55.014' N, 121° 4.642' W; UTM Zone 10, 640860.997E, Phenology: Flowers. Origin: Introduced. 5419780.74N; Source: Field notes, UTM calculated from Lat-Lon. Furthest point west of lake. South-facing, rocky slope along trail. Robert Goff 02-21 16 Jun 2002 Young stems with white bloom on stems; in fruit. Phenology: Fruits. Origin: Native. NOCA-360261, NPS accession 632, catalog 22576 Sarah Gage 7277 13 Jul 2002 Rosaceae WTU-360248, NPS accession 632, catalog 22519 Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., WASHINGTON, CHELAN COUNTY: Rosaceae North Cascades National Park; Horseshoe Basin; just below waterfalls next to snowfield on west side of basin. Rubus idaeus L. ssp. strigosus (Michx.) Focke Elev. 4772 ft. 48° 28' 42.7" N, 121° 1' 24.6" W; UTM Zone 10, 646070.532E, U.S.A., WASHINGTON, WHATCOM COUNTY: 5371156.543N; Source: Field notes, UTM calculated from Lat-Lon. Little Beaver Trail, about 1 mile east of Perry Creek; North Cascades Rocky slope with very little soil; open with a few short trees (Alnus National Park. sp., Sorbus sp., Chamaecyparis nootkatensis); lots of Valeriana sp. Elev. 2034 ft. in bloom. Scattered about 3 to 4 feet tall. Phenology: Flowers. 48° 55.3' N, 121° 8.1' W; UTM Zone 10, 636625.715E, Origin: Introduced. 5420205.349N; Source: Field notes, UTM calculated from Lat-Lon. Dry gneiss talus, adjacent Douglas-fir forest, and adjacent cold Dena Grossenbacher 02-39 10 Aug 2002 stream bank just below talus; southern aspect. Flowers white; old stems purple; small . Phenology: Flowers. Origin: Introduced. Herbarium: WTU, NPS accession 632, catalog 22683 Ben Legler 14 13 Jul 2002 Rosaceae Herbarium: NOCA, NPS accession 632, catalog 22811 Rubus parviflorus Nutt. Rosaceae U.S.A., OREGON, WALLOWA COUNTY: Wallowa-Whitman National Forest. Lostine Rubus spectabilis Pursh Canyon, Bowman Trail, where trail first crosses a feeder creek. U.S.A., WASHINGTON, WHATCOM COUNTY: Elev. 6689 ft. Little Beaver Trail, about 1 mile east of Perry Creek; North Cascades 45° 17.289' N, 117° 24.645' W National Park. Dry, north-facing outcrop; open area, break in Elev. 2034 ft. trees. Granite cliff and talus. Petals white. 48° 55.3' N, 121° 8.1' W; UTM Zone 10, 636625.715E, Common along trail. Altimeter reading 6450 ft. 5420205.349N; Source: Field notes, UTM calculated from Lat-Lon. Phenology: Flowers. Origin: Native. Dry gneiss talus, adjacent Douglas-fir forest, and adjacent cold stream bank just below talus; southern aspect. Fruit orange-red. Sarah Gage 7339 Phenology: Fruits. Origin: Native. with Don Knoke, Dale Blum, Richard Robohm Ben Legler 29 13 Jul 2002 21 Jul 2002 Herbarium: WTU WTU-360574, WTU-V-045445 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus arcticus L. U.S.A., WASHINGTON, PACIFIC COUNTY: U.S.A., WASHINGTON, OKANOGAN COUNTY: The Nature Conservancy's Ellsworth Creek preserve, 5 miles NW of [locality and coordinates withheld; status in Washington: G5T5/S1 Naselle on east side of mouth of creek, under powerlines. (Threatened; WANHP)] Elev. 100 ft. Elev. 6300 ft. 46° 24' 58.2" N, 123° 53' 8.9" W Growing on damp ground. Associates: lone Picea engelmannii, Salix Open area with grasses and weeds. Petals white. Phenology: species, Carex species, sphagnum and other mosses. Phenology: Flowers. Origin: Native. Flowers. Origin: Native. Ben Legler 490 31 May 2003 Elroy Burnett 35 20 Jun 1988 Herbarium: WTU WTU-315163

Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus bifrons Vest U.S.A., WASHINGTON, THURSTON COUNTY: U.S.A., WASHINGTON, PACIFIC COUNTY: 2 miles southwest of Yelm, along bike trail (old railroad track) East side of Sandridge Road, near Black Lake. paralleling NW side of Hwy 507 just south of Manke St. Elev. 16 ft. Elev. 400 ft. T10N R11W S28 NW 1/4 of SE 1/4; NAD 27, uncertainty: 805 m., 46° 54' 45.3" N, 122° 38' 52.1" W Source: TRS2LL, Georef'd by Ben Legler Built up railroad bed through wet forest of Fraxinus latifolia, Pinus Ditch. Weed in cranberry fields, with Vaccinium macrocarpon, Viola contorta, Pseudotsuga menziesii. Petals white, 6 mm long; stems lanceolata. Phenology: Fruits. Origin: Introduced. trailing. Phenology: Flowers. Origin: Native. P. F. Zika 14182 31 Aug 1999 Ben Legler 409 18 May 2003 WTU-340173 WTU-358509 Rosaceae Rosaceae Rubus bifrons Vest Rubus spectabilis Pursh U.S.A., WASHINGTON, CLALLAM COUNTY: U.S.A., WASHINGTON, SKAGIT COUNTY: Quillayute Prairie. Okanogan National Forest. North Cascades 47.94333°, -124.54111°; WGS 84, uncertainty: 10000 m., Source: Highway 20. Easy Pass Trail Boardwalk. GeoLocate, Georef'd by WTU Staff Elev. 3756 ft. Phenology: Fruits. Origin: Introduced. 48° 35.27' N, 120° 48.144' W; T36N R16E S25; uncertainty: 9 m. George Neville Jones 3783 Aug 1931 Bottomland with: Mountain Hemlock, Western Red Cedar, Abies amabilis, Thuja heterophylla, WTU-21620 Tsuga heterophylla, Trisetum spicatum, Viburnum edule, Athyrium filix-femina, Rubus spectabilis, Rubus parviflora, Epilobium Rosaceae latifolium, Vaccinium, Glyceria, Viola, Trillium, Clintonia, and Dryopteris. (GPS 3-4 sat.); Rubus bifrons Vest Accuracy 26-30'. Phenology: Fruits. Origin: Native. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Hoquiam. Ken Davis 03-39 27 Jul 2003 46.98111°, -123.88806°; WGS 84, uncertainty: 10000 m., Source: with Richard Robohm, Sarah Gage, Robert Goff, Margarite Goff, GeoLocate, Georef'd by WTU Staff Dale Blum Rocky low areas. Common. Phenology: Fruits. Origin: Introduced. WTU-362401, WTU-V-039429 Delzie Demaree 60908 22 Aug 1969 Rosaceae WTU-274576 Rubus parviflorus Nutt. Rosaceae U.S.A., WASHINGTON, SKAGIT COUNTY: Okanogan National Forest. North Cascades Highway 20. Easy Pass Rubus bifrons Vest Trail Boardwalk. Elev. 3756 ft. U.S.A., WASHINGTON, ISLAND COUNTY: 48° 35.27' N, 120° 48.144' W; T36N R16E S25; uncertainty: 9 m. Hoypus Point, Deception Pass State Park, Whidbey Island. Bottomland with: Mountain Hemlock, Western Red Cedar, Abies 48.39389°, -122.6475°; WGS 84, uncertainty: 10000 m., Source: amabilis, Thuja heterophylla, Tsuga heterophylla, Trisetum GeoLocate, Georef'd by WTU Staff spicatum, Viburnum edule, Athyrium filix-femina, Rubus spectabilis, Roadside. Phenology: Flowers & Fruits. Origin: Introduced. Rubus parviflora, Epilobium latifolium, Vaccinium, Glyceria, Viola, Trillium, Clintonia, and Dryopteris. (GPS 3-4 sat.); Accuracy 26-30'. Harold W. Smith 1666 31 Jul 1936 Phenology: Flowers & Fruits. Origin: Native. WTU-40536 Ken Davis 03-44 27 Jul 2003 with Richard Robohm, Sarah Gage, Robert Goff, Margarite Goff, Dale Blum WTU-362067 Rosaceae Rosaceae Rubus bifrons Vest Rubus bifrons Vest U.S.A., WASHINGTON, KLICKITAT COUNTY: U.S.A., WASHINGTON, YAKIMA COUNTY: Northwest of Bingen. At Parker. 45.715°, -121.46333°; WGS 84, uncertainty: Elev. 705 ft. 10000 m., Source: GeoLocate, Georef'd by 46.50111°, -120.46417°; WGS 84, uncertainty: 5000 m., Source: WTU Staff GeoLocate, Georef'd by WTU Staff Roadside. Phenology: Flowers & Fruits. Origin: Upper sonoran. Phenology: Flowers & Fruits. Origin: Introduced. Introduced. E. Hunn 622 5 Jul 1977 Wilhelm N. Suksdorf 12015 WTU-270931 29 May 1926 WTU-197552, WTU-V-060716 Rosaceae Rubus idaeus L. Rosaceae U.S.A., WASHINGTON, OKANOGAN COUNTY: LoupLoup campground. Rubus bifrons Vest Elev. 4002 ft. 48.38881°, -119.88934°; WGS 84, uncertainty: 500 m., Source: : Georeferenced, Georef'd by WTU Staff Canadian meadow edge. Phenology: Fruits. Origin: Unknown. Elev. 0-0 Rich deep organic soil in drained bog. Widespread in city; Flowers E. Hunn 755 19 Aug 1977 white. Phenology: Flowers & Fruits. Origin: Introduced. WTU-271083 T. F. Johnson 143 9 Jun 1987 WTU-328304 Rosaceae Rosaceae Rubus idaeus L. Rubus bifrons Vest U.S.A., WASHINGTON, PIERCE COUNTY: Longmire. Mount Rainier. U.S.A., WASHINGTON, KITSAP COUNTY: 46.75°, -121.81139°; WGS 84, uncertainty: 10000 m., Source: Bainbridge Island. GeoLocate, Georef'd by WTU Staff Elev. 0 ft. Moist soil, edge of swamp. Phenology: Fruits. Origin: Unknown. 47.65528°, -122.51667°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff F. A. Warren 1672 31 Aug 1932 Sea level. Phenology: Flowers & Fruits. Origin: Introduced. WTU-26458 Mae E. Cornelius 41 13 Jun 1948 WTU-148546 Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rosaceae U.S.A., WASHINGTON, STEVENS COUNTY: Rubus bifrons Vest Along Columbia River just west of Flat Creek. Above 1290 ft. level. 48.82359°, -117.97836°; WGS 84, uncertainty: 1000 m., Source: U.S.A., WASHINGTON, CLALLAM COUNTY: GeoLocate, Georef'd by WTU Staff Near Blyn. Along road through woods. Shrub with drooping branches up to 2 Elev. 30 ft. meters long. Phenology: Flowers. 48.0225°, -123.00472°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff H. T. Rogers 5556 29 May 1940 Phenology: Fruits. Origin: Introduced. WTU-82979 George Neville Jones 8504 5 Sep 1935 WTU-31536 Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rosaceae U.S.A., WASHINGTON, KLICKITAT COUNTY: Rubus bifrons Vest Garden, Bingen. 45.715°, -121.46333°; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, WHITMAN COUNTY: GeoLocate, Georef'd by WTU Staff 13.5 miles west of Lewiston. (from the riverbank). Phenology: Fruits. Cultivated. T11N R45E S5; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Wilhelm N. Suksdorf 8271 1 Jul 1903 Along small stream from side canyon. Forming dense thickets to 10 ft. tall. Phenology: Flowers. Origin: Introduced. WTU-197547 C. L. Hitchcock 21858 11 Jun 1959 with C. V. Muhlick WTU-195135 Rosaceae Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rubus laciniatus Willd. U.S.A., WASHINGTON, SPOKANE COUNTY: U.S.A., WASHINGTON, JEFFERSON COUNTY: Spokane. Vicinity Fisher's Ranch - Hoh River, Olumpic Forest. 47.65889°, -117.425°; WGS 84, uncertainty: 1000 m., Source: Elev. 250 ft. GeoLocate, Georef'd by WTU Staff 47.814011°, -124.176698°; WGS 84, uncertainty: 10000 m., Along streams in woods. Indefinite loan by Tacoma High School to Source: GeoLocate, Georef'd by WTU Staff University of Washington. Phenology: Fruits. Level slope in sandy loam. Common in fields. Height 2 - 8 ft. Phenology: Flowers & Fruits. Origin: Introduced. C. V. Piper 2268 2 Jul 1896 J. E. Schwartz 94 A 8 Jul 1936 WTU-33513 WTU-322835 Rosaceae Rosaceae Rubus sachalinensis H. Lév. var. sachalinensis Rubus laciniatus Willd. U.S.A., WASHINGTON, CHELAN COUNTY: Rainbow Creek trail at Bench Creek. U.S.A., WASHINGTON, KING COUNTY: 48.379651°, -120.688234°; WGS84, uncertainty: 5000 m., Source: Seattle: University of Washington, west campus parking lots and GeoLocate, Georef'd by WTU Staff margins. Rocky banks near creek. Phenology: Fruits. Origin: Unknown. 47.39361°, -122.42361°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff George H. Ward 120 22 Aug 1942 Warm, early area near canal. Flowers white; this species less common than R. discolor and outcompeted by it. Phenology: WTU-182718 Flowers & Fruits. Origin: Introduced. T. F. Johnson 159 17 Jun 1987 Rosaceae WTU-328260 Rubus sachalinensis H. Lév. var. sachalinensis U.S.A., WASHINGTON, STEVENS COUNTY: Rosaceae Kettle Falls. 48.61083°, -118.05472°; WGS 84, uncertainty: 10000 m., Source: Rubus laciniatus Willd. GeoLocate, Georef'd by WTU Staff At the falls. Phenology: Flowers. Origin: Unknown. U.S.A., WASHINGTON, MASON COUNTY: Union. Carl H Spieg 322 26 May 1923 47.35528°, -123.09972°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-28858 Phenology: Fruits. Origin: Introduced. George Neville Jones 8538 5 Sep 1935 Rosaceae WTU-31818 Rubus laciniatus Willd. U.S.A., WASHINGTON, CLALLAM COUNTY: Rosaceae Quillayute Prairie. 47.94333°, -124.54111°; WGS 84, uncertainty: 10000 m., Source: Rubus laciniatus Willd. GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Origin: Introduced. U.S.A., WASHINGTON, PIERCE COUNTY: Longmire, Mount Rainier. George Neville Jones 3782 Aug 1931 46.75°, -121.81139°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-21691 Rocky soil. Phenology: Flowers & Fruits. Origin: Introduced. F. A. Warren 1573 26 Aug 1932 Rosaceae WTU-26116 Rubus laciniatus Willd. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Rosaceae Montesano. 46.98139°, -123.60139°; WGS 84, uncertainty: 10000 m., Source: Rubus laciniatus Willd. GeoLocate, Georef'd by WTU Staff Phenology: Flowers & Fruits. Origin: Introduced. U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Island; around Egg Lake, ca. 3.5 mi directly NW of Friday George Neville Jones 8405 9 Sep 1935 Harbor;. 48.567222°, -123.081667°; WGS 84, uncertainty: 200 m., Source: WTU-31816 Georeferenced, Georef'd by WTU Staff Locally abundant; petals white;. Phenology: Flowers & Fruits. Origin: Introduced. Melinda F. Denton 3520 2 Aug 1974 WTU-259536 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus lasiococcus A. Gray U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, MASON COUNTY: King (205th and Alaska Way)/Snohomish Counties. Undeveloped Along the trail above Sindown Lakes on the way to Six Ridge Pass. property on the north border of LFP, 10 yards in from 205th. Elev. 4000 ft. 47.91306°, -122.09694°; WGS 84, uncertainty: 10000 m., Source: 47.56°, -123.42528°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Along side the trail with various trailing Rubus sp., Veronica Western exposure-open meadow habitat. Phenology: Flowers. hederaefolia and Circaea alpina. Phenology: Vegetative. Origin: Origin: Native. Introduced. William E. Brockman s.n. 14 Aug 1956 Christie S. Isler 4-4-98 26 Apr 1998 WTU-213652 WTU-335300 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Rampart Ridge, Mt. Rainier. Head of Rainbow Creek, near McAlester Pass. 46.76917°, -121.81°; WGS 84, uncertainty: 10000 m., Source: Elev. 5000 ft. GeoLocate, Georef'd by WTU Staff 48.414174°, -120.685256°; WGS84, uncertainty: 5000 m., Source: Phenology: Vegetative. Origin: Native. GeoLocate, Georef'd by WTU Staff Wooded slopes, shade. Phenology: Flowers. Origin: Native. George Neville Jones 9656 27 Sep 1936 George H. Ward 115 22 Aug 1942 WTU-101588 WTU-182720 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., WASHINGTON, SKAMANIA COUNTY: Mount Saint Helens. U.S.A., WASHINGTON, CLALLAM COUNTY: Elev. 6150 ft. Hurricane Ridge. 46.19893°, -122.18888°; WGS 84, uncertainty: 5000 m., Source: Elev. 5000 ft. GeoLocate, Georef'd by WTU Staff 47.954534°, -123.440526°; WGS 84, uncertainty: 5000 m., Steep pumice slope. On decent trail, with grass. Phenology: Source: Digital Map, Georef'd by John Haskins Flowers. Origin: Native. Shady, moist canyon. Phenology: Flowers. Origin: Native. Donald B. Lawrence s.n. 5 Aug 1937 Edith Hardin English 1163 12 Jul 1941 WTU-324123 WTU-315755 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Cascade Mountains. Along Perry Creek Trail. Stampede Pass Tunnel, east entrance. Elev. 3000-3500 ft. Elev. 2800-3200 ft. 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., 47° 16' 30" N, 121° 19' 30" W; T21N R12E S27 Source: GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. Sarah Gage G&R 301 16 Jun 1992 J. William Thompson 17466 30 Jun 1953 with Sharon Rodman WTU-171361 WTU-344569 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., WASHINGTON, YAKIMA COUNTY: U.S.A., WASHINGTON, KING COUNTY: (YIN) At Howard Lake, 55 miles west of Toppenish ACF. By trail to Snow Lake. Elev. 4887-5248 ft. Elev. 4000 ft. 46.371695°, -121.461726°; WGS 84, uncertainty: 500 m., Source: 47.46722°, -121.45472°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Hudsonian zone. Phenology: Flowers. Origin: Native. Alpine woods. Phenology: Flowers. Origin: Native. E. Hunn 547 30 Jun 1977 J. William Thompson 10652 16 Jun 1934 WTU-271271 WTU-15882JWT Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, COWLITZ COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Mount Saint Helens, Toutle Mudflow northwest of crater breach. Tacoma. 46.34444°, -122.5275°; WGS 84, uncertainty: 10000 m., Source: 47.25306°, -122.44306°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Upland. Phenology: Fruits. Origin: Native. Waste shady places. Phenology: Flowers. Origin: Native. Jon Titus s.n. 14 Jul 1994 J. B. Flett s.n. 27 May 1896 WTU-334471 WTU-36746

Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Wenatchee Forest. Mt. Dickerman. Elev. 5250 ft. Elev. 3000 ft. T26N R15E S22 NW; WGS84, uncertainty: 32000 m., Source: 48.06829°, -121.47068°; WGS 84, uncertainty: 3000 m., Source: Georeferenced, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Rocky granite soil. Slope 10 S. Associated species: Kiderberry - Wooded slopes. Phenology: Flowers & Fruits. Origin: Native. Aconite - columbine. Phenology: Flowers. Origin: Native. J. William Thompson 14738 15 Jun 1940 Frank B. Lenzie 387 29 Jun 1918 WTU-21910JWT WTU-322294 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SKAMANIA COUNTY: U.S.A., WASHINGTON, ISLAND COUNTY: Chenowith. Hoypus Point, Deception Pass State Park, Whidbey Island. 45.73187°, -121.62838°; WGS 84, uncertainty: 1600 m., Source: 48.39389°, -122.6475°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Phenology: Vegetative. Origin: Native. Phenology: Fruits. Origin: Native. Wilhelm N. Suksdorf 11901 16 Aug 1925 Harold W. Smith 1440 31 Jul 1936 WTU-197553 WTU-40541 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, YAKIMA COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Klickitat R Rd. 1.5 mi S of Camp Chaparall at 30 mi W of White Slopes above Hyas Lake, 37 miles northwest of Cle Elum. Swan. Elev. 3600 ft. Elev. 2804 ft. 47.56722°, -121.12114°; WGS 84, uncertainty: 1000 m., Source: 46.381332°, -121.359625°; WGS 84, uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Origin: Native. Canadian. Phenology: Fruits. Origin: Native. C. L. Hitchcock 8048 14 Jul 1942 E. Hunn 704 4 Aug 1977 WTU-75667 WTU-270712 Rosaceae Rosaceae Rubus pensilvanicus Poir. Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Hollywood. 0.5 mile N of Seattle. Elev. 300 ft. 47.613627°, -122.33083°; WGS 84, uncertainty: 1000 m., Source: 47.73111°, -122.12806°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff In open woods. Phenology: Flowers & Fruits. Origin: Native. Wet bank. Growing erect. Phenology: Flowers. Origin: Introduced. Walter J. Eyerdam s.n. 15 May 1949 Ruby L. C. Sovereign 452 17 May 1941 WTU-290957 WTU-316850 Rosaceae Rosaceae Rubus macrophyllus Weihe & Nees Rubus nivalis Douglas ex Hook. U.S.A., WASHINGTON, MASON COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Hoodsport. Lake Crescent. 47.40611°, -123.13806°; WGS 84, uncertainty: 10000 m., Source: 48.09472°, -123.80583°; WGS 84, GeoLocate, Georef'd by WTU Staff uncertainty: 10000 m., Source: GeoLocate, Phenology: Fruits. Origin: Introduced. Georef'd by WTU Staff Phenology: Vegetative. Origin: Native. George Neville Jones 8542 5 Sep 1935 George Neville Jones 3436 WTU-31817 Aug 1931 Rosaceae WTU-21682, WTU-V-18512 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SAN JUAN COUNTY: Rosaceae Yellow Island. 48.592222°, -123.031944°; WGS 84, uncertainty: 250 m., Source: Rubus nivalis Douglas ex Hook. Georeferenced, Georef'd by WTU Staff East meadow. Growing on dry edge of south side of meadow. U.S.A., WASHINGTON, GRAYS HARBOR Phenology: Flowers & Fruits. Origin: Introduced. COUNTY: Colonel Bob Mountain. Eliza Habegger s.n. 22 Jul 1996 47.48889°, -123.74556°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, WTU-333041 Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Rosaceae Harold E. Helmrich 225 Rubus ursinus Cham. & Schltdl. 12 Jul 1933 U.S.A., WASHINGTON, THURSTON COUNTY: WTU-306034, WTU-V-18511 Unincorporated area, St. Clair, railroad tracks near McAllister Springs. Elev. 120 ft. Rosaceae 47° 1' 30" N, 122° 42' 45" W; T18N R1E S20 In fruit. Phenology: Fruits. Origin: Introduced. Rubus nivalis Douglas ex Hook. Sarah Gage G&R 330 20 Jun 1992 U.S.A., WASHINGTON, LEWIS COUNTY: with Sharon Rodman Lower slopes of Storm King Mountain, south of Elbe. WTU-345120 Elev. 3000-4000 ft. 46.650833°, -122.186389°; NAD 27, Rosaceae uncertainty: 5000 m., Source: Georeferenced, Georef'd by WTU Staff Rubus macrophyllus Weihe & Nees In woods. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, WHITMAN COUNTY: J. William Thompson 8611 20 miles west of Lewiston, along north bank of Snake River in small side canyon, near stream. 28 Jun 1932 T13N R44E S28; WGS 84, uncertainty: 1200 m., Source: Nat. Geog. TOPO!, Georef'd by John Bassett WTU-18614JWT, WTU-V-18510 Stems strongly glaucous, the primocanes greenish; the flowering canes long and trailing (to 12 ft.) and bluish-brown or bluish-purple; spines flattened, some straight, some hooked; leaves greenish on both surfaces, glabrous on the lower surface and only slightly paler Rosaceae than the upper surface; calyx almsot glabrous, but a few stipitate glands; immature fruits greenish, not at all glaucous, glabrous. Rubus nivalis Douglas ex Hook. Phenology: Fruits. Origin: Introduced. U.S.A., WASHINGTON, PIERCE COUNTY: C. L. Hitchcock 21860 11 Jun 1959 Tahoma Creek, Mount Rainier. with C. V. Muhlick 46.737489°, -121.903173°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, WTU-197465 Georef'd by WTU Staff Deep woods. Phenology: Flowers. Origin: Native. F. A. Warren 1592 24 Jul 1932 WTU-26124, WTU-V-18509 Rosaceae Rosaceae Rubus nivalis Douglas ex Hook. Rubus parviflorus Nutt. U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Near Summer Lake, Lake Cavanaugh Road. At 8-Mile Camp, 3 miles up 8-Mile Creek from Icicle Creek Bridge. Elev. 650 ft. 47.557948°, -120.672115°; WGS84, uncertainty: 10000 m., T33N R5E S21; NAD 27, uncertainty: 805 m., Source: GeoLocate, Georef'd by WTU Staff Source: TRS2LL, Georef'd by Ben Legler Open yellow pine forest. N = 7. Phenology: Flowers. Origin: Native. Mossy forest floor in deep conifer woods. With Linnaea borealis, Goodyera oblongifolia, and A. R. Kruckeberg 5412 7 Jul 1962 Vaccinium. Leaves deep glossy green with curved prickles on veins underneath. Trailing WTU-216079 vine, flowers purple to some white. Phenology: Flowers. Origin: Native. Rosaceae Mildred Arnot 194 with Elroy Burnett Rubus parviflorus Nutt. 26 May 1990 U.S.A., WASHINGTON, FERRY COUNTY: Along Columbia River 8 miles north of Inchelium. WTU-318210, WTU-V-18515 Elev. 1290 ft. 48.41571°, -118.2161°; WGS 84, uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff On moist ground among shrubs. Phenology: Flowers. Origin: Native. Rosaceae H. T. Rogers 537 25 May 1940 Rubus nivalis Douglas ex Hook. WTU-82980 U.S.A., WASHINGTON, WHATCOM COUNTY: Near Coal Creek, Glacier. 48.90464°, -121.906838°; WGS 84, Rosaceae uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Rubus parviflorus Nutt. Coniferous forest. Phenology: Vegetative. Origin: Native. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Along shore of Quinault Lake, Olympic Forest. W. C. Muenscher 16183 Elev. 200 ft. 47.46722°, -123.84417°; WGS 84, uncertainty: 10000 m., Source: 6 Sep 1943 GeoLocate, Georef'd by WTU Staff Slope 40% west; soil gravelly clay loam. Timbered, with vine maple, WTU-118971, WTU-V-18507 salmonberry. Common in mosit soil. Phenology: Flowers. Origin: Native. J. E. Schwartz 57 5 Jun 1936 Rosaceae WTU-91429 Rubus parviflorus Nutt. U.S.A., WASHINGTON, COLUMBIA COUNTY: Rosaceae Blue Mountains: North facing slope near Tucannon Campground, within 200 meters of campground. Rubus parviflorus Nutt. Elev. 2621 ft. 46° 14' 34" N, 117° 41' 22" W; T9N R41E S21; WGS 84, U.S.A., WASHINGTON, ISLAND COUNTY: uncertainty: 551 m., Source: Digital Map, Georef'd by Richard Near Good Beach. Olmstead 48.21359°, -122.72215°; WGS 84, uncertainty: 500 m., Source: Growing on rocky outcrops; scattered, infrequent. Petals white. GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Wooded roadside. Phenology: Flowers. Origin: Native. Richard G. Olmstead 97-31 27 Jun 1997 Lloyd E. McElvain 62 25 May 1950 WTU-339060 WTU-129212 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, CLALLAM COUNTY: Mt. Angeles. U.S.A., WASHINGTON, JEFFERSON COUNTY: Elev. 5000 ft. Lower Lena Lake. 47.995278°, -123.474444°; NAD 27, uncertainty: 1000 m., Source: Elev. 2000 ft. Georeferenced, Georef'd by WTU Staff 47.61944°, -123.15833°; WGS 84, uncertainty: 10000 m., Source: Open grassy meadows and talus slopes. 2 collections on this sheet. GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. J. William Thompson 7548 19 Jul 1931 S. R. Wellings 43 18 Jul 1948 WTU-23010JWT-A WTU-183077 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KITSAP COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Port Orchard. Mount Rainier. 47.645917°, -122.609352°; WGS 84, uncertainty: 10000 m., 46.852827°, -121.760441°; WGS 84, uncertainty: 17500 m., Source: GeoLocate, Georef'd by WTU Staff Source: GeoLocate, Georef'd by Joseph Stormer Open woods. Phenology: Flowers. Origin: Native. Phenology: Fruits. Origin: Native. Walter J. Eyerdam 1574 29 May 1938 J. M. Grant s.n. Aug 1925 WTU-95843 WTU-22089

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KLICKITAT COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Keilfall, Bingen. San Juan Island, University of Washington Friday Harbor 45.715°, -121.46333°; WGS 84, uncertainty: 10000 m., Source: Laboratories. GeoLocate, Georef'd by WTU Staff 48.546944°, -123.010833°; WGS 84, uncertainty: 300 m., Source: Accession number on sheet is hard to read. May be entered wrong Georeferenced, Georef'd by WTU Staff here. Phenology: Flowers. Origin: Native. Woods of Pseudotsuga, Arbutus, and ALnus. Shrub 4-5 feet tall; flwoers white; locally common. Phenology: Flowers. Origin: Native. Wilhelm N. Suksdorf s.n. 18 May 1902 Susan Gates 40 12 May 1975 WTU-137558 WTU-263237 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KITTITAS COUNTY: Slopes above Hyas Lake, 37 miles northwest of Cle Elum. U.S.A., WASHINGTON, STEVENS COUNTY: Elev. 3600 ft. Black Lake. 47.56722°, -121.12114°; WGS 84, uncertainty: 1000 m., Source: T35N R41E S3; NAD 27, uncertainty: 805 m., Source: TRS2LL, GeoLocate, Georef'd by WTU Staff Georef'd by Ben Legler Flowers pink on about half of plants of the area. Phenology: Flowers. In open woods on west side of lake. Phenology: Flowers. Origin: Origin: Native. Native. C. L. Hitchcock 8042 14 Jul 1942 Monita Engvall 120 14 Jun 1958 WTU-75657 WTU-195327

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, SPOKANE COUNTY: Cougar Mountain. Spokane. 47.54056°, -122.09806°; WGS 84, uncertainty: 10000 m., Source: 47.65889°, -117.425°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Phenology: Vegetative. Origin: Native. In open woods. Phenology: Flowers. Origin: Native. David M. Chapin s.n. 9 Oct 1982 Sister Mary Milburge s.n. May 1931 WTU-314240 WTU-30775

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, MASON COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Route 101, Tillamook Beach, Hood Canal. Stillaguamish River 1.5 miles west of Arlington, near State Road 47.40418°, -123.14052°; WGS 84, uncertainty: 5000 m., Source: 530. GeoLocate, Georef'd by WTU Staff 48.201684°, -122.160147°; WGS 84, uncertainty: 1000 m., In moist soil. Phenology: Flowers. Origin: Native. Source: GeoLocate, Georef'd by WTU Staff Abundant on river banks. Shrub 1 meter tall. Phenology: Flowers. Phillips B. Freer 195 28 May 1950 Origin: Native. WTU-134990 Mildred Arnot 103 9 May 1988 WTU-314671 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus pedatus Sm. U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Near junction of Sauk and Suiattle Rivers. Copalis Bog Preserve. Elev. 450 ft. T21N R10W S20 SE 1/4; T21N R10W S20 NE 1/4; NAD 27, 48.475°, -121.62528°; WGS 84, uncertainty: 10000 m., Source: uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler GeoLocate, Georef'd by WTU Staff Clearcut, with Gaultheria shallon, Maianthemm dilatatum. Growing Slope level; soil deep rich sandy bottom loam. Conifer forest, with on a stump. Phenology: Flowers. Origin: Native. bluegrass, bracken fern, fireweed, vine maple, weeds. Frequent. Phenology: Flowers. Origin: Native. Eliza Habegger 618 11 Apr 1998 D. C. Ingram 1922 15 May 1925 WTU-338847 WTU-322442 Rosaceae Rosaceae Rubus pedatus Sm. Rubus parviflorus Nutt. U.S.A., WASHINGTON, JEFFERSON COUNTY: Upper Lena Trail. U.S.A., WASHINGTON, WHATCOM COUNTY: Elev. 2500 ft. Above Bagley Lake, Mt. Baker Lodge. 47.63055°, -123.1837°; WGS 84, uncertainty: 2000 m., Source: Elev. 4300-4500 ft. GeoLocate, Georef'd by WTU Staff 48.85675°, -121.68675°; WGS 84, uncertainty: 575 m., Source: Phenology: Flowers. Origin: Native. Nat. Geog. TOPO!, Georef'd by John Bassett On rocky slopes. Phenology: Flowers. Origin: Native. S. R. Wellings 50 18 Jul 1948 J. William Thompson 5358 25 Jul 1930 WTU-183082 WTU-23006JWT Rosaceae Rosaceae Rubus pedatus Sm. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KING COUNTY: By trail to Snow Lake. U.S.A., WASHINGTON, YAKIMA COUNTY: Elev. 4000 ft. (YIN) Old Maid Canyon, 15 miles west of White Swan. 47.46722°, -121.45472°; WGS 84, uncertainty: 10000 m., Source: Elev. 3493 ft. GeoLocate, Georef'd by WTU Staff 46.382628°, -121.04481°; WGS 84, uncertainty: 500 m., Source: Alpine woods. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Canadian road edge. Phenology: Flowers. Origin: Native. J. William Thompson 10654 16 Jun 1934 E. Hunn 519 30 Jul 1977 WTU-15861JWT WTU-271292 Rosaceae Rosaceae Rubus pedatus Sm. Rubus pedatus Sm. U.S.A., WASHINGTON, MASON COUNTY: Mount Elinor [Ellinor] Trail. U.S.A., WASHINGTON, CLALLAM COUNTY: 47.52205°, -123.26061°; WGS 84, uncertainty: 500 m., Source: Deer Lake Basin South Meadow, Olympic National Park. GeoLocate, Georef'd by WTU Staff Elev. 3600 ft. In dry soil, cutover land. Phenology: Flowers. Origin: Native. 47° 55' N, 123° 50' W Phenology: Flowers. Origin: Native. Phillips B. Freer 126 4 Jun 1950 J. M. Patt s.n. 23 Sep 1983 WTU-135113 WTU-313038 Rosaceae Rosaceae Rubus pedatus Sm. Rubus pedatus Sm. U.S.A., WASHINGTON, OKANOGAN COUNTY: North summit road, Twisp Cut-off route. U.S.A., WASHINGTON, CHELAN COUNTY: 48.36361°, -120.12111°; WGS 84, uncertainty: 10000 m., Source: Trail to Stuart Lake. GeoLocate, Georef'd by WTU Staff Elev. 5000 ft. In a swamp. 2 collections on this sheet. Phenology: Flowers. Origin: 47.51797°, -120.8242°; WGS 84, uncertainty: 20000 m., Source: Native. GeoLocate, Georef'd by WTU Staff Moist wooded slopes. 2 collections on this sheet. Phenology: Charles B. Fiker 913 26 Jun 1932 Flowers. Origin: Native. WTU-23013 Clarence B. Seely 31 8 Jun 1931 WTU-23008 Rosaceae Rosaceae Rubus pedatus Sm. Rubus spectabilis Pursh U.S.A., WASHINGTON, SKAMANIA COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Along N733, 6 miles east of Eagle's Cliff at Swift Reservoir. Lake Ozette. Elev. 2591-2985 ft. Elev. 50 ft. 46.06333°, -122.19667°; WGS 84, uncertainty: 5000 m., Source: 48.155°, -124.66583°; WGS 84, uncertainty: GeoLocate, Georef'd by WTU Staff 10000 m., Source: GeoLocate, Georef'd by Canadian road edge, west side of crest. Phenology: Flowers. Origin: WTU Staff Native. On hillside. Phenology: Flowers. Origin: Native. E. Hunn 695 3 Sep 1977 Ruby L. C. Sovereign 411 WTU-270750 26 Apr 1941 WTU-316078, WTU-V-18373 Rosaceae Rubus pedatus Sm. Rosaceae U.S.A., WASHINGTON, SNOHOMISH COUNTY: Cascade Mountains. Along Perry Creek Trail. Rubus spectabilis Pursh Elev. 3000-3500 ft. 48.063294°, -121.513675°; WGS 84, uncertainty: 10000 m., U.S.A., WASHINGTON, COWLITZ COUNTY: Source: GeoLocate, Georef'd by WTU Staff On South Toutle Road 1 mile on left side of Phenology: Flowers. Origin: Native. spirit Lake Highway. 46.325°, -122.73528°; WGS 84, uncertainty: J. William Thompson 17465 30 Jun 1953 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-171360 Loam soil. With horsetails and ferns. Phenology: Flowers & Fruits. Origin: Native. Rosaceae Marianne Dapp 14 Rubus pedatus Sm. 29 Apr 1980 U.S.A., WASHINGTON, WHATCOM COUNTY: WTU-315454, WTU-V-18385 Trail to Mt. Ann. Elev. 4400 ft. 48.82672°, -121.64608°; WGS 84, uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff Rosaceae At timberline. Trailing in underbrush. Phenology: Flowers. Origin: Native. Rubus spectabilis Pursh J. William Thompson 5345 25 Jul 1930 U.S.A., WASHINGTON, CHELAN COUNTY: Along State Creek near junction with west fork WTU-23007JWT of Bridge Creek. 47.562003°, -120.779518°; WGS84, uncertainty: 10000 m., Source: GeoLocate, Rosaceae Georef'd by WTU Staff Frest. Shrub to 3 ft. tall. Phenology: Fruits. Rubus pubescens Raf. Origin: Native. U.S.A., WASHINGTON, FERRY COUNTY: George H. Ward 611 East shore of Swan Lake. 48.51333°, -118.83583°; WGS 84, uncertainty: 10000 m., Source: 27 Jul 1946 GeoLocate, Georef'd by WTU Staff Deep shaded fertile soil. Phenology: Flowers. Origin: Native. WTU-182725, WTU-V-18383 D. Sanford 102 31 May 1936 with J. Jorgensen Rosaceae WTU-9007 Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Rubus pubescens Raf. Along the banks of Humptulips River. 47.22956°, -123.956466°; WGS 84, U.S.A., WASHINGTON, STEVENS COUNTY: uncertainty: 10000 m., Source: GeoLocate, Lake Thomas. Georef'd by WTU Staff 48.62774°, -117.53941°; WGS 84, uncertainty: 1500 m., Source: Phenology: Flowers & Fruits. Origin: Native. GeoLocate, Georef'd by WTU Staff In moist forest along lake shore. Phenology: Flowers. Origin: Native. George Neville Jones 4589 Earle F. Layser 1760 Aug 1971 8 Apr 1934 WTU-266101 WTU-23574, WTU-V-18382 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, ISLAND COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Near Rhododendron Park. Watmough Bay, southeast end of Lopez Island. 48.20259°, -122.64998°; WGS 84, T34N R1W S21; WGS 84, uncertainty: 10000 uncertainty: 700 m., Source: GeoLocate, m., Source: GeoLocate, Georef'd by WTU Georef'd by WTU Staff Staff Woods. Phenology: Flowers. Origin: Native. Much branched shrub ca. 3 meters tall, on edge of Pseudotsuga menziesii, Alnus rubra, Thuja Lloyd E. McElvain 42 plicata woods. Flowers bright pink. Phenology: Flowers. Origin: Native. 12 Aug 1950 Dorothy Naas 5755 WTU-129122, WTU-V-18381 with Mildred Arnot 19 Apr 1996 Rosaceae WTU-333481, WTU-V-18377 Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, JEFFERSON COUNTY: Rubus spectabilis Pursh Upper Lena Trail. Elev. 2500 ft. U.S.A., WASHINGTON, SNOHOMISH 47.63055°, -123.1837°; WGS 84, uncertainty: COUNTY: 2000 m., Source: GeoLocate, Georef'd by 2 miles east of Gold Bar along Forest Service WTU Staff Road 6010; along Skykomish River. Phenology: Flowers. Origin: Native. Elev. 1000 ft. T27N R9E S9; NAD 27, uncertainty: 805 m., S. R. Wellings 57 Source: TRS2LL, Georef'd by Ben Legler Alnus rubra stand with Scot's broom, along 18 Jul 1948 primitive road; soil of gravelly glacial till; near river. Phenology: Flowers & Fruits. Origin: WTU-194252, WTU-V-18380 Native. L. Hegstrom 18 Rosaceae 10 May 1986 Rubus spectabilis Pursh WTU-313225, WTU-V-18376 U.S.A., WASHINGTON, KING COUNTY: Kings Lake, 1 mile west of Boyle Lake, Rosaceae northeast of Snoqualmie Falls. Elev. 951 ft. Rubus spectabilis Pursh T24N R8E; WGS 84, uncertainty: 997 m., Source: Digital Map, Georef'd by Joseph U.S.A., WASHINGTON, SKAMANIA COUNTY: Stormer Cold Springs Indian Camp, 18 miles northwest of Trout Lake. Shrub, 1-2 meters high; corolla rose-pink. Elev. 3493 ft. Growing along road near bog with Sambucus 46.12694°, -121.73056°; WGS 84, uncertainty: 10000 m., Source: racemosa. Phenology: Flowers & Fruits. Origin: GeoLocate, Georef'd by WTU Staff Native. Canadian zone. Phenology: Fruits. Origin: Native. G. Lebednik 359 E. Hunn 673 2 Sep 1977 10 May 1973 WTU-271212 WTU-261052, WTU-V-18379 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, WHATCOM COUNTY: Goshen. U.S.A., WASHINGTON, LEWIS COUNTY: 48.85472°, -122.33972°; WGS 84, Narada Falls of Paradise River, Mt. Rainier uncertainty: 10000 m., Source: GeoLocate, National Park. Georef'd by WTU Staff Elev. 3800 ft. Growing along a stream. Phenology: Flowers. 46.775683°, -121.745691°; WGS 84, Origin: Native. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff W. C. Muenscher 9767 In heavy thickets above creek bed. Phenology: Flowers. Origin: Native. 1 May 1939 Joseph Andorfer Ewan 10638 WTU-55606, WTU-V-18375 20 Aug 1937 WTU-52535, WTU-V-18378 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, KITSAP COUNTY: West side of Wenatchee River at upper end of Tumwater Canyon. Port Orchard. T25N R17E; WGS84, uncertainty: 10000 m., Source: GeoLocate, 47.645917°, -122.609352°; WGS 84, uncertainty: 10000 m., Georef'd by WTU Staff Source: GeoLocate, Georef'd by WTU Staff Steep hillside. Phenology: Flowers. Origin: Native. Growing on logs in burnt over area. Phenology: Flowers. Origin: Native. George H. Ward 491 6 Jun 1946 Walter J. Eyerdam 1594 29 May 1938 WTU-188714 WTU-292480 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, CLALLAM COUNTY: Crystal Creek. U.S.A., WASHINGTON, KLICKITAT COUNTY: Elev. 2500 ft. Mountains near Bingen. 47.977296°, -123.684108°; WGS 84, uncertainty: 10000 m., 45.715°, -121.46333°; WGS 84, uncertainty: 5000 m., Source: Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Dense forest. Phenology: Vegetative. Origin: Native. Phenology: Flowers. Origin: Native. George Neville Jones 8446 24 Aug 1935 Wilhelm N. Suksdorf 1757 8 Jun 1901 WTU-31783 WTU-171451

Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: McCleary. Scatter Creek Trail, 2 miles from junction with Fish Creek Road. 47.05333°, -123.26417°; WGS 84, uncertainty: 10000 m., Source: Elev. 4200 ft. GeoLocate, Georef'd by WTU Staff 47.507151°, -121.059494°; WGS 84, uncertainty: 10000 m., Phenology: Flowers. Origin: Native. Source: GeoLocate, Georef'd by WTU Staff On open rocky knoll near stream. Phenology: Flowers. Origin: George Neville Jones 4592 7 Apr 1934 Native. WTU-23600 C. L. Hitchcock 8011 6 Jul 1942 WTU-75329 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, ISLAND COUNTY: Near Fort Casey. U.S.A., WASHINGTON, LEWIS COUNTY: 48.16444°, -122.6775°; WGS 84, uncertainty: 5000 m., Source: 1 mile south of first bridge over Cowlitz River, on road out of GeoLocate, Georef'd by WTU Staff Packwood. Wooded roadside. Phenology: Flowers. Origin: Native. 46.460887°, -122.778497°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Lloyd E. McElvain 67 23 May 1950 Phenology: Flowers. Origin: Native. WTU-129207 Ruby L. C. Sovereign 68 12 May 1940 WTU-315754 Rosaceae Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, KING COUNTY: Rubus ursinus Cham. & Schltdl. Bellevue. 47.61056°, -122.19944°; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, PIERCE COUNTY: GeoLocate, Georef'd by WTU Staff American Lake. Phenology: Flowers. Origin: Native. 47.1325°, -122.56361°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff I. L. Norberg s.n. 1 May 1934 Phenology: Flowers. Origin: Native. WTU-26656 George Neville Jones 4602 29 Apr 1934 WTU-26456 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus pedatus Sm. U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, FERRY COUNTY: Mt. Dickerman. Okanogan Valley: 0.5 miles along Forest Service Road 9565 (South Elev. 3000 ft. Deadman). 4.1 miles from Highway 20, off south side of road about 48.06829°, -121.47068°; WGS 84, uncertainty: 3000 m., Source: 100 yards near old fire road. GeoLocate, Georef'd by WTU Staff Elev. 4900 ft. Wooded slopes. Phenology: Flowers. Origin: Native. T37N R35E S31; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler J. William Thompson 14739 15 Jun 1940 Englemann spruce bog; open/partial cover; slopes with bunchberry, Vaccinium, Alnus, lodgepole pine, Arctostaphylos. Phenology: WTU-21914JWT Flowers. Origin: Native. Suzanne Bagshaw 204 11 Jul 1999 Rosaceae WTU-349598 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SAN JUAN COUNTY: Rosaceae Lopez Island; Iceberg Point, 7 miles SSE of Lopez Village. T34N R2W S24; WGS 84, uncertainty: 500 m., Source: Rubus parviflorus Nutt. Georeferenced, Georef'd by WTU Staff Edge of PSME woods. With Rosa nutkana, Sambucus racemosa, U.S.A., WASHINGTON, FERRY COUNTY: grasses. Trailing , petals white. Phenology: Flowers. Origin: Colville National Forest: Vicinity of Swan Lake. Native. Elev. 3600 ft. 48° 31' 4" N, 118° 50' 16" W; T35N R32E S20; WGS 84, Dorothy Naas 5717 17 May 1995 uncertainty: 532 m., Source: Digital Map, Georef'd by Richard Olmstead WTU-331954 Open mixed conifer forest - Douglas fir, Ponderosa pine, Larch. Phenology: Flowers. Origin: Native. Rosaceae Richard G. Olmstead 99-106 8 Jul 1999 Rubus ursinus Cham. & Schltdl. WTU-349299 U.S.A., WASHINGTON, SKAMANIA COUNTY: Rosaceae Below Flattop Mountain, on N78, 25 miles west of Glenwood. Elev. 3050-3149 ft. Rubus spectabilis Pursh 46.02833°, -121.61278°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., WASHINGTON, KING COUNTY: Canadian zone. Phenology: Fruits. Origin: Native. Cascade Mountains of Western Washington: Cedar River Watershed. Rock Creek wetland system. Head into south end from E. Hunn 650 2 Sep 1977 rd 40. Drop into system at Binker culvert and head into west side of the wetland. WTU-271159 Elev. 800 ft. T22N R7E S16 NE; NAD 27, uncertainty: 500 m., Source: Georeferenced, Georef'd by WTU Staff Rosaceae Rock Creek wetland system. Head into south end from rd 40. Drop into system at Binker culvert and head into west side of the wetland. Rubus bifrons Vest Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Clayton J. Antieau 01-5 26 May 2001 Along the banks of Humptulips River. with Linda Cooley, Kristie Dinkin 47.22956°, -123.956466°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-350619 Climbing among shrubs on banks of river. Phenology: Flowers. Origin: Introduced. Rosaceae George Neville Jones 4593 8 Apr 1934 Rubus ursinus Cham. & Schltdl. WTU-15905 U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington. Cedar River Watershed. Little Mountain Bog. Rosaceae Elev. 1600 ft. T22N R9E S27; NAD 27, uncertainty: 500 m., Source: Rubus parviflorus Nutt. Georeferenced, Georef'd by WTU Staff Marshy and boggy area (actually a fen) surrounded by forested bog. U.S.A., WASHINGTON, FERRY COUNTY: The fen is a mosaic of sedge marsh, sphagnum moss, and open Colville National Forest: Forest Service Road 53 to 5330 to water. Growing in open fen area dominated by Ledum, Kalmia, and 5330-100. Approximately 1 mile from Ferry Lake Fen. Myrica. Phenology: Flowers. Origin: Native. T35N R32E S16; WGS 84, uncertainty: 805 m., Source: Digital Map, Georef'd by Joseph Stormer Clayton J. Antieau 01-26 23 Jun 2001 Phenology: Flowers. Origin: Native. with Nancy Job, Sandra Whiting, Jeffery Walker Suzanne Bagshaw 35 9 Jul 1999 WTU-350672 WTU-349608 Rosaceae Rosaceae Rubus nivalis Douglas ex Hook. Rubus spectabilis Pursh U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, KING COUNTY: COUNTY: Cascade Mountains; Sutton Lake. Lost Ridge Trail #646, ca. 1 mile above Elev. 3650 ft. trailhead. North Fork Sauk River drainage. 47° 17.769' N, 121° 32.05' W; T21N R10E S21 NE 1/4 Elev. 2500 ft. Herbaceous meadow surrounding the lake, the adjacent forest, and 48° 6' 1" N, 121° 19' 35" W an older clear cut. Dominant trees: , Abies Dense forest of Tsuga heterophylla, amabilis. Low growing (<1 M) at rocky edge of lake. Petl;as deep Pseudotsuga menziesii. South facing slope. pink, starting to fruit. Phenology: Flowers & Fruits. Origin: Native. Common along trail, in desne patches. No flowers or fruits seen. Leaf material removed for Sarah Gage 7253 14 Jul 2001 RADseq analysis by Megan Ruffley at WTU, with Barb DeCaro, Ian Stewart, Jenny Stewart February 2016. Phenology: Vegetative. Origin: Native. WTU-350840 Ben Legler 65 Rosaceae 21 Sep 2002 Rubus lasiococcus A. Gray WTU-350372, WTU-V-019106 U.S.A., WASHINGTON, KING COUNTY: Rosaceae Cascade Mountains; Sutton Lake. Elev. 3650 ft. Rubus parviflorus Nutt. 47° 17.769' N, 121° 32.05' W; T21N R10E S21 NE 1/4 Herbaceous meadow surrounding the lake, the adjacent forest, and U.S.A., WASHINGTON, KING COUNTY: an older clear cut. Dominant trees: Tsuga mertensiana, Abies Cascade Mountains of Western Washington: Rattlesnake Ledge amabilis. White petals. Phenology: Flowers. Origin: Native. trail. Elev. 1491 ft. Sarah Gage 7266 14 Jul 2001 47° 25.979' N, 121° 46.79' W; T23N R8E S33 with Barb DeCaro, Ian Stewart, Jenny Stewart Open area along trail. Common. Phenology: Flowers. Origin: Native. WTU-350841 Sarah Gage 7224 23 Jun 2001 with Cindy Spurgeon, Janet Sailer Rosaceae WTU-350860 Rubus pedatus Sm. Rosaceae U.S.A., WASHINGTON, KING COUNTY: Rubus ursinus Cham. & Schltdl. Cascade Mts., Pacific Crest Trail above Twilight Lake. Elev. 3600-4000 ft. U.S.A., WASHINGTON, KING COUNTY: T21N R10E S5; NAD 27, uncertainty: 1000 m., Source: Cascade Mountains of Western Washington: Rattlesnake Ledge Georeferenced, Georef'd by WTU Staff trail. Old growth forest. Abies amabilis, Abies procera, Tsuga heterophylla Elev. 1491 ft. with Xerophyllum tenax, Rhodendron albiflorum, and Vaccinium 47° 25.979' N, 121° 46.79' W; T23N R8E S33 species in the understory. In fruit. Phenology: Fruits. Origin: Native. Open area along trail. Starting to friut. Phenology: Flowers & Fruits. Origin: Native. Clayton J. Antieau 01-12 11 Aug 2001 with Katie Saurer, Mike Mazer Sarah Gage 7219 23 Jun 2001 with Cindy Spurgeon, Janet Sailer WTU-350883 WTU-350859 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus parviflorus Nutt. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Cedar River bridge at Cascade Mountains; Sutton Lake. Barneston. Elev. 3650 ft. Elev. 1000 ft. 47° 17.769' N, 121° 32.05' W; T21N R10E S21 NE 1/4 47° 23.33' 33" N, 121° 51.68' 51" W; T22N R7E S14 Herbaceous meadow surrounding the lake, the adjacent forest, and Woods, riparian area, and roadside near bridge. Pinkish-red fruit. an older clear cut. Dominant trees: Tsuga mertensiana, Abies Phenology: Flowers & Fruits. Origin: Native. amabilis. White petals. Phenology: Flowers. Origin: Native. P. F. Zika 16034 26 May 2001 Sarah Gage 7269 14 Jul 2001 with Jeff Walker, David Kiefer, Kari Stices, Bill Winije with Barb DeCaro, Ian Stewart, Jenny Stewart WTU-350660 WTU-350842 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus lasiococcus A. Gray U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Cedar River bridge at Cascade Mountains of Western Washington: Findley Lake. Barneston. Elev. 3400-3800 ft. Elev. 1000 ft. T21N R10E S7; NAD 27, uncertainty: 300 m., Source: 47° 23.33' 33" N, 121° 51.68' 51" W; T22N R7E S14 Georeferenced, Georef'd by WTU Staff Woods, riparian area, and roadside near bridge. Trailing, about 8 East-facing talus slope. Phenology: Flowers & Fruits. Origin: Native. meters tall. White flowers. Phenology: Flowers. Origin: Native. Nancy Job 01-38 11 Aug 2001 P. F. Zika 16007 26 May 2001 with Bill Belknap, Julie Belknap, Susan Rafanelli, Kathryn Sobocinski with Jeff Walker, David Kiefer, Kari Stices, Bill Winije WTU-351093 WTU-350703 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Findley Lake. Williams Creek. Wetland southeast of Road 30 and southwest of the Elev. 3400-3800 ft. creek. Cascade Mountains of Western Washington. T21N R10E S7; NAD 27, uncertainty: 300 m., Source: Elev. 800 ft. Georeferenced, Georef'd by WTU Staff T22N R7E S13 NE 1/4; NAD 27, uncertainty: 500 m., Source: East-facing talus slope. Small, white flower. Phenology: Flowers. Georeferenced, Georef'd by WTU Staff Origin: Native. Plants in area: Alnus rubra, Psuedotsuga menziesii, sword fern, braken fern, Athyrium sp., Carex aquatilis, Acer macrophyllum, Nancy Job 01-33 11 Aug 2001 Digitalis purpurea, Sambucus racemosa, Plantago major. with Bill Belknap, Julie Belknap, Susan Rafanelli, Kathryn Sobocinski Phenology: Flowers. Origin: Native. WTU-351113 Alan C. Yen 01-4 26 May 2001 WTU-350942 Rosaceae Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, KING COUNTY: Rubus spectabilis Pursh Halmar Gate Wetland; Cascade Mountains of Western Washington. U.S.A., WASHINGTON, KING COUNTY: Elev. 1400 ft. Cascade Mountains of Western Washington: Wetlands along road T22N R7E S1; NAD 27, uncertainty: 500 m., 30; near headwaters of Williams Creek. Source: Georeferenced, Georef'd by WTU Elev. 1000 ft. Staff T22N R7E S2 NE 1/4; NAD 27, uncertainty: 805 m., Source: Ditches filled with water, drier hummocks with TRS2LL, Georef'd by Ben Legler Spiraea sp., Juncus effusus, Juncus supinus. Other plants in vicinity: Thuja plicata, Tsuga heterophylla, Other plants in vicinity: Salix sp., Lysichitum Psuedotsuga menziesii, Acer circinatum, Galium sp., Glycyrrhiza sp. americanum. Gaultheria shallon growing from Phenology: Flowers. Origin: Native. stumps; Menziesia ferruginea common. Upland island with doug fir, western hemlock, red alder. Alan C. Yen 01-53 26 May 2001 Golden yellow fruit. Phenology: Fruits. Origin: Native. WTU-350943 Suzanne Bagshaw 01-19 23 Jun 2001 with Roz Glasser, Mary Clare Schroeder, Neil Dijrani Rosaceae WTU-351092, WTU-V-044847 Rubus laciniatus Willd. Rosaceae U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Powerline along Rubus pedatus Sm. Kerriston road. Elev. 1000 ft. U.S.A., WASHINGTON, KING COUNTY: T22N R7E S2 NE 1/4; NAD 27, uncertainty: 805 m., Source: Cascade Mountains; Sutton Lake. TRS2LL, Georef'd by Ben Legler Elev. 3650 ft. Other plants in area: Alnus rubra, bracken fern, Rubus parviflora, 47° 17.769' N, 121° 32.05' W; T21N R10E S21 NE 1/4 Plantago major, Juncus effusus, Gaultheria shallon, Maianthum Herbaceous meadow surrounding the lake, the adjacent forest, and dilatatum, Dicentra sp. Phenology: Vegetative. Origin: Introduced. an older clear cut. Dominant trees: Tsuga mertensiana, Abies amabilis. White petals. Phenology: Flowers. Origin: Native. Alan C. Yen 01-41 26 May 2001 Sarah Gage 7274 14 Jul 2001 WTU-350946 with Barb DeCaro, Ian Stewart, Jenny Stewart WTU-350791 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Powerline along Cascade Mountains of Western Washington: Findley Lake. Kerriston road. Elev. 3400-3800 ft. Elev. 1000 ft. T21N R10E S7; NAD 27, uncertainty: 300 m., Source: T22N R7E S2 NE 1/4; NAD 27, uncertainty: 805 m., Source: Georeferenced, Georef'd by WTU Staff TRS2LL, Georef'd by Ben Legler East-facing talus slope. Phenology: Fruits. Origin: Native. Other plants in area: Alnus rubra, bracken fern, Rubus parviflora, Plantago major, Juncus effusus, Gaultheria shallon, Maianthum Nancy Job 01-14 11 Aug 2001 dilatatum, Dicentra sp. Phenology: Flowers. Origin: Native. with Bill Belknap, Julie Belknap, Susan Rafanelli, Kathryn Sobocinski Alan C. Yen 01-32 26 May 2001 WTU-351221 WTU-351289 Rosaceae Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Christmas Lake. U.S.A., WASHINGTON, KING COUNTY: Elev. 1000 ft. Cascade Mountains of Western Washington: Powerline along 47° 26.008' N, 121° 45.693' W; T23N R8E S34 Kerriston road. Upland areas are dominated by alder, some doug fir, cedar and Elev. 1000 ft. hemlock. Lake has a large amount of floating logs, some standing T22N R7E S2 NE 1/4; NAD 27, uncertainty: 805 m., Source: snags. In fruit. Growing in upland. Phenology: Fruits. Origin: TRS2LL, Georef'd by Ben Legler Introduced. Other plants in area: Alnus rubra, bracken fern, Rubus parviflora, Plantago major, Juncus effusus, Gaultheria shallon, Maianthum Suzanne Bagshaw 01-206 11 Aug 2001 dilatatum, Dicentra sp. Phenology: Flowers & Fruits. Origin: Native. with Michelle Koehler, Julie Hall, Sage Chaiya Pechara Alan C. Yen 01-26 26 May 2001 WTU-351207 WTU-351225 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Mt. Baldy. U.S.A., WASHINGTON, KING COUNTY: Elev. 4911 ft. Williams Creek. Wetland southeast of Road 30 47° 20.93' N, 121° 30.086' W; T22N R10E S35 and southwest of the creek. Cascade Mountains Plants in area: Abies amabalis, Smilacina stellata, Rubus pedatus, of Western Washington. Rhododendron albiflorum, Vaccinium sp., Xerophyllum tenax. White Elev. 800 ft. flowers; trailing in shade under Abies amabilis. Phenology: Flowers. T22N R7E S13 NE 1/4; NAD 27, uncertainty: Origin: Native. 500 m., Source: Georeferenced, Georef'd by WTU Staff Richard G. Olmstead 01-85 11 Aug 2001 Plants in area: Alnus rubra, Psuedotsuga with Hillary Hayden, Adam Ryen, April Schiller, Mary Clare Schroder menziesii, sword fern, braken fern, Athyrium sp., Carex aquatilis, Acer macrophyllum, Digitalis WTU-351457 purpurea, Sambucus racemosa, Plantago major. Phenology: Flowers & Fruits. Origin: Native. Rosaceae Alan C. Yen 01-3 Rubus spectabilis Pursh 26 May 2001 U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: WTU-351224, WTU-V-18367 Law's Ledge. Elev. 3400 ft. T22N R8E S10 NE; NAD 27, uncertainty: 805 Rosaceae m., Source: TRS2LL, Georef'd by Ben Legler Old clear cut, rock, gravel. Some firs. Rubus parviflorus Nutt. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: David Kiefer 01-030 Cascade Mountains of Western Washington: Findley Lake. with Sharon Brown, Bob Antieau, Rena Elev. 3400-3800 ft. Hamburger T21N R10E S7; NAD 27, uncertainty: 300 m., Source: Georeferenced, Georef'd by WTU Staff 23 Jun 2001 East-facing talus slope. Phenology: Fruits. Origin: Native. WTU-351233, WTU-V-18368 Nancy Job 01-16 11 Aug 2001 with Bill Belknap, Julie Belknap, Susan Rafanelli, Kathryn Sobocinski WTU-351142 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus lasiococcus A. Gray U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Cascade Mountains of Western Washington: Mount Gardner. Law's Ledge. Elev. 3700 ft. Elev. 3400 ft. 47° 22' N, 121° 33' 5" W; T22N R10E S29 NE T22N R8E S10 NE; NAD 27, uncertainty: 805 Collected at the intersection of roads 155, 155.5, and 155.6. Plants m., Source: TRS2LL, Georef'd by Ben Legler in area: Tsuga mertensiana, Pseudotsuga menziesii, Abies Old clear cut, rock, gravel. Some firs. lasiocarpa, Abiea amabalis, Alnus rubra, Polystichum munitum, Phenology: Flowers. Origin: Native. Rubus spectabilis, Gaultheria shallon. Growing in well drained rocky soil in old clear cut with bear grass, Epilobium sp., Carex sp., Alnus David Kiefer 01-016 rubra, Alnus sinuata. Phenology: Flowers. Origin: Native. with Sharon Brown, Bob Antieau, Rena Hamburger Tammy Stout TLS2 23 Jun 2001 with Denise Liguori, Gail McHenry-Teller, Steve Rentmeester 23 Jun 2001 WTU-351650 WTU-351317, WTU-V-045612 Rosaceae Rosaceae Rubus pedatus Sm. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Mount Gardner. U.S.A., WASHINGTON, KING COUNTY: Elev. 3700 ft. Cascade Mountains of Western Washington: Cedar River. 47° 22' N, 121° 33' 5" W; T22N R10E S29 NE Elev. 601 ft. Collected at the intersection of roads 155, 155.5, and 155.6. Plants 47° 23' 28.9" N, 121° 55' 12.47" W; T22N R7E S16 SE 1/4 in area: Tsuga mertensiana, Pseudotsuga menziesii, Abies Along Cedar River at road 40.3, north of the river. Field notes lasiocarpa, Abiea amabalis, Alnus rubra, Polystichum munitum, include an extensive plant list. Phenology: Flowers. Origin: Native. Rubus spectabilis, Gaultheria shallon. Growing in wet alpine meadow; lots of course woody debris, full sun. Phenology: Richard G. Olmstead 01-7 26 May 2001 Vegetative. Origin: Native. with Jeremiah Ralston, Rich Gwodz, Liz Stenning, Mark Joselyn Tammy Stout SAR2 23 Jun 2001 WTU-351440 with Denise Liguori, Gail McHenry-Teller, Steve Rentmeester WTU-351649 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: U.S.A., WASHINGTON, KING COUNTY: Cedar River. Cascade Mountains of Western Washington: Elev. 601 ft. Eagle Ridge Creek Marsh. 47° 23' 28.9" N, 121° 55' 12.47" W; T22N Elev. 1620 ft. R7E S16 SE 1/4 47° 21' 42.1" N, 121° 37' 59.4" W; T22N R9E Along Cedar River at road 40.3, north of the S27 NE 1/4 river. Field notes include an extensive plant list. Collected between road 100-300 and midway Nearly past flowering. Deep pink flowers. into the marsh, near east end of marsh. Phenology: Flowers & Fruits. Origin: Native. Phenology: Flowers & Fruits. Origin: Native. Richard G. Olmstead 01-14 Judy Harpel 01-7 with Jeremiah Ralston, Rich Gwodz, Liz with Wayne Harpel, Rena Hamburger Stenning, Mark Joselyn 26 May 2001 26 May 2001 WTU-351495, WTU-V-18346 WTU-351433, WTU-V-038474

Rosaceae Rubus lasiococcus A. Gray U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Rex River headwaters, along both sides of road 700, approximately 1 mile north of Rex Pond. Elev. 3600 ft. T21N R9E S13 NW 1/4; WGS 84, uncertainty: 500 m., Source: Georeferenced, Georef'd by WTU Staff Collected along both sides of road 700, approximately 1 mile north of Rex Pond. Subalpine meadow with young Abies amabalis. White flowers. Abundant. Phenology: Flowers. Origin: Native. Eric Beach 01-7-EWB 14 Jul 2001 with Kate Bricket, Linda Cooley, Kristie Dunkin, Denise Liguori WTU-351628 Rosaceae Rosaceae Rubus pedatus Sm. Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, SKAMANIA COUNTY: Cascade Mountains of Western Washington: Rex River headwaters, Gifford - Pinchot National Forest. Riparian corridor along unnamed along both sides of road 700, approximately 1 mile north of Rex creek - Forest Service Road 6615 at bridge crossing. Pond. Elev. 3054 ft. Elev. 3600 ft. 45° 55.298' N, 121° 42.983' W T21N R9E S13 NW 1/4; WGS 84, uncertainty: 500 m., Source: Populus balsamifera, Picea engelmanii, and Pseudotsuga menziesii. Georeferenced, Georef'd by WTU Staff Magenta-pink flowers - pendent. Growing in partial shade thickets on Collected along both sides of road 700, approximately 1 mile north of creek bank. Phenology: Flowers. Origin: Native. Rex Pond. Subalpine meadow with young Abies amabalis. Abundant. Phenology: Flowers. Origin: Native. Sharon Rodman 573 16 Jun 2001 Eric Beach 01-112-KTB 14 Jul 2001 WTU-352050 with Kate Bricket, Linda Cooley, Kristie Dunkin, Denise Liguori WTU-351632 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus spectabilis Pursh U.S.A., WASHINGTON, SKAMANIA COUNTY: Trout Lake Campground. U.S.A., WASHINGTON, KING COUNTY: Elev. 2237 ft. Cascade Mountains of Western Washington: 46° 3' N, 121° 36.896' W; T7N R10E S31 Lost Creek (historic). Phenology: Flowers. Origin: Native. Elev. 2600 ft. T22N R8E S4 & 11; NAD 27, uncertainty: 805 Don Knoke 144 17 Jun 2001 m., Source: TRS2LL, Georef'd by Ben Legler Collected west of creek and north of road 800. WTU-352027 Tsuga heterophylla old growth forest. Growing along road. Leaf material removed for RADseq analysis by Megan Ruffley at WTU, February Rosaceae 2016. Phenology: Flowers. Origin: Native. Rubus spectabilis Pursh Denise Liguori 01-9 with Cindy Spurgeon, Kris DeLancey, Tara U.S.A., WASHINGTON, SKAGIT COUNTY: Fletcher 2 miles SW of Conway; WDFW access point at end of Mann Road, along dike between parking area and slough of Skagit River. 26 May 2001 Elev. 10 ft. 48° 19' 29.6" N, 122° 22' 27.6" W WTU-351677, WTU-V-019102 Under Alnus rubra. Common understory plant; flowers deep pink. Phenology: Flowers & Fruits. Origin: Native. Ben Legler 134 23 Mar 2003 Rosaceae WTU-352530 Rubus parviflorus Nutt. U.S.A., WASHINGTON, SKAMANIA COUNTY: Rosaceae Gifford Pinchot National Forest: Trout Lake Campground. Elev. 2237 ft. Rubus pedatus Sm. 46° 3' N, 121° 36.896' W; T7N R10E S31 Uneven ground along stream. Pseudotsuga menziesii, Acer U.S.A., WASHINGTON, WHATCOM COUNTY: circinatum, Corylus cornuta, Symphoricarpos mollis. Various North Cascades National Park; Thunder Creek Drainage; west of locations in campground. Phenology: Flowers. Origin: Native. footbridge at south end of Thunder Arm (Diablo Lake). Elev. 1241 ft. Don Knoke 01-114 14 Jun 2001 48° 40.712' N, 121° 4.79' W; UTM Zone 10, 641348.618E, 5393280.981N; Source: Field notes, UTM calculated from Lat-Lon. WTU-351892 East facing slope; mixed hemlock, Douglas fir, and red-cedar forest with understory of Acer circinatum. Creeping plants; 5 white petals. GPS accuracy 38 feet. Phenology: Flowers. Origin: Native. Rosaceae Sarah Gage 7230 14 Jun 2002 Rubus lasiococcus A. Gray WTU-353289, NPS accession 632, catalog 22398 U.S.A., WASHINGTON, SKAMANIA COUNTY: Gifford Pinchot National Forest: Forest Road 60 at Dry Creek Bridge. T5N R9E S6; WGS 84, uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff Roadside near stream, gravelly soil. Douglas fir, Alnus rubra, Berberis nervosa. Roqdside under shrubs. Phenology: Flowers. Origin: Native. Don Knoke 01-141 16 Jun 2001 WTU-351968 Rosaceae Rosaceae Rubus pedatus Sm. Rubus parviflorus Nutt. U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, WHATCOM COUNTY: Little Beaver Trail, about 1 mile east of Perry Creek; North Cascades North Cascades National Park; Stetattle Creek National Park. Trail, along Stetattle Creek just north of the Elev. 2034 ft. trailhead. 48° 55.3' N, 121° 8.1' W; UTM Zone 10, 636625.715E, Elev. 1329 ft. 5420205.349N; Source: Field notes, UTM calculated from Lat-Lon. 48° 43.783' N, 121° 9.342' W; UTM Zone 10, Dry gneiss talus, adjacent Douglas-fir forest, and adjacent cold 635626.483E, 5398832.286N; Source: Field stream bank just below talus; southern aspect. Creeping; growing at notes, UTM calculated from Lat-Lon. shaded edge of forest. Phenology: Flowers. Origin: Native. Overstory of Acer macrophyllum, Alnus rubra; understory of Rubus parviflorus, Rubus Ben Legler 27 13 Jul 2002 spectabilis. Common along trailside. Phenology: Flowers. Origin: Native. NOCA-353680, NPS accession 632, catalog 22841 Jeff Walker 16 with Sarah Gage, Nancy Job, Amy Dearborn Rosaceae 15 Jun 2002 Rubus pedatus Sm. WTU-354027, WTU-V-045454, NPS accession U.S.A., WASHINGTON, SNOHOMISH COUNTY: 632, catalog 23600 1 mile SE of Granite Falls; on private residence along Coon Creek accessed from 84th St NE on east side of Robe-Mentzel road. Elev. 380 ft. Rosaceae 48° 4' 22" N, 121° 57' 34" W Ditched creek and wetland in open wet field with young shrubs, Rubus ursinus Cham. & Schltdl. surrounded by forested slopes. Petals white, 8 mm long; growing in moss under conifers. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, WHATCOM COUNTY: Ross Lake National Recreation Area; Pyramid Lake, west of Colonial Ben Legler 350 4 May 2003 Creek Campground. Elev. 2685 ft. WTU-353647 48° 41.69' N, 121° 7.404' W; UTM Zone 10, 638097.116E, 5395012.922N; Source: Field notes, UTM calculated from Lat-Lon. Rocky slopes and dry coniferous forest surrounding lake. Growing Rosaceae around edge of lake and on a rock face at south end of lake. Phenology: Flowers. Origin: Native. Rubus pedatus Sm. Cindy Spurgeon CHS 11 15 Jun 2002 U.S.A., WASHINGTON, CHELAN COUNTY: North Cascades National Park; Southeast of Cascade Pass; WTU-354102, NPS accession 632, catalog 23917 begining of Ptarmigan Traverse; northeast-facing slope. Elev. 5588 ft. 48° 27.909' N, 121° 3.586' W; UTM Zone 10, 643428.125E, Rosaceae 5369601.027N; Source: Field notes, UTM calculated from Lat-Lon. Rocky (granite) slope with mostly Phyllodoce empetriformis and Rubus spectabilis Pursh other sub-shrubs; some short Tsuga mertensiana. Petals white, stamens white; on steep incline, partly on rock under tree. U.S.A., WASHINGTON, KITSAP COUNTY: Phenology: Flowers. Origin: Native. Mouth of Big Beef Creek at University of Washington's Big Beef Creek Field Station, 9 Sarah Gage 7377 10 Aug 2002 miles NW of Bremerton. with Diane Hennessey, Laura Becca Elev. 5 ft. 47° 39' 7.3" N, 122° 46' 58.8" W WTU-354125, NPS accession 632, catalog 22358 Moist gravelly soil around facilities; disturbed ground. Common shrub to 2 meters tall; flowers deep pink. Phenology: Flowers. Origin: Native. Rosaceae Ben Legler 312 Rubus pedatus Sm. with David Giblin U.S.A., WASHINGTON, WHATCOM COUNTY: 3 May 2003 North Cascades National Park; Perry Creek Shelter, near confluence of Perry Creek and Little Beaver Creek. WTU-353851, WTU-V-18370 Elev. 2350 ft. 48° 55.2' N, 121° 8.9' W; UTM Zone 10, 635653.508E, 5419996.201N; Source: Field notes, UTM calculated from Lat-Lon. Shade of Thuja plicata and Pseudotsuga menziesii, with thick duff and loose soil. Phenology: Flowers. Origin: Native. Justin Hohn 19 13 Jul 2002 WTU-353803, NPS accession 632, catalog 23159 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, WAHKIAKUM U.S.A., WASHINGTON, ISLAND COUNTY: COUNTY: Whidbey Island, Ebey's Landing; Hill Road c. 1 4 miles north of Oneida, along Deep River km South of main parking lot, just below summit Valley Road ca. 1.5 miles north of junction with of road. Bluff to west of road. Anderson road. Elev. 171 ft. Elev. 50 ft. 48° 11' 17" N, 122° 42' 5" W; NAD 27 46° 22' 43.5" N, 123° 40' 8.9" W Steep bluff dominated by native and exotic Wet deciduous forest along small stream, grasses with Lupinus, Eriophyllum, and Achillea; dominated by Alnus rubra, Rubus spectabilis, backed by conifer woods. In woods behind bluff. Lysichiton americanum. Flowers deep pink; Phenology: Flowers. Origin: Native. shrub to ca. 2 meters tall. Phenology: Flowers & Fruits. Origin: Native. Peter W. Dunwiddie PWD 77 Ben Legler 522 30 Apr 2004 1 Jun 2003 WTU-355527, WTU-V-044008 WTU-354322, WTU-V-18369 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, ISLAND COUNTY: U.S.A., WASHINGTON, WAHKIAKUM COUNTY: Whidbey Island, The Nature Conservancy's 4 miles north of Oneida, along Deep River Valley Road ca. 1.5 miles Ebey's Landing Preserve; along top of bluff north of junction with Anderson road. above Perego's Lagoon at about midway point Elev. 50 ft. of lagoon. 46° 22' 43.5" N, 123° 40' 8.9" W Elev. 200 ft. Wet deciduous forest along small stream, dominated by Alnus rubra, 48° 12' 7.6" N, 122° 43' 54.1" W Rubus spectabilis, Lysichiton americanum. Petals white; trailing Open grassy bluff and Pseudotsuga woods. shrub along road. Phenology: Flowers. Origin: Native. Petals deep pink; shrub in woods. Phenology: Flowers. Origin: Native. Ben Legler 523 1 Jun 2003 Ben Legler 1586 WTU-354674 with David Giblin, Barbara Williams, Molly Bernstein Rosaceae 7 May 2004 Rubus parviflorus Nutt. WTU-355891, WTU-V-044009 U.S.A., WASHINGTON, WAHKIAKUM COUNTY: 4 miles north of Oneida, along Deep River Valley Road ca. 1.5 miles north of junction with Anderson road. Rosaceae Elev. 50 ft. 46° 22' 43.5" N, 123° 40' 8.9" W Rubus spectabilis Pursh Wet deciduous forest along small stream, dominated by Alnus rubra, Rubus spectabilis, Lysichiton americanum. Petals white; shrub U.S.A., WASHINGTON, PACIFIC COUNTY: growing along road. Phenology: Flowers. Origin: Native. The Nature Conservancy's Ellsworth Creek Preserve along East bank of upper Ellsworth Creek. Ben Legler 524 1 Jun 2003 Elev. 200 ft. 46° 22' 55" N, 123° 52' 37.4" W WTU-354673 Deep shade, old growth Tsuga/Thuja, west aspect, moderate slope. Deep pink flower, fruit green, spiny shrub. Phenology: Flowers & Fruits. Origin: Native. Rosaceae Ben Legler 481 31 May 2003 Rubus ursinus Cham. & Schltdl. WTU-356840 U.S.A., WASHINGTON, ISLAND COUNTY: Whidbey Island, Ebey's Landing National Historic Preserve Bluff Trail. 2 KM north from main parking lot. Along Trail and nearby Rosaceae woods at top of bluff. 48° 11.82' N, 122° 43.183' W Rubus parviflorus Nutt. Sandy, west-facing, exposed bluff dominated by grasses, Achillea, Vicia; wind-swept Pseudotsuga forest with Ribes, Berberis. Common U.S.A., WASHINGTON, PACIFIC COUNTY: in understory. Phenology: Flowers. Origin: Native. The Nature Conservancy's Ellsworth Creek preserve, 5 miles NW of Naselle on east side of mouth of creek, under powerlines. Peter W. Dunwiddie PWD 45 30 Apr 2004 Elev. 100 ft. 46° 24' 58.2" N, 123° 53' 8.9" W WTU-355624 Open area with grasses and weeds. Petals white. Phenology: Flowers. Origin: Native. Ben Legler 492 31 May 2003 WTU-356837 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, ISLAND COUNTY: U.S.A., WASHINGTON, PACIFIC COUNTY: Whidbey Island, Ebey's Landing Nature Conservancy Preserve; The Nature Conservancy's Ellsworth Creek preserve, 5 miles NW of south end of Perego's Lagoon. Naselle on east side of mouth of creek, under powerlines. Elev. 5 ft. Elev. 100 ft. 48° 11' 50" N, 122° 43' 30" W 46° 24' 58.2" N, 123° 53' 8.9" W One patch; in fruit. Phenology: Fruits. Origin: Introduced. Open area, with grasses and weeds. Petals white. Phenology: Flowers. Origin: Native. Jeff Walker 287 10 Sep 2004 Ben Legler 490 31 May 2003 WTU-357059 WTU-357095 Rosaceae Rosaceae Rubus bifrons Vest Rubus sachalinensis H. Lév. U.S.A., WASHINGTON, ISLAND COUNTY: Whidbey Island, Ebey's Landing Nature Conservancy Preserve; east RUSSIAN FEDERATION, SAKHALIN REGION: side of Perego's Lagoon. Sakhalin Island, central part; circa 6 kilometers north-northwest of Elev. 5 ft. Tymovskoye and 2 kilometers southwest of Voskressnovka, on east 48° 12' 6" N, 122° 43' 53" W bank of Tym River. Steep dry grassy slope; 2 meters tall, fruit black. Phenology: Fruits. Elev. 328 ft. Origin: Introduced. 50° 53' 26.22" N, 142° 38' 33.12" E Cut hay fields and deciduous riparian forest of Populus, Salix. Fruit Jeff Walker 269 10 Sep 2004 red; shrub 1 meter tall on edge of field. Phenology: Fruits. Origin: Native. WTU-357071 Ben Legler 1000 29 Jul 2003 Rosaceae WTU-357301 Rubus chamaemorus L. Rosaceae RUSSIAN FEDERATION, SAKHALIN REGION: Sakhalin Island, along west coast 5 km north of Krasnogorsk, at Rubus ursinus Cham. & Schltdl. south end of Lake Aynskoye near outlet. Elev. 0 ft. U.S.A., WASHINGTON, PACIFIC COUNTY: 48° 27' 13.14" N, 142° 3' 36.24" E The Nature Conservancy's Ellsworth Creek Preserve. Old logging Open grassy marsh on edge of lake. Creeping sub-shrub; berries road 0.5 km north of first gate on main road, west side of estuary. turning orange. Phenology: Fruits. Origin: Native. Elev. 70 ft. 46° 24.424' N, 123° 53.376' W Ben Legler 769 21 Jul 2003 Edge of clearcut and riparian strip; Alnus and Tsuga overstory; Rubus and Sambucus shrub layer. Trillium along main road. WTU-357008 Running vine; flowers white; abundant. Phenology: Flowers. Origin: Native. Rosaceae Jessie Johanson 03-23 31 May 2003 Rubus arcticus L. WTU-357430 RUSSIAN FEDERATION, SAKHALIN REGION: Rosaceae Sakhalin Island, along west coast 5 km north of Krasnogorsk, at south end of Lake Aynskoye near outlet. Rubus spectabilis Pursh Elev. 0 ft. 48° 27' 13.14" N, 142° 3' 36.24" E U.S.A., WASHINGTON, PACIFIC COUNTY: Open grassy marsh on edge of lake. Flowers pink-red; fruit The Nature Conservancy's Ellsworth Creek orange-red; scattered through marsh. Phenology: Flowers & Fruits. Preserve. Old logging road 0.5 km north of first Origin: Native. gate on main road, west side of estuary. Elev. 70 ft. Ben Legler 786 21 Jul 2003 46° 24.424' N, 123° 53.376' W Edge of clearcut and riparian strip; Alnus and WTU-356979 Tsuga overstory; Rubus and Sambucus shrub layer. Trillium along main road. Leaf material removed for RADseq analysis by Megan Ruffley at WTU, February 2016. Phenology: Flowers & Fruits. Origin: Native. Jessie Johanson 03-24 31 May 2003 WTU-357431, WTU-V-019104 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, PACIFIC COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: The Nature Conservancy's Ellsworth Creek Preserve. Old logging Along shore of Quinault Lake, Olympic Forest. road 0.5 km north of first gate on main road, west side of estuary. Elev. 200 ft. Elev. 70 ft. 47.46722°, -123.84417°; WGS 84, uncertainty: 10000 m., Source: 46° 24.424' N, 123° 53.376' W GeoLocate, Georef'd by WTU Staff Edge of clearcut and riparian strip; Alnus and Tsuga overstory; Slope; 40% W. Soil; gravelly clay loam. Type; timber. Browse. Rubus and Sambucus shrub layer. Trillium along main road. Large Associated plants; Vine Maple, Salmonberry, etc. Use; moderate. white flowers; common. Phenology: Flowers. Origin: Native. Distribution; moist soils. Abundance; common. Density in tenths; 8. Forage value; good. Grazed by E & D. Season grazed; winter- Jessie Johanson 03-28 31 May 2003 spring. Usual size of plant; 3?-7?. Flowering period; June. Period of seed dissemination; Fall. Herbarium of the Forest Service #80694. WTU-357435 Ingram. Phenology: Flowers. Origin: Native. J. E. Schwartz 57A 5 Jun 1936 Rosaceae WTU-355196 Rubus laciniatus Willd. U.S.A., WASHINGTON, PACIFIC COUNTY: Rosaceae The Nature Conservancy's Ellsworth Creek Preserve. Entrance to Yellow Gate Road. Rubus lasiococcus A. Gray Elev. 3 ft. 46° 25.6' N, 123° 53.675' W U.S.A., WASHINGTON, CHELAN COUNTY: Estuary with gravel road through it. Dominant species: Carex Okanogan National Forest. Lake Ann, north end of lake, 100 meters obnupta, C. lyngbei, and Holcus lanatus. Low growing to 30 cm west of outlet. height; flowers white. Phenology: Flowers. Origin: Introduced. Elev. 5520 ft. 48.506967°, -120.755633°; WGS 84, uncertainty: 500 m., Source: Marianne Spahr 03-11 9 Jul 2003 GeoLocate, Georef'd by WTU Staff Mesic meadow dominated by Erigeron, Arnica, Salix, Phyllodoce. WTU-357442 Stoloniferous; flowers white. Phenology: Flowers. Origin: Native. David Tank 03-10 27 Jul 2003 Rosaceae with Jessie Johanson, Cindy Spurgeon, Marianne Powell, David Giblin, Kara Ardern, Suzanne Bagshaw Rubus ursinus Cham. & Schltdl. WTU-362674 U.S.A., WASHINGTON, PACIFIC COUNTY: The Nature Conservancy's Ellsworth Creek Preserve. Waypoint #36. Along trail in forest tract just above logging spur road. Rosaceae Elev. 161 ft. 46° 23.68' N, 123° 53.054' W Rubus parviflorus Nutt. In disturbed opening along spur road. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, CHELAN COUNTY: Okanogan National Forest. Lake Ann Trail, 1 km. north of lake. Richard G. Olmstead 2003-38 31 May 2003 Elev. 5477 ft. 48.5105°, -120.746283°; WGS 84, uncertainty: 500 m., Source: WTU-357519 GeoLocate, Georef'd by WTU Staff Trailside. Forest edge and sloping subalpine meadow with occasional pockets of talus. In rocks; full sun. Phenology: Flowers. Rosaceae Origin: Native. Rubus spectabilis Pursh David Tank 03-56 27 Jul 2003 with Jessie Johanson, Cindy Spurgeon, Marianne Powell, David U.S.A., WASHINGTON, PACIFIC COUNTY: Giblin, Kara Ardern, Suzanne Bagshaw The Nature Conservancy's Ellsworth Creek Preserve. Waypoint #36. Along trail in forest Herbarium: WTU tract just above logging spur road. Elev. 161 ft. Rosaceae 46° 23.68' N, 123° 53.054' W In disturbed opening along spur road. Red Rubus lasiococcus A. Gray flowers. Phenology: Flowers & Fruits. Origin: Native. U.S.A., WASHINGTON, CHELAN COUNTY: Okanogan National Forest. Lake Ann Trail, 1 km. north of lake. Richard G. Olmstead 2003-40 Elev. 5477 ft. 48.5105°, -120.746283°; WGS 84, uncertainty: 500 m., Source: 31 May 2003 GeoLocate, Georef'd by WTU Staff Trailside. Forest edge and sloping subalpine meadow with WTU-357521, WTU-V-020803 occasional pockets of talus. Stoloniferous; flowers white. Phenology: Flowers. Origin: Native. David Tank 03-60 27 Jul 2003 with Jessie Johanson, Cindy Spurgeon, Marianne Powell, David Giblin, Kara Ardern, Suzanne Bagshaw WTU-363131 Rosaceae Rosaceae Rubus pedatus Sm. Rubus bifrons Vest U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Okanogan National Forest. Rainy Lake Outlet. San Juan Islands National Wildlife Refuge and Elev. 5010 ft. Wilderness Area. Castle Island. UTM Zone 10, 667387E, 5374601N; NAD 27, uncertainty: 50 m., Elev. 0-98 ft. Source: Georeferenced, Georef'd by WTU Staff 48.42104° N, 122.82176° W; T34N R1W Sunny glade; "seep" according to Suzanne; lady ferns, Athyrium. S21; WGS 84, uncertainty: 100 m., Source: Dominant trees: silver fir, mountain hemlock, subalpine fir, Engelman Digital map spruce. From tree-island growing with Yen 03-041; 5 leaf bramble. Steep, cliffy island with many rock ledges, Phenology: Flowers. Origin: Native. cracks and beaches. Few scattered individuals on south cliff, at summit. Phenology: Fruits. Alan C. Yen 03-042 25 Jul 2003 Origin: Introduced. with Richard Robohm, Suzanne Bagshaw, Liz Van Volkenburgh, Therese Ohlson Ben Legler 2572 WTU-363220 28 Apr 2005 WTU-384318, WTU-V-060739 Rosaceae Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, CHELAN COUNTY: Okanogan National Forest. Rainy Lake Outlet. Rubus bifrons Vest Elev. 5010 ft. UTM Zone 10, 667387E, 5374601N; NAD 27, uncertainty: 50 m., U.S.A., WASHINGTON, SAN JUAN COUNTY: Source: Georeferenced, Georef'd by WTU Staff San Juan Islands National Wildlife Refuge and Wilderness Area. Sunny glade; "seep" according to Suzanne; lady ferns, Athyrium. Colville Island. Dominant trees: silver fir, mountain hemlock, subalpine fir, Engelman Elev. 0-65 ft. spruce. Salmonberry. Phenology: Flowers & Fruits. Origin: Native. 48° 24' 55" N, 122° 49' 21" W; Source: Specimen label Small island with vertical cliffs on all sides. Topography is Alan C. Yen 03-042a 25 Jul 2003 undulating, with large outcrops of smooth rock, covered with foliose with Richard Robohm, Suzanne Bagshaw, Liz Van Volkenburgh, and crustose lichens. All aspects represented. Vegetation is Therese Ohlson dominated by exotic taxa including large swards of Bromus sterilis and thickets of Rubus discolor and Rubus lacinatus. Native WTU-363221 vegetation is scattered and patchy. Thickets of Rosa nutkana occur in swales. Common; scattered. Phenology: Flowers. Origin: Introduced. Rosaceae David Giblin 05-3 28 Apr 2005 Rubus parviflorus Nutt. f. parbifarius Fassett WTU-383632 U.S.A., CALIFORNIA, ALAMEDA COUNTY: Berkeley, 0.7 mile above mouth of Strawberry Rosaceae Creek Canyon. Elev. 1100 ft. Rubus spectabilis Pursh 37.879502°, -122.245819°; WGS 84, uncertainty: 1358 m., Source: Digital Map, U.S.A., WASHINGTON, SAN JUAN COUNTY: Georef'd by Sara Legler San Juan Islands National Wildlife Refuge and Thickets on north slope of canyon. Associated Wilderness Area. Colville Island. with Umbellularia, Quercus agrifolia, Corylus, Elev. 0-65 ft. Salix, Sambucus, Physocarpus, 48° 24' 55" N, 122° 49' 21" W; Source: Symphoricarpos, Aralia, and Rubus vitifolius. Specimen label Shrub 5-8 feet high, with exfoliating brown bark, Small island with vertical cliffs on all sides. berries red. Phenology: Fruits. Origin: Native. Topography is undulating, with large outcrops of Type Specimen: Isotype. smooth rock, covered with foliose and crustose lichens. All aspects represented. Vegetation is L. Constance 2397 4 Aug 1938 dominated by exotic taxa including large swards of Bromus sterilis and thickets of Rubus discolor WTU-275322, WTU-V-000397 and Rubus lacinatus. Native vegetation is scattered and patchy. Thickets of Rosa nutkana occur in swales. Uncommon on north face. Rosaceae Phenology: Flowers. Origin: Native. Rubus ursinus Cham. & Schltdl. David Giblin 05-22 28 Apr 2005 U.S.A., WASHINGTON, SAN JUAN COUNTY: WTU-384857, WTU-V-18372 San Juan Islands National Wildlife Refuge and Wilderness Area. Castle Island. Elev. 0-98 ft. 48.42104° N, 122.82176° W; T34N R1W S21; WGS 84, uncertainty: 100 m., Source: Digital map Steep, cliffy island with many rock ledges, cracks and beaches. Occasional; clambering over rock and grassland. Phenology: Flowers. Origin: Native. Ben Legler 2549 28 Apr 2005 WTU-383230 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus laciniatus Willd. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. Katon Road at Peel Slough, north side of Chehalis River, 2 airmiles Colville Island. southwest of Montesano. Elev. 0-65 ft. Elev. 16 ft. 48° 24' 55" N, 122° 49' 21" W; Source: Specimen label 46° 57.4' N, 123° 38.3' W; T17N R8W S14 Small island with vertical cliffs on all sides. Topography is Wet edge of floodplain forest; associated with Rubus armeniacus, undulating, with large outcrops of smooth rock, covered with foliose Alnus rubra. Pink flowers, most fruit unripe, 2.5 meters arching and crustose lichens. All aspects represented. Vegetation is adventive shrubs. Phenology: Flowers & Fruits. Origin: Introduced. dominated by exotic taxa including large swards of Bromus sterilis and thickets of Rubus discolor and Rubus lacinatus. Native Peter F. Zika 20228 15 Sep 2004 vegetation is scattered and patchy. Thickets of Rosa nutkana occur in swales. Phenology: Flowers. Origin: Native. WTU-359712 David Giblin 05-29a 28 Apr 2005 Rosaceae WTU-384879 Rubus laciniatus Willd. Rosaceae U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Small tributary to Wynoochee River, Wyoochee Tracts Road, 1.3 air Rubus ursinus Cham. & Schltdl. miles west of Montesano. Elev. 16 ft. U.S.A., WASHINGTON, SAN JUAN COUNTY: 46° 58.8' N, 123° 38' W; T17N R8W S12 San Juan Islands National Wildlife Refuge and Associated with Equisetum telmateia, Rubus armeniacus, Glyceria, Wilderness Area. Aleck Rock. South of Lopez Iris pseudacorus. Black fruit, 2 meters arching adventive shrubs, by Island. creek. Phenology: Flowers & Fruits. Origin: Introduced. Elev. 0-32 ft. 48° 25.399' N, 122° 51.038' W; Source: Peter F. Zika 20218 15 Sep 2004 Specimen label Grassy meadow with rocky outcrops. WTU-359715 Phenology: Flowers. Origin: Native. Joseph Arnett 05-15 Rosaceae 26 Apr 2005 Rubus bifrons Vest WTU-384262, WTU-V-021000 U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Small tributary to Wynoochee River, Wyoochee Tracts Road, 1.3 air miles west of Montesano. Elev. 16 ft. Rosaceae 46° 58.8' N, 123° 38' W; T17N R8W S12 Associated with Equisetum telmateia, Rubus laciniatus, Glyceria, Iris Rubus laciniatus Willd. × Rubus macrophyllus Weihe & pseudacorus. Black fruit, common 2 meter arching adventive shurbs, Nees by creek. Phenology: Fruits. U.S.A., WASHINGTON, PACIFIC COUNTY: Peter F. Zika 20217 15 Sep 2004 Both sides of Route 101, 5 air kilometers east northeast of Stoney Point, Willapa Bay, Stuart Slough. WTU-359722 Elev. 16 ft. 46° 41.2' N, 123° 52.3' W; T14N R10W S24 Roadside thickets, full sun, with Rubus armeniacus, Rubus Rosaceae laciniatus, Rubus ursinus, Rubus macrophyllus. Uncommon adventive evergreen shrub 1.7 meters tall; infl glandular; fruit black, Rubus laciniatus Willd. shiny, truncate base, 17 X 18 milimeters wide. Phenology: Fruits. Origin: Introduced. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Small tributary to Wynoochee River, Wyoochee Tracts Road, 1.3 air Peter F. Zika 20360 29 Sep 2004 miles west of Montesano. Elev. 16 ft. WTU-359826 46° 58.8' N, 123° 38' W; T17N R8W S12 Associated with Equisetum telmateia, Rubus armeniacus, Glyceria, Rosaceae Iris pseudacorus. Black fruit, 2 meters arching adventive shrubs, by creek. Phenology: Fertile. Origin: Introduced. Rubus laciniatus Willd. Peter F. Zika 20218 15 Sep 2004 U.S.A., WASHINGTON, PACIFIC COUNTY: Both sides of Route 101, 5 air kilometers east northeast of Stoney WTU-359723 Point, Willapa Bay, Stuart Slough. Elev. 16 ft. 46° 41.2' N, 123° 52.3' W; T14N R10W S24 Roadside thickets, full sun, with Rubus armeniacus, Rubus ursinus, Rubus macrophyllus. Uncommon adventive evergreen shrub, canes dark red, arching, 1.5 meters high; fruit black, shiny, ovoid globose, truncate base, 11-16 X 12-16 milimeters wide; infl. glandular; leaves pale green below, glandular. Phenology: Fruits. Origin: Introduced. Peter F. Zika 20355 29 Sep 2004 WTU-359647 Rosaceae Rosaceae Rubus bifrons Vest Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Katon Road at Peel Slough, north side of Chehalis River, 2 airmiles San Juan Islands National Wildlife Refuge and Wilderness Area. southwest of Montesano. Flint Beach Island. Elev. 16 ft. Elev. 0-49 ft. 46° 57.4' N, 123° 38.3' W; T17N R8W S14 48° 25.202' N, 122° 52.036' W; Source: Specimen label Wet edge of floodplain forest, with Rubus laciniatus, Alnus rubra. Trailing across Rosa and grassland; occasional. Phenology: Overripe black fruit, 2.5 meters arching adventive shrubs. Flowers. Origin: Native. Phenology: Fruits. Peter F. Zika 20497 27 Apr 2005 Peter F. Zika 20229 15 Sep 2004 WTU-384000 WTU-359162 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus laciniatus Willd. U.S.A., WASHINGTON, COWLITZ COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: East bank of Carrolls Channel. Opposite south end of Cottonwood Katon Road at Peel Slough, north side of Chehalis River, 2 airmiles Island, Columbia River. southwest of Montesano. Elev. 32 ft. Elev. 16 ft. 46° 3.7' N, 122° 52' W; T7N R2W S30 46° 57.4' N, 123° 38.3' W; T17N R8W S14 Steep rocky east bank of channel. Occassional. Flowers white, 1 Wet edge of floodplain forest, with Rubus armeniacus, Rubus meter erect to arching shrub, with Rubus armeniacus, R. laciniatus. laciniatus, Alnus rubra. Black fruit 18-24 X 17-19 milimeters weight; Phenology: Flowers. Origin: Native. 2.5 meters arching adventive shrubs. Phenology: Fruits. Peter F. Zika 19470 4 May 2004 Peter F. Zika 20230 15 Sep 2004 WTU-360022 WTU-359157 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, PACIFIC COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Both sides of Route 101, 5 air kilometers east northeast of Stoney San Juan Islands National Wildlife Refuge and Wilderness Area. Point, Willapa Bay, Stuart Slough. Boulder Island. Elev. 16 ft. Elev. 0-65 ft. 46° 41.2' N, 123° 52.3' W; T14N R10W S24 48° 25.92' N, 122° 48.158' W; Source: Specimen label Roadside thickets, full sun, with Rubus armeniacus, R. ursinus, R. Open grassy 4 acre(?) island with rock/cliff shores. Only trees are macrophyllus. Uncommon adventive evergreen shrub, canes dark several dozen stunted Pseudotsuga menziesii. Uncommon over red, arching, 1.5 meters high; fruit black, shiny, ovoid globose, rocky outcrops. Phenology: Flowers. Origin: Native. truncate base, 11-16 x 12-16 millimeters wide; infl. glandular; leaves pale green below, glandular. Phenology: Flowers & Fruits. Origin: Peter Dunwiddie 05-35 26 Apr 2005 Introduced. WTU-384159 Peter F. Zika 20355 29 Sep 2004 WTU-360180 Rosaceae Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: Rubus laciniatus Willd. × Rubus macrogongylus Focke San Juan Islands National Wildlife Refuge and Wilderness Area. Charles Island, southwest corner of island. U.S.A., WASHINGTON, PACIFIC COUNTY: Elev. 15 ft. Both sides of Route 101, 5 air kilometers east northeast of Stoney 48° 26.601' N, 122° 54.804' W; Source: Specimen label Point, Willapa Bay, Stuart Slough. Grassy headland. Scattered; clambering over Symphoricarpos and Elev. 16 ft. Rosa. Phenology: Flowers. Origin: Native. 46° 41.2' N, 123° 52.3' W; T14N R10W S24 Roadside thickets, full sun, with Rubus armeniacus, R. laciniatus, R. Steve Hahn 05-29 27 Apr 2005 ursinus, R. macrophyllus. Uncommon adventive evergreen shrub 1.7 meters tall. Infloresecense glandular; fruit black, shinny, truncate WTU-385174 base, 17 x 18 millimeters wide. [Sheet 2 of 2]. Phenology: Vegetative. Origin: Introduced. Peter F. Zika 20360 29 Sep 2004 WTU-360171 Rosaceae Rosaceae Rubus vestitus Weihe & Nees Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, PACIFIC COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Both sides of Route 101, 5 air kilometers east northeast of Stoney San Juan Islands National Wildlife Refuge and Wilderness Area. Point, Willapa Bay, Stuart Slough. Buck Island. Southwest edge of Lopez Island, with frequent rock Elev. 16 ft. outcroppings on top. 46° 41.2' N, 123° 52.3' W; T14N R10W S24 Elev. 9-39 ft. Roadside thickets, full sun, with Rubus armeniacus, R. laciniatus, R. 48° 27.146' N, 122° 55.194' W; Source: Specimen label ursinus. Occasional adventive evergreen shrub, canes purple; Flat-topped meadow dominated by Rosa, Festuca, Grindelia, infloresecense glandular; fruit black; shiny, truncate ovate, 11-19 x Elymus, and Holcus. Vegetation compacted in many areas from 14-17 millimeters wide; 8-36 seed per fruit; leaves green below. animal activities. A few scattered plants. Phenology: Flowers. Origin: Phenology: Fruits. Origin: Introduced. Native. Peter F. Zika 20359 29 Sep 2004 Elizabeth Bell 05-48 29 May 2005 WTU-360181 WTU-382292 Rosaceae Rosaceae Rubus bifrons Vest Rubus bifrons Vest U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, PACIFIC COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. Both sides of Route 101, 5 air kilometers east northeast of Stoney Rum Island. Point, Willapa Bay, Stuart Slough. Elev. 16-32 ft. Elev. 16 ft. 48° 28.798' N, 122° 49.753' W; WGS 84 46° 41.2' N, 123° 52.3' W; T14N R10W S24 Island is narrow and runs north-south direction with forested area in Roadisde thickets, full sun, with Rubus laciniatus, R. ursinus, R. center. Dominated by Arbutus, Abies, Pseudotsuga, Quercus macrphyllus. Common adventive evergreen shrub, canes dark overstory, and Bromus pacificus, Bromus mollis, Bromus rigidus purple-red, arching, 2 meters high; fruit black, shiny, ovate, truncate understory. One plant in middle of island. Phenology: Flowers. base, 14-20 x 13-18 millimeters wide; 17-44 sd/frt; leaves white Origin: Introduced. tomentose below. Phenology: Flowers & Fruits. Jessie Johanson 05-10 29 May 2005 Peter F. Zika 20357 29 Sep 2004 with Joe Johanson WTU-360182 WTU-381674

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, ASOTIN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Fields Spring State Park, found in picnic area San Juan Islands National Wildlife Refuge and Wilderness Area. near "Field's Spring." Rum Island. T7N R45E S16; WGS 84, uncertainty: 200 m., Elev. 16-32 ft. Source: Georeferenced, Georef'd by WTU 48° 28.798' N, 122° 49.753' W; WGS 84 Staff Island is narrow and runs north-south direction with forested area in Hillside with Pseudotsuga menziesii, Abies center. Dominated by Arbutus, Abies, Pseudotsuga, Quercus grandis in overstory. Growing on slope with overstory, and Bromus pacificus, Bromus mollis, Bromus rigidus Trautvetteria caroliniensis and Acer glabrum. understory. Frequent in understory. Phenology: Flowers. Origin: Phenology: Flowers & Fruits. Origin: Native. Native. Jeff Walker 186 Jessie Johanson 05-15 29 May 2005 with Joe Johanson 26 Jun 2004 WTU-381675 WTU-360420, WTU-V-045447 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus spectabilis Pursh U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. San Juan Islands National Wildlife Refuge and Wilderness Area. Fortress Island. Buck Island. Southwest edge of Lopez Island, with frequent rock Elev. 16-65 ft. outcroppings on top. 48° 27.922' N, 122° 50.284' W; Source: Specimen label Elev. 9-39 ft. Treeless, 3.2 acre, island with steep north-facing slope. South-facing 48° 27.146' N, 122° 55.194' W; Source: Specimen label slope dominated by Grindelia, Sedum, Holcus lanatus. Shrubs Flat-topped meadow dominated by Rosa, Festuca, Grindelia, include Rosa nutkana and Holodiscus discolor; moist swales and Elymus, and Holcus. Vegetation compacted in many areas from seeps sporadic. Uncommon on northwest-face. Phenology: Flowers. animal activities. A couple of shrubs on northeast side, next to cliff. Origin: Native. Phenology: Flowers. Origin: Native. L. A. Brooking 05-38 28 May 2005 Elizabeth Bell 05-43 29 May 2005 WTU-384807 WTU-382296 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus lasiococcus A. Gray U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. Mount Rainier National Park, upper Stevens Ridge 0.5 miles due Deadman Island. north of Fairy Falls. Elev. 9-32 ft. Elev. 6260 ft. 48° 27.5' N, 122° 56.629' W; Source: Specimen label 46° 48' 7.3" N, 121° 42' 10.1" W; UTM Zone 10, 598983E, Bedrock knob ca. [circa] 10 meters tall, with numerous crevices, 5183980N; WGS 84, uncertainty: 5 m., Source: GPS several shrub thickets. Occasional; trailing. Phenology: Flowers. Gentle south-facing mesic meadow, with Phlox diffusa, Potentilla Origin: Native. drummondii, Carerx sp., scattered trees on edge. Flowers white; common under trees. Phenology: Flowers. Origin: Native. Louisa C. Dunwiddie 05-10 30 May 2005 Deanna Goldy 05-15 2 Aug 2005 WTU-382085 with Sharon Rodman, Ben Legler, Don Knoke, Pete Del Zotto WTU-366223, NPS accession 489, catalog 15511 Rosaceae Rubus pubescens Raf. Rosaceae U.S.A., WASHINGTON, PEND OREILLE COUNTY: Rubus bifrons Vest Cedar swale on south side of Forest Service Road 2695 ca. 1/4 mile east of east end of Big Meadow Lake and 0.2 miles west of junction U.S.A., WASHINGTON, SPOKANE COUNTY: of Rd 2695 with Rd 430. Southwest end of Finch Arboretum, near Woodland Boulevard, Elev. 3470 ft. Spokane. 48° 43' 48" N, 117° 32' 42" W; WGS 84 Elev. 1984 ft. Gently sloping gully with small sluggish stream under dense shade 47° 38.6' N, 117° 27.7' W of mature Thuja plicata and scattered Populus balsamifera, with Partial shade, with Acer platanoides. 1.5 meters tall arching shrubs, Botrychium virginianum, B. montanum, Asarum caudatum, Smilacina rare weed. Phenology: Flowers. stellata, Actaea rubra, Rubus pubescens, Alnus sinuata. Small patch of plants in muddy saturated soil on edge of shady pool. Phenology: Peter F. Zika 20309 19 Sep 2004 Fertile. Origin: Native. WTU-360958 Ben Legler 2876 24 Jul 2005 WTU-361230 Rosaceae Rubus bifrons Vest Rosaceae U.S.A., WASHINGTON, WHITMAN COUNTY: Rubus ursinus Cham. & Schltdl. Mouth of Wawawai Canyon on east shore of Snake River, Wawawai. Elev. 721 ft. U.S.A., WASHINGTON, SAN JUAN COUNTY: 46° 38.2' N, 117° 22.5' W; T13N R43E S2 San Juan Islands National Wildlife Refuge and Wilderness Area. Edge of cattail dominated wetland, with Typha latifolia, Salix exigma, Iceberg Island. Xanthium, Scirpus microcarpus. Fruit black, leaves white beneath; Elev. 0-30 ft. common arching shrub. Phenology: Flowers & Fruits. 48° 25.625' N, 122° 53.354' W; WGS 84 Island is low, ca. [circa] 30 feet high, relatively flat, with extensive P. F. Zika 20124 30 Aug 2004 bedrock outcrops throughout. Sedum lanceolatum is abundant with R. Old across the bedrock. A large swale in mid-island is dense Rosa nutkana thicket. A couple of small vernal pools in the bedrock near WTU-361026 the south side are noteworthy. Scrambling vine; uncommon. Phenology: Flowers & Fruits. Origin: Native. Rosaceae Lisa L. Bellefond 05-28 29 May 2005 Rubus bifrons Vest WTU-382015 U.S.A., WASHINGTON, DOUGLAS COUNTY: Above east bank of Columbia River, Route 97. Rosaceae Elev. 721 ft. 47° 33.6' N, 120° 15.1' W Rubus ursinus Cham. & Schltdl. With Elaeagnus angustifolia, full sun. 3 meters tall arching evergreen shrubs, common, leaves white below, black fruit 18-21 x 19-21 U.S.A., WASHINGTON, SAN JUAN COUNTY: milimeters wide, globose, shiny, stipitate-glandular . San Juan Islands National Wildlife Refuge and Wilderness Area. Phenology: Fruits. Origin: Introduced. Near Decatur Island, Unnamed Island #3. Elev. 6-32 ft. Peter F. Zika 20311 21 Sep 2004 48° 29.097' N, 122° 49.27' W; Source: Specimen label Unforested grassland; steep northwest facing outcrops. Rare. WTU-360952 Phenology: Flowers. Origin: Native. Peter F. Zika 22109 30 May 2005 WTU-383981 Rosaceae Rosaceae Rubus nigerrimus (Greene) Rydb. Rubus arcticus L. U.S.A., WASHINGTON, WHITMAN COUNTY: CANADA, YUKON TERRITORY: [locality and coordinates withheld; status in Washington: G2/S2 Ogilvie Mountains: mountain east of mile 50-54. (Threatened; WANHP)] Elev. 2900 ft. Elev. 1640 ft. 64.595645°, -138.224096°; WGS 84, uncertainty: 5000 m., Dry, north-facing slope below road, with Philadelphus, Rosa woodsii, Source: Digital Map, Georef'd by Joseph Stormer Rhus glabra. Three arching shrubs 1.5 meters tall; canes bright Phenology: Flowers. blue-white, glaucous; leaves green beneath. Phenology: Fruits. R. T. Porsild 91 4 Jul 1966 P. F. Zika 20123 30 Aug 2004 with R. Old WTU-231489 WTU-361032 Rosaceae Rosaceae Rubus arcticus L. Rubus bifrons Vest CANADA, YUKON TERRITORY: Canol Rd.: Mile 95. Upper Rose R. valley. U.S.A., WASHINGTON, KING COUNTY: Elev. 3600 ft. Magnusum Park, north side of Sand Point, Lake Washington, 61.395687°, -133.059111°; WGS 84, uncertainty: 10000 m., Seattle. Source: Digital Map, Georef'd by Sara Legler Elev. 16 ft. Grassy places on river flat. Flowers scented. Phenology: Flowers. T25N R4E S2 NE 1/4; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler A. E. Porsild 10368 11 Jul 1944 Black fruit, bird-sown. Sheet 2 of 2. Phenology: Fruits. with A. J. Breitung A. L. Jacobson s.n. 9 Nov 1999 WTU-127242 WTU-340112 Rosaceae Rosaceae Rubus arcticus L. Rubus ulmifolius Schott U.S.A., ALASKA COUNTY: U.S.A., WASHINGTON, KING COUNTY: Kodiak Island. Olga Bay. Cannery Station, near Akalura Cabin. Magnusum Park, north side of Sand Point, Lake Washington, T35S R31W S9; uncertainty: 5000 m., Georef'd by Joseph Seattle. Stormer Elev. 16 ft. Wet. Flowers pink. Phenology: Flowers. Origin: Native. T25N R4E S2 NE 1/4; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Ethel H. Looff 154 11 Jul 1981 Black fruit, weed near sculptures. Sheet 1 of 2. Phenology: Vegetative. WTU-314309 A. L. Jacobson s.n. 9 Nov 1999 Rosaceae WTU-340114 Rubus arcticus L. Rosaceae U.S.A., ALASKA COUNTY: Central Brooks Range, Walker Lake. Rubus arcticus L. Elev. 2600 ft. 59.395013°, -136.146003°; WGS 84, uncertainty: 1000 m., CANADA, YUKON TERRITORY: Source: Digital Map, Georef'd by Sara Shiina Canol Road: Rose-Lapie R. Pass, mile 105; Schist Mountain east of Wet depression. Phenology: Flowers. Origin: Native. Lake. Elev. 5000 ft. Karen Gustafson s.n. 9 Jul 1979 61.77214°, -133.084269°; uncertainty: 1500 m., Georef'd by Joseph Stormer WTU-282287 Alpine meadow. Phenology: Flowers. A. E. Porsild 10917 21 Jul 1944 Rosaceae with A. J. Breitung Rubus arcticus L. WTU-127244 U.S.A., ALASKA, YUKON KOYUKUK COUNTY: Camp Denali; Moose Creek Area. Elev. 1750 ft. 63° 31' 6.6" N, 150° 53' 36.6" W Boggy bottom land. Phenology: Flowers. Origin: Native. Sally Dickman s.n. 23 Jun 1967 WTU-287245 Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. CANADA, BRITISH COLUMBIA: U.S.A., ALASKA COUNTY: Haines Road (kilometers 91), Three Guardsmen Mountain. Along spit bordering small lake near mouth of Sadie Creek, Elev. 5002 ft. Kotzebue Sound. 59° 35' N, 136° 29' W 66.444246°, -162.83461°; uncertainty: 85295 m., Georef'd by On mineral soil in alpine. Phenology: Flowers. Joseph Stormer On damp banks. Common; mostly in flower now, with only few in G. W. Douglas 11059 27 Jun 1979 fruit. Phenology: Flowers. Origin: Native. with M. J. Ratcliffe Jean H. Langenheim 4296 15 Aug 1956 WTU-282004 WTU-177139 Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA, SITKA COUNTY: Sitka. U.S.A., ALASKA COUNTY: 56.966317°, -135.207777°; WGS 84, uncertainty: 106622 m., Hinchinbrook Id. Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Phenology: Fruits. Origin: Native. W. J. Eyerdam s.n. 6 Jun 1937 A. A. Heller 14913 11 Jul 1929 WTU-292195 WTU-89109 Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA COUNTY: CANADA, BRITISH COLUMBIA: Below Fishhook Inn, Lit. Susitna valley. One mile south of south end of Summit Lake along Hart Highway Roadside. Type locality. Phenology: Fertile. north of Prince George. 54.312184°, -122.664576°; WGS 84, uncertainty: 2000 m., J. P. Anderson 6981 27 Jun 1941 Source: Digital Map, Georef'd by Sara Shiina Common at margin of bog among alders and willows; occasional in WTU-292745 open part of bog. Flowers pink. Phenology: Flowers. J. A. Calder 12490 4 Jul 1954 Rosaceae with D. B. O. Savile, J. M. Ferguson Rubus arcticus L. WTU-194977 CANADA, YUKON TERRITORY: Northeast shoulder of Mount Sheldon; saddle between Mount Rosaceae Sheldon and Mount Riddle. Elev. 4500 ft. Rubus arcticus L. × Rubus arcticus L. var. acaulis (Michx.) Timbered lower slopes; open avalanche paths near timberline. Boivin Phenology: Flowers. U.S.A., ALASKA COUNTY: A. E. Porsild 11083 13 Aug 1944 Flat north of Savage River; McKinley National Park. with A. J. Breitung Elev. 2952 ft. 63.766705°, -149.294778°; uncertainty: 20000 m., Georef'd by WTU-254242 Joseph Stormer In dwarf birch thicket. Flowers cerise; common. Phenology: Flowers. Rosaceae Ynes Mexia 2088 9 Jul 1928 Rubus arcticus L. WTU-30035 CANADA, YUKON TERRITORY: Northeast shoulder of Mount Sheldon and glacial cirque. Rosaceae Elev. 4500 ft. Forming dense undergrowth in alder thicket near timberline. Rubus arcticus L. Phenology: Vegetative. U.S.A., ALASKA, KENAI PENINSULA COUNTY: A. E. Porsild 11082 13 Aug 1944 Skilak Lake. with A. J. Breitung 60° 28' N, 150° 28' W Among rocks on upper part of beach. Quite common; flowers pink. WTU-254237 Phenology: Flowers. J. A. Calder 4986 31 May 1951 WTU-194985 Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. CANADA, YUKON TERRITORY: U.S.A., ALASKA, NOME COUNTY: Northeast shoulder of Mount Sheldon and glacial cirque. Seward Peninsula, Cape Nome. Elev. 4500 ft. 64.473659°, -164.997849°; WGS 84, uncertainty: 10000 m., Open avalanch paths near timberline. Phenology: Flowers. Source: Digital Map, Georef'd by Joseph Stormer Rocky slope. Phenology: Flowers. Origin: Native. A. E. Porsild 11083 13 Aug 1944 with A. J. Breitung George Neville Jones 9121 Jul 1936 WTU-254233 WTU-36124

Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. CANADA, YUKON TERRITORY: CANADA, BRITISH COLUMBIA: Canol Road. MacMillian Pass. Mackenzie-Yukon divide, mile Blackwater Lake, about 32 miles north of Golden. 284-290. Elev. 2150 ft. Elev. 4600-5500 ft. 51.623437°, -117.406929°; uncertainty: 1000 m. In a river meadow where it appeared to be sterile. Phenology: On Sphagnum island. Phenology: Flowers. Vegetative. C. L. Hitchcock 7651 11 Jul 1941 A. E. Porsild 11243 31 Aug 1944 with J. S. Martin WTU-254232 WTU-68305 Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. CANADA, YUKON TERRITORY: Northeast shoulder of Mount Sheldon; saddle between Mount U.S.A., ALASKA COUNTY: Sheldon and Mount Riddle. St. Paul, Pribilof Islands. Elev. 4500 ft. 57.17308°, -170.300654°; WGS 84, uncertainty: 7500 m., Source: Open avalanche paths near timberline. Phenology: Flowers. Digital Map, Georef'd by Sara Shiina Phenology: Flowers. Origin: Native. A. E. Porsild 11083 13 Aug 1944 with A. J. Breitung T. Kincaid s.n. Aug 1897 WTU-127241 WTU-101573

Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA COUNTY: CANADA, BRITISH COLUMBIA: Approximately 4 miles northwest of Cooper Center. Mountains 10-15 miles south of Telegraph Creek. Roadside. Phenology: Flowers. In lower alpine meadows under shadow of small semi-scrub Abies. Flowers crimson-lake. Phenology: Flowers. R. John Taylor 3048 5 Jul 1965 T. T. McCabe 8879 15 Aug 1941 WTU-224866 WTU-71768 Rosaceae Rosaceae Rubus arcticus L. Rubus bartonianus M. Peck U.S.A., ALASKA COUNTY: St. Paul Island, Pribilof Group. U.S.A., IDAHO, IDAHO COUNTY: 57.178849°, -170.273364°; uncertainty: 13000 m. [locality and coordinates withheld; status in Idaho: S2 (Imperiled; Phenology: Flowers. Origin: Native. IDNHP)] Rock slide by trail, with Philadephus lewisii. Phenology: Fruits. M. A. Baranoff 26 15 Jul 1938 Douglass Henderson 3041 23 May 1976 WTU-47206 WTU-358365 Rosaceae Rosaceae Rubus arcticus L. Rubus bartonianus M. Peck U.S.A., ALASKA, NOME COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Seward Peninsula: Nome; along Dry Creek. [locality and coordinates withheld; status in Idaho: S2 (Imperiled; 64.522564°, -165.351569°; WGS 84, uncertainty: 4149 m., IDNHP)] Source: Digital Map, Georef'd by Joseph Stormer Deep protect-ravines along small stream. Shrub, 4 to 8 feet, tall. Tundra. Phenology: Fertile. Origin: Native. Phenology: Flowers. George Neville Jones 9122 Jul 1936 Fred G. Meyer 846 13 May 1937 WTU-12024JWT WTU-16886JWT

Rosaceae Rosaceae Rubus bartonianus M. Peck Rubus arcticus L. U.S.A., IDAHO, LEMHI COUNTY: CANADA, BRITISH COLUMBIA: [locality and coordinates withheld; status in Idaho: S2 (Imperiled; Hart Highway, Summit Lake. IDNHP)] Drained peat bog. Phenology: Flowers. Outlying ridge at timberline. Phenology: Fruits. Origin: Native. T. M. C. Taylor 12 7 Jun 1956 J. H. Christ 14850 15 Jul 1945 with A. F. Szczawinski, M. Bell WTU-265969 WTU-172424

Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA, NOME COUNTY: CANADA, BRITISH COLUMBIA: Seward Peninsula: Nome; along Dry Creek. Haines Road, mile 75. 64.522564°, -165.351569°; WGS 84, uncertainty: 4149 m., Elev. 3116 ft. Source: Digital Map, Georef'd by Joseph Stormer 59° 30' N, 136° 30' W Tundra. Phenology: Flowers. Origin: Native. Damp creek bank. Phenology: Flowers. George Neville Jones 9122 Jul 1936 T. M. C. Taylor 745 5 Jul 1956 WTU-36125 WTU-172425

Rosaceae Rosaceae Rubus idaeus L. Rubus bartonianus M. Peck U.S.A., IDAHO, CAMAS COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Along Forest Service road 094; along Little Smoky Creek. [locality and coordinates withheld; status in Idaho: S2 (Imperiled; Elev. 7000 ft. IDNHP)] 44° 33' N, 114° 48.5' W; WGS 84, uncertainty: 1500 m., Source: Brushy thickets. Slender shrub, 6 to 8 feet high. Phenology: Flowers. Nat. Geog. TOPO!, Georef'd by John Bassett Talus slope above road, by stream. Corollas white. Phenology: L. Constance 1818 13 May 1937 Fertile. Origin: Native. with D. Hedrick, D. Peters J. F. Smith 4250 22 Jun 2002 WTU-13431JWT WTU-352744 Rosaceae Rosaceae Rubus idaeus L. Rubus bartonianus M. Peck U.S.A., WASHINGTON, PIERCE COUNTY: Longmire. Mount Rainier. U.S.A., IDAHO, IDAHO COUNTY: 46.75°, -121.81139°; WGS 84, uncertainty: 10000 m., Source: [locality and coordinates withheld; status in Idaho: S2 (Imperiled; GeoLocate, Georef'd by WTU Staff IDNHP)] Moist soil, edge of swamp. Phenology: Fertile. Brushy thickets. Slender shrub, 6 to 8 feet high. Phenology: Flowers. F. A. Warren 1672 31 Aug 1932 L. Constance 1818 13 May 1937 with D. Hedrick, D. Peters WTU-36698 WTU-41447 Rosaceae Rosaceae Rubus arcticus L. Rubus idaeus L. U.S.A., ALASKA, NOME COUNTY: U.S.A., IDAHO COUNTY: Seward Peninsula, Cape Nome. 1 1/2 miles west C. T. P. A. Headquarters. 64.473659°, -164.997849°; WGS 84, uncertainty: 10000 m., Ridge top. Phenology: Fertile. Origin: Native. Source: Digital Map, Georef'd by Joseph Stormer Rocky slope. Phenology: Flowers. Origin: Native. Gene F. Payne G-59 28 Jun 1942 George Neville Jones 9121 Jul 1936 WTU-296025 WTU-12023JWT Rosaceae Rosaceae Rubus chamaemorus L. Rubus arcticus L. CANADA, YUKON TERRITORY: Canol Road: Mile 132. Lower Lapie River Crossing. CANADA, BRITISH COLUMBIA: Elev. 2750 ft. Vicinity of Upper Hat Creek, base of Marble Mountains. 61.90854°, -132.594276°; uncertainty: 5000 m., Georef'd by Elev. 3000 ft. Joseph Stormer 50.817167°, -121.574099°; uncertainty: 8000 m. In a muskeg. Male plant only. Phenology: Flowers. Phenology: Fertile. A. E. Porsild 9582 17 Jun 1944 J. William Thompson 367 12 Jul 1938 with A. J. Breitung with Emily M. Thompson WTU-128621 WTU-17384JWT Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. CANADA, YUKON TERRITORY: U.S.A., ALASKA COUNTY: Canol Road: Mile 132, Lower Lapie River Crossing. North slope Borough, Atkasook, Meade River, 60 miles south of Elev. 2750 ft. Barrow. 61.90854°, -132.594276°; uncertainty: 5000 m., Georef'd by 70.483962°, -157.403845°; WGS 84, uncertainty: 10000 m., Joseph Stormer Source: Digital Map, Georef'd by Joseph Stormer In a muskeg. Phenology: Flowers. Phenology: Flowers. Origin: Native. A. E. Porsild 9960 27 Jun 1944 Beatrice R. Vogel s.n. 1 Jul 1978 with A. J. Breitung WTU-336549 WTU-128620

Rosaceae Rosaceae Rubus chamaemorus L. Rubus arcticus L. CANADA, YUKON TERRITORY: U.S.A., ALASKA COUNTY: Ogilvie Mountains; lake west of Dempster Road near mile 58. Flat north of Savage River; McKinley National Park. Elev. 3800 ft. Elev. 2952 ft. 64.650381°, -138.394384°; WGS 84, uncertainty: 2000 m., 63.766705°, -149.294778°; uncertainty: 20000 m., Georef'd by Source: Digital Map, Georef'd by Joseph Stormer Joseph Stormer In willows around outlet from lake. Phenology: Flowers. In dwarf birch thicket. Phenology: Flowers. Origin: Native. R. T. Porsild 334 26 Jul 1966 Ynes Mexia 2088 9 Jul 1928 WTU-231488 WTU-86074

Rosaceae Rosaceae Rubus arcticus L. Rubus chamaemorus L. CANADA, BRITISH COLUMBIA: U.S.A., ALASKA COUNTY: 25 miles north of Fort St. James. Adjacent Canada, Attu Island, on Terrible Mountain. Willow bog, growing in sphagnum in shade of dense willows. Petals 52.855598°, 173.154721°; WGS 84, uncertainty: 1000 m., Source: strong uniform pink. Phenology: Flowers. Digital Map, Georef'd by Sara Shiina Hillside, in deep moss and low grass. Abundant. Phenology: T. T. McCabe 7535 25 Jun 1940 Flowers. Origin: Native. WTU-72186 Maxcine Williams 2789 12 Jul 1971 WTU-255177 Rosaceae Rosaceae Rubus arcticus L. Rubus chamaemorus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA, WRANGELL PETERSBURG COUNTY: Fairbanks. Pillar Bay, Kuiu Island. 64.837778°, -147.716389°; WGS 84, uncertainty: 10000 m., 56.629169°, -134.173422°; WGS 84, uncertainty: 6643 m., Source: GeoLocate, Georef'd by Joseph Stormer Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Upstream bogs. Phenology: Fertile. Origin: Native. Christine Heller s.n. 16 Jun 1946 G. B. Rigg 20 11 Aug 1929 WTU-110974 WTU-36222

Rosaceae Rosaceae Rubus chamaemorus L. Rubus idaeus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Camp Denali, Taiga Trail. Tongas Village. Elev. 1750-3500 ft. Site of abandoned native village. Phenology: Fruits. Origin: Native. 63° 31' 6.6" N, 150° 53' 36.6" W Phenology: Flowers. Origin: Native. Mr. Ernest P. Walker 890 3 Aug 1915 with Mrs. Ernest P. Walker Sally Dickman s.n. 25 Jun 1967 WTU-76668 WTU-313813 Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. U.S.A., ALASKA, WRANGELL PETERSBURG COUNTY: U.S.A., ALASKA COUNTY: Pillar Bay, Kuiu Island. Hydaburg. 56.629169°, -134.173422°; WGS 84, uncertainty: 6643 m., Elev. 0 ft. Source: Digital Map, Georef'd by Joseph Stormer 55.212366°, -132.806068°; WGS 84, uncertainty: 3500 m., Phenology: Fruits. Origin: Native. Source: Digital Map, Georef'd by Joseph Stormer Muskeg. Phenology: Fruits. Origin: Native. G. B. Rigg 30 11 Aug 1929 Helen H. Norton 642-H 3 Jul 1979 WTU-36223 WTU-281090 Rosaceae Rosaceae Rubus chamaemorus L. Rubus idaeus L. U.S.A., ALASKA COUNTY: Near Miller House. U.S.A., IDAHO COUNTY: 65.524167°, -145.216111°; WGS 84, uncertainty: 10000 m., Cedar Mountain. Source: GeoLocate, Georef'd by Joseph Stormer 47.350204°, -115.842379°; WGS 84, uncertainty: 1500 m., Bank of a small stream. Phenology: Fruits. Origin: Native. Source: GeoLocate, Georef'd by Sara Shiina Phenology: Fertile. Origin: Native. R. John Taylor 3194 31 Jul 1965 C. V. Piper 2879 Jul 1898 WTU-226534 WTU-20701 Rosaceae Rosaceae Rubus idaeus L. ssp. idaeus Rubus chamaemorus L. U.S.A., OREGON, KLAMATH COUNTY: Aspen Mountain, Cascade Mountains. U.S.A., ALASKA COUNTY: East slope, dry yellow pine forest. Phenology: Fertile. 54.912677°, -158.050781°; WGS 84, uncertainty: 1879152 m., Elmer I. Applegate 3753 30 Jul 1923 Source: Digital Map, Georef'd by Joseph Stormer Phenology: Fertile. Origin: Native. WTU-8831JWT G. B. Rigg 70 6 Aug 1929 WTU-39228 Rosaceae Rosaceae Rubus chamaemorus L. Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Along East side of Oarlock Island, Dease Inlet. Coopers Landing. 71° N, 156° W; Misleading coordinates given only to nearest Phenology: Fertile. Origin: Native. degree. Sides and between high-center polygon bummocks, almost covering J. P. Anderson 6862 19 Jun 1941 area, tundra. Phenology: Flowers & Fruits. Origin: Native. WTU-292748 Ira L. Wiggins 12775 25 Aug 1951 WTU-191946 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., WASHINGTON, PIERCE COUNTY: Mount Tacoma. U.S.A., WASHINGTON COUNTY: Elev. 4000 ft. Klickitat River basin. Indefinite loan by Tacoma High School to University of WAshington. Dsense mossy woods. Indefinite loan by Tacoma High School to Phenology: Flowers. Origin: Native. University of WAshington. Phenology: Fertile. Origin: Native. J. B. Flett s.n. 28 Aug 1895 J. B. Flett 1336 13 Jul 1899 WTU-33524 WTU-33510 Rosaceae Rosaceae Rubus chamaemorus L. Rubus idaeus L. U.S.A., ALASKA COUNTY: CANADA, BRITISH COLUMBIA: Umiat. Haines Road (km 68), near Alaska border. 69° 22' N, 152° 8' W Elev. 918 ft. Tundra of both hills and river bottoms. Common. 59° 28' N, 136° 23' W Reference specimen for illustration in Revision In Populus trichocarpa stand. Phenology: Fertile. of Flora of the Pacific Northwest, WTU Herbarium, 2016. Phenology: Flowers. Origin: G. W. Douglas 11029 26 Jun 1979 Native. with M. J. Ratcliffe George Lindsay 2275 WTU-281992 7 Jul 1952 Rosaceae WTU-193643, WTU-V-024779 Rubus chamaemorus L. Rosaceae U.S.A., ALASKA COUNTY: Point Barrow, east shore of Sungorak Lake, 18 miles south of Rubus chamaemorus L. Barrow Base. 71° 5' N, 156° 26' 30" W U.S.A., ALASKA COUNTY: Mossy sides of polygons. Staminate and pistillate plants. Phenology: Umiat. Flowers & Fruits. Origin: Native. 69° 22' N, 152° 8' W In tundra. Common. Phenology: Fruits. Origin: Native. Henry J. Thompson 1420 29 Aug 1950 George Lindsay 2362 12 Aug 1952 WTU-193642 WTU-193644 Rosaceae Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rubus idaeus L. U.S.A., IDAHO, BONNER COUNTY: Along Pend Oreille River, four miles east of Priest River. U.S.A., IDAHO, BONNER COUNTY: 48.200171°, -116.90305°; WGS 84, uncertainty: 10000 m., Packrat Gorge, 15 miles northwest of Nordman. Source: GeoLocate, Georef'd by WTU Staff 48.81359°, -117.00264°; WGS 84, uncertainty: 6750 m., Source: Open pine woods. Erect shrub. Phenology: Fertile. Origin: Native. Nat. Geog. TOPO!, Georef'd by John Bassett Granite walls. Phenology: Fertile. Origin: Native. William H. Baker 8968 31 May 1952 William H. Baker 15176 22 Jun 1958 WTU-295928 WTU-295712 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., WASHINGTON, MASON COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Mount Elinor [Ellinor]. Mount Dickerman, Cascade Mountains. Elev. 5500 ft. Elev. 5000 ft. 47.52205°, -123.26061°; WGS 84, uncertainty: 500 m., Source: 48.06829°, -121.47068°; WGS 84, uncertainty: 3000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff In thicket. Phenology: Flowers. Origin: Native. Moist banks. Phenology: Flowers. Origin: Native. Phillips B. Freer 375 3 Sep 1950 J. William Thompson 8870 17 Jul 1932 WTU-135201 WTU-23011JWT

Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: 1 mile inland [circa] 5 miles southwest of Barrow Base. Point Barrow, north shore of Iko Bay, 18 miles southeast of Barrow 71° 15' N, 156° 51' W Base. Sides of high-center polygons. Phenology: Flowers. Origin: Native. 71° 10.5' N, 156° 5' W Peaty hummocks with Vaccinium vitis-idaea. Plants apparently Geo. H. Ward 1102 13 Jul 1952 spreading vegetatively, leaves forming nearly complete ground with I. L. Wiggins cover, many of the foliaceous. Phenology: Flowers. Origin: Native. WTU-193645 John H. Thomas 2208 16 Aug 1950 Rosaceae WTU-193647 Rubus lasiococcus A. Gray Rosaceae U.S.A., WASHINGTON, YAKIMA COUNTY: Rubus lasiococcus A. Gray Snoqualmie National Forest, Shoe Lake. Elev. 6100 ft. U.S.A., WASHINGTON COUNTY: T13N R11E S25 NE 1/4; WGS 84, uncertainty: 10000 m., Source: Stevens Pass Region, Cascade Mountains. GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Flat, shallow sand and gravel in open grass park. Heavy useage. In Carex type. Distribution: high altitude; common. Phenology: Fertile. J. M. Grant s.n. Aug 1929 Origin: Native. WTU-43174 E. H. Reid 49 22 Aug 1936 WTU-322443 Rosaceae Rubus lasiococcus A. Gray Rosaceae U.S.A., WASHINGTON COUNTY: Rubus chamaemorus L. Along trail to Snow Lake, Cascade Mountains. Elev. 4000 ft. U.S.A., ALASKA COUNTY: Alpine woods. Phenology: Flowers. North side of Meade River, circa. 50 miles south of Barrow. J. William Thompson 10652 16 Jun 1934 70° 40' N, 156° 55' W In wet moss near edge of pond. Reference WTU-25414 specimen for illustration in Revision of Flora of the Pacific Northwest, WTU Herbarium, 2016. Phenology: Flowers. Origin: Native. Rosaceae George H. Ward 1198 Rubus lasiococcus A. Gray 18 Jul 1952 U.S.A., WASHINGTON, PIERCE COUNTY: Mount Rainier. WTU-193646, WTU-V-024778 46.852827°, -121.760441°; WGS 84, uncertainty: 17500 m., Source: GeoLocate, Georef'd by Joseph Stormer Phenology: Fruits. Origin: Native. C. V. Piper s.n. Aug 1888 WTU-9558 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Scenic. Western base of Iron Peak, upper North Fork Teanaway Creek, Elev. 2100 ft. Wenatchee Mountains. 47.71167°, -121.15444°; WGS 84, uncertainty: 10000 m., Source: 47.411667°, -120.948889°; NAD 27, uncertainty: 3000 m., Source: GeoLocate, Georef'd by WTU Staff Georeferenced, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. In woods. Phenology: Flowers. Origin: Native. I. C. Otis 925 16 Jun 1920 A. R. Kruckeberg 2584 7 Aug 1950 WTU-15876JWT WTU-182438

Rosaceae Rosaceae Rubus chamaemorus L. Rubus lasiococcus A. Gray U.S.A., ALASKA COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: 1 mile southwest of Barrow Base, inland from coast a few hundred Mountain Rainier. yards. 46.852133°, -121.761399°; WGS 84, uncertainty: 17500 m., 71° 19' N, 156° 43' W Source: Digital Map, Georef'd by Joseph Stormer On tops and sides of hummocks, appearing to spread by vegetative Phenology: Flowers. Origin: Native. means only. Petals with 1 main vein from base to apex, others derived from this, and some bifurcate at distal ends. Phenology: A. S. Pope s.n. 6 Aug 1904 Flowers. Origin: Native. WTU-9534 John H. Thomas 2161 8 Aug 1950 WTU-193648 Rosaceae Rubus chamaemorus L. Rosaceae U.S.A., ALASKA COUNTY: Rubus lasiococcus A. Gray Just west of base at Umiat: flats adjacent to Colville River, north of river. U.S.A., WASHINGTON, CLALLAM COUNTY: Elev. 915 ft. Mount Angeles. 69° 23' N, 152° 10' W 48.00365°, -123.445867°; WGS 84, uncertainty: 5330 m., Source: Flat damp area. In fruit but not yet ripe, fruit reddish and yellow, 2 to Digital Map, Georef'd by Joseph Stormer 2.5 cemtimeters. Phenology: Fruits. Origin: Native. Phenology: Flowers. Origin: Native. Henry J. Thompson 1212 25 Jul 1950 E. B. Webster 1633 Aug 1911 WTU-193640 WTU-32286 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., WASHINGTON, CLALLAM COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Mount Angeles, Olympic Mountains. Ice Creek. Elev. 3000 ft. Elev. 4000 ft. 47.99536°, -123.46725°; WGS 84, uncertainty: 1000 m., Source: 48.105535°, -120.737582°; WGS84, uncertainty: 10000 m., GeoLocate, Georef'd by WTU Staff Source: GeoLocate, Georef'd by WTU Staff Lower wooded slopes. Phenology: Flowers. Origin: Native. Woods. Phenology: Fruits. Origin: Native. J. William Thompson 7572 20 Jul 1931 G. E. Morrill 407 23 Aug 1933 WTU-23009JWT WTU-27884 Rosaceae Rosaceae Rubus idaeus L. Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., OREGON, WALLOWA COUNTY: U.S.A., MONTANA, DEER LODGE COUNTY: Imnaha Grade. Anaconda-Pintlar Range, north ridge of West Goat Peak. Dry bank. Phenology: Fruits. Origin: Native. Elev. 8900 ft. Common in coarse talus and around rockpiles. Phenology: Flowers. Lilla Leach 4252 19 Jul 1931 Origin: Native. WTU-255247 Klaus H. Lackschewitz 5681 23 Aug 1974 WTU-267275 Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Overlooking west bank of chandler River circa. 5 Aleutians, Attu Island, northeast Alexi Pass. miles south of its mouth and circa. 4 miles south Elev. 300 ft. of Gubic Camp. 52.852564°, 173.318684°; WGS 84, uncertainty: 1500 m., Source: 69° 23' N, 151° 25' W Digital Map, Georef'd by Joseph Stormer Under Alders, hillsides. Reference specimen for Crowberry slope. Phenology: Fruits. Origin: Native. illustration in Revision of Flora of the Pacific Northwest, WTU Herbarium, 2016. Phenology: Robert M. Hardy s.n. 3 Sep 1945 Fruits. Origin: Native. WTU-106630 Kenton L. Chambers 227 24 Jul 1951 Rosaceae WTU-193641, WTU-V-024776 Rubus chamaemorus L. U.S.A., ALASKA, NORTH SLOPE COUNTY: Northeast Alaska; Okpilak Valley, 1 kilometer west of West Okpilak Rosaceae Lake. Elev. 1902 ft. Rubus lasiococcus A. Gray 69° N, 144° W; Misleading coordinates given only to nearest degree. U.S.A., WASHINGTON, PIERCE COUNTY: Upland tundra, wet shrub tundra. Phenology: Flowers. Origin: Mt. Rainier, Paradise Valley. Native. 46.782778°, -121.728611°; NAD 27, uncertainty: 1000 m., Source: Georeferenced, Georef'd by WTU Staff, Invalid coordinates N. E. Grulke 98 Aug 1984 (located outside county bounds). with D. J. Marrett Near snow fields. Phenology: Flowers. Origin: Native. WTU-305170 A. S. Foster Foster No. 1136 8 Aug 1909 Rosaceae WTU-9784 Rubus idaeus L. U.S.A., IDAHO, BONNER COUNTY: Rosaceae Priest River Experimental Forest. 48.355678°, -116.787586°; WGS 84, uncertainty: 4000 m., Rubus lasiococcus A. Gray Source: Digital Map, Georef'd by Melissa Bowers Some leaves pinnate. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON COUNTY: Mount Adams. R. F. Daubenmire 43366 21 Jul 1943 46.197222°, -121.491389°; WGS 84, uncertainty: 10000 m., Source: Digital map, Georef'd by Ben Legler WTU-251476 Phenology: Fertile. Origin: Native. Wilhelm N. Suksdorf 11925 5 Sep 1925 Rosaceae WTU-197551 Rubus lasiococcus A. Gray U.S.A., OREGON, WASCO COUNTY: Rosaceae Near Fifteen Mile Meadow, Mount Hood National Forest. Elev. 4500 ft. Rubus lasiococcus A. Gray Damp woods. Phenology: Fertile. Origin: Native. U.S.A., WASHINGTON, KITTITAS COUNTY: G. N. Jones 4119 19 Jul 1933 Just west of Hawkins Mountain on Huckleberry Peak trail, Upper Cle Elum River drainage, Wenatchee National Forest. WTU-28735 Elev. 4500 ft. 47.176973°, -120.991239°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Rosaceae Open meadows in subalpine fir-mountain hemlock forest type. Phenology: Flowers. Origin: Native. Rubus lasiococcus A. Gray A. R. Kruckeberg 5653 13 Jul 1963 U.S.A., OREGON, HOOD RIVER COUNTY: Lost Lake. WTU-214204 45.489349°, -121.821695°; uncertainty: 3000 m. Open woods. Phenology: Flowers. Origin: Native. Sister Mary Milburge 1466 23 Jun 1938 WTU-17472JWT Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus lasiococcus A. Gray U.S.A., ALASKA COUNTY: U.S.A., OREGON COUNTY: Klondike River. Mount Hood. Woods. Indefinite loan by Tacoma High School to University of Elev. 4000 ft. Washington. Phenology: Flowers. 45.373343°, -121.695728°; WGS 84, uncertainty: 2000 m., Source: GeoLocate, Georef'd by Sara Shiina J. B. Flett 1336 13 Jul 1899 In woods. Phenology: Flowers. Origin: Native. WTU-36221 T. C. Frye s.n. 30 Aug 1907 WTU-9865 Rosaceae Rubus lasiococcus A. Gray Rosaceae U.S.A., OREGON, CLACKAMAS COUNTY: Rubus chamaemorus L. Around Mirror Lake, Mount Hood National Forest. Elev. 4000 ft. U.S.A., ALASKA COUNTY: 45.297067°, -121.793085°; uncertainty: 1000 m. Arctic Slope District: 70 air miles south of Point Barrow, near eskimo Alpine slopes. Phenology: Flowers. Origin: Native. village of Atkasook, 2.5 kilometers south of Meade River Camp. Elev. 49 ft. J. William Thompson 1642 10 Aug 1926 70.481641°, -157.416014°; uncertainty: 10000 m., Georef'd by Joseph Stormer WTU-23004JWT-A Tussock tundra. Phenology: Flowers. Origin: Native. Vera Komarkova 325 14 Jul 1976 Rosaceae with Holly Hansell, Karen Seabert Rubus chamaemorus L. WTU-271686 U.S.A., ALASKA COUNTY: Baranof Island, west coast. About 2 miles northeast of Sitka near the Rosaceae Indian River Trail. Elev. 50 ft. Rubus lasiococcus A. Gray 57.077223°, -135.289851°; WGS 84, uncertainty: 2000 m., Source: Digital Map, Georef'd by Joseph Stormer U.S.A., OREGON, HOOD RIVER COUNTY: Large muskeg, near trail. Near margin of muskeg, along musked Lost Lake. edges. Flower white, common. Phenology: Fruits. Origin: Native. Elev. 3140 ft. 45.489349°, -121.821695°; WGS 84, uncertainty: 2000 m., Mary Clay Muller 4277 30 Jun 1981 Source: GeoLocate, Georef'd by Joseph Stormer Open woods. Phenology: Flowers. Origin: Native. WTU-289027 Sister Mary Milburge 1466 23 Jun 1938 Rosaceae WTU-53659 Rubus idaeus L. ssp. strigosus (Michx.) Focke Rosaceae CANADA, YUKON TERRITORY: South slope of Moosehide Mountain, Dawson. Rubus chamaemorus L. Along open trail in Aspen woods. Flowers white, height 1 foot - 3 1/2 - canes upright. Phenology: Flowers. U.S.A., ALASKA, ALEUTIANS WEST COUNTY: Pribilof Group. Saint Paul Island. J. A. Calder 3509 4 Jul 1949 57.180474°, -170.250996°; uncertainty: 13000 m. with L. G. Billard Phenology: Flowers. Origin: Native. WTU-291126 M. A. Baranoff 53 23 Jul 1938 WTU-47338 Rosaceae Rubus lasiococcus A. Gray Rosaceae U.S.A., OREGON, LANE COUNTY: Rubus chamaemorus L. McKenzie Pass, Cascade Mountains. Elev. 4000 ft. U.S.A., ALASKA COUNTY: Alpine slopes. Phenology: Flowers. Origin: Native. Aleutian Islands, Attu Island. 52.9°, 172.932959°; WGS 84, uncertainty: 35000 m., Source: J. William Thompson 48 11 Aug 1925 Digital Map, Georef'd by Joseph Stormer At low elevations. In quantity, but in 1944 blooms very few, in one or WTU-230004JWT-B two restricted areas fruit plentiful. Phenology: Fruits. Origin: Native. G. B. Van Schaack 36b Aug 1944 WTU-155473 Rosaceae Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rubus lasiococcus A. Gray CANADA, BRITISH COLUMBIA: U.S.A., OREGON, JOSEPHINE COUNTY: 21 Mile Creek, near Rainbow Falls, Alta Lake. Lake Mountain, Oregon Caves National Monument. Phenology: Flowers. 42.090139°, -123.359861°; WGS 84, uncertainty: 2000 m., Source: Digital map, Georef'd by Joseph Stormer F. Comte 5487 31 Jul 1967 Edge of snow. Phenology: Flowers. Origin: Native. WTU-329403 William H. Baker 330 3 Jul 1949 with George C. Ruhle Rosaceae WTU-274604 Rubus lasiococcus A. Gray Rosaceae U.S.A., OREGON, KLAMATH COUNTY: Crater Lake National Park. Rubus chamaemorus L. Along the rim of the lake in sandy soil. Hesperopeuce Mertensiana occurs almost exclusively, there being a few trees of PInus albicaulis U.S.A., ALASKA COUNTY: on the crest of the rim. There are practically no undershrubs except Sitka. infrequent clumps of elder and prostrate mates of Ribes 57.094858°, -135.272322°; WGS 84, uncertainty: 10000 m., erythrocarpum. Phenology: Fertile. Origin: Native. Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. A. A. Heller 12576 29 Aug 1916 T. Kincaid s.n. 16 Jun 1899 WTU-83663 WTU-101660 Rosaceae Rosaceae Rubus chamaemorus L. Rubus lasiococcus A. Gray U.S.A., ALASKA COUNTY: Point Lay. Along north bank of Kukpowruk River between 1 and 2 U.S.A., OREGON, JOSEPHINE COUNTY: miles from its mouth on Kasegaluk Lagoon. Forest Service, Camp Grayback, Upper Biglow Lake, Oregon Caves 69° 36' N, 163° W National Monument & Vicinity. On tundra, berries ripe. Common locally, 15 to 20 milimeters in Woods. Phenology: Fertile. Origin: Native. diameter, salmon colored. Phenology: Fruits. Origin: Native. William H. Baker 309 3 Jul 1949 John H. Thomas 2256 2 Sep 1950 with George C. Ruhle WTU-152777 WTU-274596

Rosaceae Rosaceae Rubus idaeus L. Rubus chamaemorus L. U.S.A., IDAHO, KOOTENAI COUNTY: U.S.A., ALASKA COUNTY: Southern shore of Spirit Lake. Aleutians, Attu Island, Cape Khlebnikof. 47.94111°, -116.89389°; WGS 84, uncertainty: 10000 m., Source: Elev. 100 ft. GeoLocate, Georef'd by WTU Staff 52.919193°, 173.287185°; WGS 84, uncertainty: 2000 m., Source: Grassy slope. Phenology: Flowers. Origin: Native. Digital Map, Georef'd by Joseph Stormer In deep crowberry and moss. Phenology: Flowers. Origin: Native. William H. Baker 13426 13 Jul 1955 Robert M. Hardy 236 20 Jul 1945 WTU-295675 WTU-97420 Rosaceae Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rubus chamaemorus L. U.S.A., WASHINGTON, CHELAN COUNTY: Near Anthem Creek, Entiat Valley. U.S.A., ALASKA COUNTY: Elev. 3200 ft. Nome City. 48.044756°, -120.692953°; WGS84, uncertainty: 5000 m., Source: 64.501111°, -165.406389°; WGS 84, uncertainty: 1500 m., GeoLocate, Georef'd by WTU Staff Source: GeoLocate, Georef'd by Sara Shiina Phenology: Fertile. Along seashore tundra. Common. Phenology: Flowers & Fruits. Origin: Native. G. E. Morrill 408 23 Aug 1933 J. B. Flett 1616 20 Jul 1900 WTU-27754 WTU-101574 Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA, SITKA COUNTY: Saint George, Pribilof Island. Back of Sitka. 56.602126°, -169.545272°; WGS 84, uncertainty: 1000 m., 57.117915°, -135.23906°; WGS 84, uncertainty: 10000 m., Source: Digital Map, Georef'd by Sara Shiina Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Small tundra on the plateau. Phenology: Fruits. Origin: Native. T. Kincaid s.n. 25 Jul 1897 A. A. Heller 14913 11 Jul 1929 WTU-101572 WTU-88783

Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. U.S.A., ALASKA, DILLINGHAM COUNTY: CANADA, BRITISH COLUMBIA: Dillingham. 17 miles north of Fort Saint James. 59.053227°, -158.49665°; WGS 84, uncertainty: 4500 m., Source: Deep sphagnum under dwarfed spruce. Petals white, very abundant. Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Flat. Phenology: Flowers. Origin: Native. T. T. McCabe 7571 25 Jun 1940 D. Martel 107 9 Jun 1935 WTU-71683 WTU-101657 Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. CANADA, BRITISH COLUMBIA: U.S.A., ALASKA COUNTY: Along highway between Prince Rupert and Galloway Rapids. Mainland, Port Houghton. 54.292243°, -130.271636°; WGS 84, uncertainty: 3990 m., 57.305727°, -133.25112°; WGS 84, uncertainty: 15755 m., Source: Digital Map, Georef'd by Joseph Stormer Source: Digital Map, Georef'd by Joseph Stormer In sphagnumbog, at edge of coniferous woods. Flowers white, Grassy edge of woods. Phenology: Fruits. Origin: Native. common. Phenology: Fruits. Mr. Ernest P. Walker 862 27 Jul 1915 J. A. Calder 13202 18 Jul 1954 with Mrs. Ernest P. Walker with D. B. O. Savile, J. M. Ferguson WTU-76670 WTU-175473

Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. U.S.A., ALASKA COUNTY: CANADA, BRITISH COLUMBIA: Mount McKinley [Denali] National Park; near Park Headquarters. Queen Charlotte Islands, 7 miles north of Port Clements o road to 63.720978°, -148.965722°; uncertainty: 5000 m., Georef'd by Masset, Graham Island. Joseph Stormer Large sphagnum bog especially on hummocks near margin. Under willows in Sphagnum. Phenology: Flowers. Origin: Native. Common. Phenology: Flowers. Aven Nelson 3528 24 Jun 1939 J. A. Calder 34700 6 Jun 1964 with Ruth A. Nelson with R. L. Taylor, L. C. Sherk WTU-75133 WTU-242729

Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. U.S.A., ALASKA, KENAI PENINSULA COUNTY: CANADA, BRITISH COLUMBIA: About 2 miles north of Divide. Queen Charlotte Islands, about half a mile southwest of Towhill, 14 60° 17' N, 149° 1' W miles east of Massett, Graham Island. Wet, boggy-marshy area by pond. Flowers white, common. Bog, scattered throughout Sphagnum bog. Common, flowers white. Phenology: Flowers. Origin: Native. Phenology: Flowers. J. A. Calder 5328 20 Jun 1951 J. A. Calder 21157 4 Jun 1957 with D. B. O. Savile, R. L. Taylor WTU-173254 WTU-240953 Rosaceae Rosaceae Rubus chamaemorus L. Rubus bifrons Vest U.S.A., ALASKA COUNTY: U.S.A., WASHINGTON, KING COUNTY: Northeast Alaska. Okpilak Valley, West Okpilak Lake. Seattle, Madrona Drive. Elev. 1902 ft. 47.60833°, -122.29417°; WGS 84, uncertainty: 10000 m., Source: 69° N, 144° W; Misleading coordinates given only to nearest GeoLocate, Georef'd by WTU Staff degree. Phenology: Flowers & Fruits. Origin: Introduced. Moist tundra. Phenology: Flowers. Origin: Native. Gray 6034 6 Sep 1975 N. E. Grulke 1083 21 Jun 1985 with D. J. Marrett WTU-330329 WTU-312731 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus bifrons Vest CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, KITTITAS COUNTY: Along PGE tracks, Aeta Lake. Along north Thorp Highway near junction with SR 10. Phenology: Fruits. Elev. 1740 ft. T19N R17E S34; WGS 84, uncertainty: 1000 m., Source: Garman 5477 23 Jul 1967 GeoLocate, Georef'd by WTU Staff South-facing slope of road bank, sand and gravel, with Populus WTU-329144 trichocarpa, Lomaium dissectum, Epilobium paniculatum, Amsinckia lycopsoides. Phenology: Flowers. Origin: Introduced. Rosaceae Don Knoke 447 11 Jun 2004 Rubus leucodermis Douglas ex Torr. & A. Gray WTU-357409 U.S.A., OREGON, JOSEPHINE COUNTY: Along the Caves Nature Trail, Caves National Monument & Vicinity. Rosaceae Phenology: Fruits. Origin: Native. Rubus bifrons Vest William H. Baker 199 30 Jun 1949 with George C. Ruhle CANADA, BRITISH COLUMBIA: Queen charlotte Islands, Queen Charlotte City, Graham Island, WTU-274605 places between Tlell and Skidegate. A escape in all the settled areas, Masset, Sandspit, Port Clements and in a few places. Widely introduced. Phenology: Flowers. Rosaceae J. A. Calder 36946 14 Aug 1964 Rubus leucodermis Douglas ex Torr. & A. Gray with R. L. Taylor CANADA, BRITISH COLUMBIA: WTU-242720 Junction of Ewart Creek and Ashnola River. Shaded roadside. Phenology: Flowers. Rosaceae R. Hainault 7768 7 Feb 1976 Rubus bifrons Vest WTU-274078 U.S.A., WASHINGTON, KLICKITAT COUNTY: Vila. Rosaceae 45.70102°, -121.3883°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by Rubus leucodermis Douglas ex Torr. & A. Gray WTU Staff Phenology: Flowers. Origin: Introduced. U.S.A., OREGON COUNTY: Umatilla River near Bingham Springs (for flowers). Wallowa Wilhelm N. Suksdorf 12002 Mountains (for fruit). 45.740935°, -118.233709°; uncertainty: 4000 m., Georef'd by 20 May 1926 Joseph Stormer Nowhere abundant. Flowers collected on May 21 1908. Fruit WTU-196418, WTU-V-060714 collected on Jul 1908. Phenology: Fruits. Origin: Native. Wm. C. Cusick 3237c 21 May 1908 Rosaceae WTU-185980 Rubus bifrons Vest U.S.A., WASHINGTON, KING COUNTY: Seattle, Madrona Drive. 47.60833°, -122.29417°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Origin: Introduced. Gray 6034 6 Sep 1975 WTU-330330 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., OREGON, WASHINGTON COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: Near Forest Grove. Siskiyou National Forest. Along Forest Road 4105, 7.4 miles from 45.519836°, -123.110663°; WGS 84, uncertainty: 7500 m., junction with Forest Road 4103. Source: GeoLocate, Georef'd by Joseph Stormer Elev. 2650 ft. Open slopes. Phenology: Flowers & Fruits. Origin: Native. 42° 20' 29" N, 123° 44' 7" W; T37S R9W S23 Full sun, dry soil. Shrub to 1 meter, arching stems, leaf underside J. William Thompson 581 15 Apr 1926 white. Phenology: Fruits. Origin: Native. WTU-23003JWT Roger N. Fuller 98-12 27 Jun 1998 WTU-345721 Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rosaceae U.S.A., IDAHO, BOISE COUNTY: Rubus leucodermis Douglas ex Torr. & A. Gray Harris Creek Road, east of Horseshoe Bend 4.2 miles west of Harris Creek Summit, 1.2 miles east of Appletree Gulch. U.S.A., OREGON, JOSEPHINE COUNTY: Elev. 4000 ft. Forest Highway 25, about 1 mile north of Serpentine Point, Siskiyou T6N R3E S11 NW 1/4; WGS 84, uncertainty: 1000 m., Source: National Forest. GeoLocate, Georef'd by WTU Staff Elev. 3798 ft. Pine and douglas-fir forest on granite, granite roadcut. Phenology: 42° 23' 5" N, 123° 38' 23" W Flowers. Origin: Native. Roadside and meadow, on bank above road with Ceanothus, madrone, manzanita, etc. Upright, vigorous. Phenology: Flowers. Barbara Ertter 4250 19 Jun 1981 Origin: Native. with Larry Ertter Jessie Johanson 32 27 Jun 1998 WTU-286638 with Joe Johanson WTU-342594 Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rosaceae U.S.A., IDAHO, BONNER COUNTY: Rubus leucodermis Douglas ex Torr. & A. Gray Priest River Experimental Forest. 48.355678°, -116.787586°; WGS 84, uncertainty: 4000 m., U.S.A., OREGON, JOSEPHINE COUNTY: Source: Digital Map, Georef'd by Melissa Bowers Forest road 176 off State Route 46, circa 1 mile west of Oregon Arborvitae-hemlock vegetation zone, seral, open forest. Phenology: Caves National Monument. Siskiyou National Forest. Fruits. Origin: Native. Elev. 3650 ft. 42° 6' 35" N, 123° 24' 37" W; T40S R6W S4 or 9 R. F. Daubenmire 44492 17 Sep 1944 North-facing slope, mixed fir/oak forest, along 176 road bank. Long reddish new wood. Phenology: Flowers. Origin: Native. WTU-251477 Jessie Johanson 06 26 Jun 1998 with Joe Johanson Rosaceae WTU-342670 Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., OREGON, WALLOWA COUNTY: Rosaceae Imnaha. 45.559323°, -116.833207°; WGS 84, uncertainty: 5000 m., Rubus leucodermis Douglas ex Torr. & A. Gray Source: GeoLocate, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. U.S.A., OREGON, JOSEPHINE COUNTY: Forest road 176 off State Route 46, circa 1 mile west of Oregon Wm. Sherwood 128 4 Jun 1923 Caves National Monument. Siskiyou National Forest. Elev. 3650 ft. WTU-23005JWT 42° 6' 35" N, 123° 24' 37" W; T40S R6W S4 or 9 North-facing slope, mixed fir/oak forest. Phenology: Flowers. Origin: Native. Rosaceae Jessie Johanson 10 26 Jun 1998 Rubus leucodermis Douglas ex Torr. & A. Gray with Joe Johanson U.S.A., IDAHO, BONNER COUNTY: WTU-342658 Along Pend Oreille River, four miles east of Priest River. 48.200171°, -116.90305°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Open pine woods. Erect shrub. Phenology: Flowers. Origin: Native. William H. Baker 8968 31 May 1952 WTU-154726 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., OREGON, YAMHILL COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: In the Coast Range at Walker Flat, about 11.5 airline miles west of Siskiyou National Forest. Along Forest Road 4105, 7.4 miles from Carlton. junction with Forest Road 4103. Elev. 1810 ft. Elev. 2650 ft. T3S R6W S22; NAD 27, uncertainty: 805 m., Source: TRS2LL, 42° 20' 29" N, 123° 44' 7" W; T37S R9W S23 Georef'd by Ben Legler Full sun, dry soil. Shrub to 1 meter, arching stems, leaf underside On clear-cut hillside, with Acer, Cirsium, Polystichum, Luzula, white. Phenology: Fruits. Origin: Native. Senecio, Pteridium. Arching canes to 1.7 meters tall. Phenology: Fruits. Origin: Native. Roger N. Fuller 98-12 27 Jun 1998 Richard R. Halse 4943 5 Jul 1995 WTU-349144 WTU-333603 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., OREGON, JOSEPHINE COUNTY: Siskiyou National Forest. Along Forest Road 4105, 7.4 miles from U.S.A., OREGON COUNTY: junction with Forest Road 4103. Umatilla River near Bingham Springs (for flowers). Wallowa Elev. 2650 ft. Mountains (for fruit). 42° 20' 29" N, 123° 44' 7" W; T37S R9W S23 45.740935°, -118.233709°; uncertainty: 4000 m., Georef'd by Full sun, dry soild. Shrub to 1 meter, arching stems, leaf underside Joseph Stormer white. Phenology: Fruits. Origin: Native. Nowhere abundant. May 21 1908 (for flowers), Jul 1908 (for fruit). Phenology: Flowers & Fruits. Origin: Native. Roger N. Fuller 98-12 27 Jun 1998 Wm. C. Cusick 3237c 21 May 1908 WTU-347852 WTU-140028 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, PIERCE COUNTY: Mount Rainier, Glacier Road. U.S.A., OREGON, CLACKAMAS COUNTY: Elev. 5000 ft. 2 miles east of Zigzag. 47.19083°, -122.58139°; WGS 84, uncertainty: 10000 m., Source: Elev. 2300 ft. GeoLocate, Georef'd by WTU Staff, Invalid coordinates (misplaced Sandy soil in open in Pseudotsuga-Pinus contorta forest. Phenology: by georeferencing program). Flowers. Origin: Native. Phenology: Fruits. Origin: Native. C. L. Hitchcock 4782 20 Jun 1939 J. M. Grant s.n. Aug 1925 with J. S. Martin WTU-22088 WTU-53633 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Marysville. Badger Creek Study Area, Lochsa River Canyon, about 40 miles 48.05194°, -122.17583°; WGS 84, uncertainty: 3570 m., Source: southwest of Missoula, Montana. GeoLocate, Georef'd by Maria Yousoufian 46.504525°, -114.822919°; WGS 84, uncertainty: 10000 m., Field. Phenology: Flowers. Origin: Native. Source: GeoLocate, Georef'd by WTU Staff Along logging roads in clear-cut areas, western red cedar/white pine J. M. Grant s.n. May 1930 forest on old river terrace. Phenology: Fruits. Origin: Native. WTU-42896 P. F. Stickney 853 26 Jul 1963 WTU-295869 Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rosaceae U.S.A., WASHINGTON, LEWIS COUNTY: Rubus leucodermis Douglas ex Torr. & A. Gray Cowlitz Valley. 46.538271°, -121.852778°; WGS 84, uncertainty: 20000 m., U.S.A., IDAHO, LATAH COUNTY: Source: Digital Map, Georef'd by Dan Post Tomer's Butte, 4 miles southwest of Moscow. No collection date given. Phenology: Flowers. 46.706979°, -116.923111°; uncertainty: 1500 m., Georef'd by Joseph Stormer R. Ernesti E2134 Wooded draw on east slope. Phenology: Fruits. Origin: Native. WTU-316469 William H. Baker 14508 20 Jul 1956 WTU-295926 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus parviflorus Nutt. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Tye. Seattle. Elev. 3100 ft. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., 47.74944°, -121.11944°; WGS 84, uncertainty: 10000 m., Source: Source: Digital Map, Georef'd by Joseph Stormer GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Phenology: Fruits. Origin: Native. E. S. Meany s.n. 1898 G. N. Jones 5709 27 Jul 1934 WTU-36216 WTU-101589 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Campus State University. Seattle. 47.66667°, -122.30861°; WGS 84, 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., uncertainty: 10000 m., Source: GeoLocate, Source: Digital Map, Georef'd by Joseph Stormer Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. C. V. Piper 68 13 Jun 1888 T. C. Frye s.n. WTU-9599 6 Oct 1904 WTU-9745, WTU-V-045450 Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rosaceae U.S.A., WASHINGTON, KING COUNTY: Seattle. Rubus parviflorus Nutt. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer U.S.A., WASHINGTON, KING COUNTY: Phenology: Flowers. Origin: Native. University of Washington Campus. 47.66667°, -122.30861°; WGS 84, Chas. V. Piper 68 Jun 1891 uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-9597 Phenology: Flowers. W. V. Lovitt s.n. Rosaceae May 1909 Rubus parviflorus Nutt. WTU-9651, WTU-V-045627 U.S.A., WASHINGTON, KING COUNTY: Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, Rosaceae Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Rubus parviflorus Nutt. C. V. Piper 460 U.S.A., WASHINGTON, KING COUNTY: Seward Park, Seattle. 9 Jun 1897 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, WTU-9603, WTU-V-045453 Georef'd by WTU Staff Phenology: Flowers. Origin: Native. G. L. Wittrock s.n. Rosaceae 3 Dec 1927 Rubus parviflorus Nutt. WTU-28128, WTU-V-045624 U.S.A., WASHINGTON, KING COUNTY: Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. E. S. Meany s.n. 1898 WTU-36215, WTU-V-045611 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: South Seattle. At 6 mile mark on trail to 8-Mile Lake, just above 47.60639°, -122.33083°; WGS 84, Little 8-Mile Lake, Icicle Creek Drainage. uncertainty: 10000 m., Source: GeoLocate, 47.557948°, -120.672115°; WGS84, Georef'd by WTU Staff uncertainty: 10000 m., Source: GeoLocate, Semi-open slopes and along streams. Georef'd by WTU Staff Phenology: Flowers. Origin: Native. In shallow stony soil of massive peridotite talus. Leaves unusually bullate, deeply rugose and J. William Thompson 11573 veined. Plants low, very glandular. Phenology: Flowers. Origin: Native. 1 Jun 1935 A. R. Kruckeberg 5423 WTU-32306, WTU-V-045452 7 Jul 1962 WTU-217554, WTU-V-045622 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., WASHINGTON, SNOHOMISH COUNTY: Rubus parviflorus Nutt. 1 mile out from Big 4 Inn. Elev. 1800 ft. U.S.A., WASHINGTON, ISLAND COUNTY: 48.06765°, -121.515°; WGS 84, uncertainty: Goose Rock, Deception Pass State Park, 1800 m., Source: GeoLocate, Georef'd by Whidbey Island. WTU Staff 48.39389°, -122.6475°; WGS 84, uncertainty: (Puget Sound Botanical Club accession number 10000 m., Source: GeoLocate, Georef'd by 54.). Phenology: Flowers. Origin: Native. WTU Staff Meadows. Phenology: Flowers. Origin: Native. Ruby L. C. Sovereign 120 Harold W. Smith 949 26 May 1940 9 Jun 1936 WTU-315752, WTU-V-045451 WTU-40628, WTU-V-045443 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., WASHINGTON, KING COUNTY: 8546 - 30th Avenue NW, Seattle. Rubus parviflorus Nutt. 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, U.S.A., WASHINGTON, PIERCE COUNTY: Georef'd by WTU Staff Tacoma. Shady woods. (Puget Sound Botanical Club 47.25306°, -122.44306°; WGS 84, accession number 120.). Phenology: Fruits. uncertainty: 10000 m., Source: GeoLocate, Origin: Native. Georef'd by WTU Staff Shady ravines. Indefinite loan by Tacoma High Edith Hardin English 1042 School to University of Washington. Phenology: Flowers. Origin: Native. 30 Jul 1940 J. B. Flett s.n. WTU-315753, WTU-V-045448 27 May 1896 WTU-36736, WTU-V-045446 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., WASHINGTON, SNOHOMISH COUNTY: Rubus parviflorus Nutt. 2 miles east of Gold Bar along Forest Service Road 6010; along Skykomish River. U.S.A., WASHINGTON, CLALLAM COUNTY: Elev. 1000 ft. Mount Angeles. T27N R9E S9; NAD 27, uncertainty: 805 m., Elev. 2500 ft. Source: TRS2LL, Georef'd by Ben Legler 47.99538°, -123.46784°; WGS 84, Deciduous forest of Acer and Populus; uncertainty: 2500 m., Source: GeoLocate, understory of ferns and other Rosaceae; near Georef'd by WTU Staff river, soil of gravely glacial till. Phenology: Phenology: Flowers. Origin: Native. Flowers. Origin: Native. G. B. Rigg s.n. L. Hegstrom 13 22 Jul 1929 10 May 1986 WTU-36214, WTU-V-045449 WTU-313224, WTU-V-045619 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, SPOKANE COUNTY: U.S.A., WASHINGTON, SNOHOMISH Clark Springs, 10 miles north of Spokane. COUNTY: 47.803635°, -117.425°; WGS 84, uncertainty: Silverton. 1000 m., Source: GeoLocate, Georef'd by 48.07844°, -121.567062°; WGS 84, WTU Staff uncertainty: 5000 m., Source: GeoLocate, Deep fertile soil. Phenology: Flowers. Origin: Georef'd by Joseph Stormer Native. Phenology: Flowers. Origin: Native. F. O. Kreager 37 L. S. Bouch s.n. 26 Jun 1902 10 Jun 1895 WTU-68594, WTU-V-045616 WTU-9847, WTU-V-045615

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, ISLAND COUNTY: Rainbow Creek trail at Bench Creek. Whidbey Island. 48.379651°, -120.688234°; WGS84, 48.146201°, -122.552685°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, uncertainty: 30000 m., Source: Digital Map, Georef'd by WTU Staff Georef'd by Joseph Stormer Rocky banks near creek. Phenology: Fruits. Phenology: Flowers. Origin: Native. Origin: Native. N. L. Gardner s.n. George H. Ward 119 20 Jun 1897 22 Aug 1942 WTU-9739, WTU-V-045614 WTU-182719, WTU-V-045613

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., IDAHO, BLAINE COUNTY: U.S.A., IDAHO, BOUNDARY COUNTY: Alpine Creek. Smith Creek Region, along Smith Creek, 10 miles due west of Port Elev. 8956 ft. Hill. 43° 54.3' N, 114° 57.2' W Elev. 4500 ft. Soil boggy and wet under trees, Pinus contorta, Abies forest 48.963645°, -116.53169°; WGS 84, uncertainty: 10000 m., intermixed with Artemisia between forest patches. Corrolla white, Source: GeoLocate, Georef'd by WTU Staff shrub to 0.5 m. Phenology: Flowers. Origin: Native. Virgin coniferous forest. Phenology: Flowers. Origin: Native. J. F. Smith 4689 12 Jul 2003 G. B. Rossbach 204 12 Jul 1936 with R. P. Rossbach WTU-354965 WTU-160208 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., IDAHO, CLEARWATER COUNTY: 8 miles northeast of Elk River. U.S.A., WASHINGTON, ISLAND COUNTY: 46.86544°, -116.059296°; WGS 84, uncertainty: 1000 m., Source: Langley. GeoLocate, Georef'd by WTU Staff 48.04028°, -122.405°; WGS 84, uncertainty: Steep, wooded slope. Phenology: Vegetative. Origin: Native. 10000 m., Source: GeoLocate, Georef'd by WTU Staff Mary Hafercamp 119 21 Jul 1948 Phenology: Flowers. Origin: Native. with Xerpha M. Gaines J. M. Grant s.n. WTU-132730 May 1923 WTU-34731, WTU-V-045618 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Three Tree Point. Seattle, W.T. 47.45056°, -122.38111°; WGS 84, 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Georef'd by WTU Staff Open woods. Phenology: Flowers. Origin: Phenology: Flowers. Origin: Native. Native. E. S. Meany s.n. Lyman Benson s.n. May 1885 10 May 1929 WTU-9640, WTU-V-045617 WTU-40681, WTU-V-045628

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., MONTANA, FLATHEAD COUNTY: U.S.A., WASHINGTON, KING COUNTY: Along north side of South Fork of Flathead River, about 7 miles east Seattle. of Columbia Falls. 47.613617°, -122.310098°; WGS 84, 48.365521°, -114.040337°; WGS 84, uncertainty: 5000 m., uncertainty: 15750 m., Source: Digital Map, Source: Digital Map, Georef'd by Joseph Stormer Georef'd by Joseph Stormer Woods. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. H. T. Rogers 1045 12 Jul 1942 Lizzy Hardy s.n. with J. M. Rogers 25 May 1890 WTU-241574 WTU-9772, WTU-V-045442 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., WASHINGTON, KING COUNTY: Rubus parviflorus Nutt. [No locality given on label]. 47.47242°, -121.86538°; WGS 84, U.S.A., WASHINGTON, KING COUNTY: uncertainty: 65000 m., Source: GeoLocate, Seattle. Georef'd by WTU Staff 47.613617°, -122.310098°; WGS 84, [Collection year on label unclear, written;-188-]. uncertainty: 15750 m., Source: Digital Map, Phenology: Flowers. Origin: Native. Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. C. V. Piper s.n. Chas. V. Piper 66 3 Jun 1885 2 Jun 1891 WTU-9601, WTU-V-045629 WTU-9602, WTU-V-045626

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., OREGON, COOS COUNTY: Isthmus Slough. U.S.A., IDAHO, LATAH COUNTY: Elev. 25 ft. Thatuna Ridge. T26S R13W S23; NAD 27, uncertainty: 805 m., Source: TRS2LL, Phenology: Flowers. Origin: Native. Georef'd by Ben Legler Douglas Fir Forest, east hill slope, moderately moist clay-loam in R. F. Daubenmire 37382 28 Jun 1937 moderate shade. Association: Pseudotsuga taxifolia- Tsuga heterophylla- Thuja plicata- Vaccinium parvifolium- V. ovatum. WTU-239885 Phenology: Flowers. Origin: Native. L. E. Detling 4098 21 Apr 1940 WTU-74211 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Seattle. Friday Harbor. 47.613617°, -122.310098°; WGS 84, 48.534266°, -123.017124°; WGS 84, uncertainty: 15750 m., Source: Digital Map, uncertainty: 5000 m., Source: GeoLocate, Georef'd by Joseph Stormer Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Phenology: Flowers & Fruits. Origin: Native. Emma A. Shumway s.n. A. S. Pope s.n. May 1892 23 Jun 1904 WTU-112691, WTU-V-045456 WTU-9540, WTU-V-045465

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., MONTANA, FLATHEAD COUNTY: CANADA, BRITISH COLUMBIA: 1 mile east of Bigfork on Swan Lake road. Goldstream, Vancouver Island. Elev. 3200 ft. Phenology: Flowers. T29N R19W Growing under dense coniferous timber and along roadbed where C. F. Newcombe s.n. 31 May 1924 shaded. Grows up to 3 feet high, woody stem, white corolla. Phenology: Flowers. Origin: Native. WTU-133296 Fred Zwickel 25 24 Jun 1949 Rosaceae WTU-182182 Rubus parviflorus Nutt. Rosaceae U.S.A., WASHINGTON, STEVENS COUNTY: Northeast of Leadpoint. Frisco Mine Road. Rubus parviflorus Nutt. Elev. 4250 ft. 48.90944°, -117.58639°; WGS 84, U.S.A., WASHINGTON, CLALLAM COUNTY: uncertainty: 10000 m., Source: GeoLocate, Olympic Peninsula; Lake Crescent. Georef'd by WTU Staff 48.073659°, -123.827282°; WGS 84, Northwest slope, moist rocky bank of stream, uncertainty: 6900 m., Source: Digital Map, open forest. Phenology: Flowers. Origin: Native. Georef'd by Joseph Stormer Phenology: Fruits. Origin: Native. Geo. G. Hedgcock s.n. Geo. N. Jones 3474 7 Jul 1933 Aug 1931 WTU-108123, WTU-V-045464 WTU-21683, WTU-V-045462 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: King/Snohomish County border. 205th and Merritt. Alaska Way. Undeveloped property on the north Elev. 2500 ft. border of LFP. 47.78111°, -120.84194°; WGS84, uncertainty: 47.91306°, -122.09694°; WGS 84, 10000 m., Source: GeoLocate, Georef'd by uncertainty: 10000 m., Source: GeoLocate, WTU Staff Georef'd by WTU Staff Phenology: Flowers. Origin: Native. By chain-link fence along cedar way by rock-drainage structure on stream. Phenology: James D. Currie 109 Flowers. 20 Jun 1948 Christie S. Isler 55-5-98 WTU-189360, WTU-V-045463 16 May 1998 WTU-335281, WTU-V-045467 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus lasiococcus A. Gray U.S.A., OREGON, BENTON COUNTY: U.S.A., OREGON, HOOD RIVER COUNTY: East foot of Alsea Mountain. Mount Hood. 44.468878°, -123.488983°; WGS 84, uncertainty: 5000 m., 45.373343°, -121.695728°; WGS 84, uncertainty: 10000 m., Source: Digital map, Georef'd by Joseph Stormer Source: GeoLocate, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. Garland M. Powell s.n. 27 May 1939 Thos. Howell 1355 24 Aug 1892 WTU-131738 WTU-22617JWT-A

Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus parviflorus Nutt. U.S.A., OREGON, JOSEPHINE COUNTY: CANADA, BRITISH COLUMBIA: Above Bolan Lake, Siskiyou Mountains, near California border. Subalpine slopes near Lake Bootahnie [Botanie Lake], Marble Elev. 5800 ft. Mountains. On rocky, springy soil. Phenology: Flowers. Origin: Native. Elev. 5000 ft. 50.383684°, -121.541882°; uncertainty: 1464 m., Georef'd by C. L. Hitchcock 5266 7 Jul 1939 Joseph Stormer with J. S. Martin Phenology: Flowers. WTU-53166 J. William Thompson 190 20 Jun 1938 with Emily M. Thompson Rosaceae WTU-16932JWT Rubus lasiococcus A. Gray Rosaceae U.S.A., OREGON, CLACKAMAS COUNTY: Mount Hood. Government Camp. Rubus lasiococcus A. Gray Elev. 4100 ft. 45.304007°, -121.7548°; WGS 84, uncertainty: 3036 m., Source: U.S.A., OREGON, HOOD RIVER COUNTY: GeoLocate, Georef'd by Joseph Stormer Near Elk Meadows, Mount Hood. Phenology: Flowers. Origin: Native. Elev. 4000 ft. 45.344364°, 0°; uncertainty: 2000 m. Lewis S. Rose 37433 6 Jul 1937 Alpine slopes. Phenology: Flowers. Origin: Native. WTU-51713 J. William Thompson 3291 31 Jul 1927 WTU-22617JWT-B Rosaceae Rubus lasiococcus A. Gray Rosaceae U.S.A., OREGON COUNTY: Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Mount Hood. Elev. 4000 ft. CANADA, BRITISH COLUMBIA: 45.373343°, -121.695728°; WGS 84, uncertainty: 2000 m., Lillooet District: Bold northwest-facing dunite outcrop and talus Source: GeoLocate, Georef'd by Sara Shiina above Cadwallader Creek Canyon and just south of Piebiter Creek; In woods. Indefinite loan by Tacoma High School to the University of about 10 miles south and above Bralorne. Washington. Phenology: Fertile. Origin: Native. Elev. 5200 ft. Lone ultra-basic outcrop surrounded by massive T. C. Frye s.n. 30 Aug 1907 argillite-chert-limestone beds; lodgepole pine and Juniperus communis dominant woody species here. Phenology: Fertile. WTU-36220 A. R. Kruckeberg 5736 10 Sep 1963 Rosaceae WTU-214280 Rubus sachalinensis H. Lév. var. sachalinensis U.S.A., ALASKA COUNTY: Near Fairbanks. Spruce forest, mossy open plain. Flowers white, bushes to 2 feet tall. Phenology: Flowers. Origin: Native. Nora Nicely 71 2 Jul 1970 WTU-266075 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., IDAHO, IDAHO COUNTY: West side of Wenatchee River at upper end of Southwest facing slope, Canteen Meadows, 1/2 mile northwest of Tumwater Canyon. Big Fog Mountain; Gedney Creek to Selway River drainage, Nez T25N R17E; WGS84, uncertainty: 10000 m., Perce National Forest. Source: GeoLocate, Georef'd by WTU Staff Elev. 7500 ft. Steep hillside. Phenology: Flowers. Origin: 46.144959°, -115.231695°; WGS 84, uncertainty: 1000 m., Native. Source: GeoLocate, Georef'd by WTU Staff In crevices of loose boulders. Phenology: Fertile. George H. Ward 492 A. R. Kruckeberg 4110 8 Jul 1957 6 Jun 1946 WTU-191986 WTU-182724, WTU-V-045468 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus lasiococcus A. Gray U.S.A., IDAHO, IDAHO COUNTY: Middle Fork of Clearwater River at mouth of 3 Devils Creek, 4 miles U.S.A., OREGON, KLAMATH COUNTY: below Lowell, Nez Perce National Forest. Castle Mountain, south side of Crater Lake. 46.086257°, -115.59472°; WGS 84, uncertainty: 1000 m., Source: Forest of black hemlock. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Rich humus under coniferous climax. Phenology: Flowers. Origin: Elmer I. Applegate 175 12 Aug 1896 Native. WTU-8827JWT L. Constance 1614 31 May 1936 with R. C. Rollins Rosaceae WTU-9587JWT Rubus lasiococcus A. Gray Rosaceae U.S.A., WASHINGTON COUNTY: Emmons Woods, Mount Rainer. Rubus lasiococcus A. Gray Elev. 6000 ft. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, CHELAN COUNTY: Steven's Pass. J. M. Grant s.n. Aug 1925 Elev. 4000-6000 ft. 47.744438°, -121.090111°; WGS84, uncertainty: 600 m., Source: WTU-22087 Georeferenced, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Rosaceae J. H. Sandberg s.n. Aug 1893 with J. B. Leiberg Rubus parviflorus Nutt. WTU-9522 U.S.A., WASHINGTON, CLALLAM COUNTY: Lake Crescent; on Mount Storm King. Elev. 600-4300 ft. Rosaceae 48.09472°, -123.80583°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Rubus parviflorus Nutt. Georef'd by WTU Staff Partially shaded places. Phenology: Flowers. U.S.A., IDAHO, CLEARWATER COUNTY: Origin: Native. Along the Clearwater River, 5 miles south of Orofino. W. H. Lawrence 332 46.407053°, -116.25417°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, 23 Jul 1904 Georef'd by WTU Staff, Invalid coordinates (located outside county bounds). WTU-96188, WTU-V-045466 Granite cliffs. Phenology: Flowers. Origin: Native. William H. Baker 13977 27 May 1956 WTU-295841, WTU-V-045470 Rosaceae Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., IDAHO, LEMHI COUNTY: U.S.A., MONTANA, CARBON COUNTY: 1 mile southeast of Sleeping Deer Mountain, 37 miles northwest of Along Rock Creek, 10 miles south of Red Lodge. Challis. In semi-meadow. Fruits red (not distinct from R. strigosus-R. 44.757259°, -114.676137°; WGS 84, uncertainty: 1000 m., melanolasius). Phenology: Flowers & Fruits. Source: GeoLocate, Georef'd by WTU Staff Among rocks on high ridge. Phenology: Fertile. C. L. Hitchcock 13565 12 Aug 1945 with C. V. Muhlick C. L. Hitchcock 11351 22 Aug 1944 with C. V. Muhlick WTU-98057 WTU-91508 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus parviflorus Nutt. U.S.A., WASHINGTON, CLALLAM COUNTY: Boulder Creek. U.S.A., OREGON, DOUGLAS COUNTY: 47.98167°, -123.60139°; WGS 84, uncertainty: 10000 m., Source: Roseburg Quadrangle. GeoLocate, Georef'd by WTU Staff 43.25°, -123.5°; WGS 84, uncertainty: 49000 Phenology: Flowers. Origin: Native. m., Source: Digital map, Georef'd by Joseph Stormer George Neville Jones 8492 6 Sep 1935 Phenology: Flowers. Origin: Native. WTU-31538 W. C. Cusick 4187 22 Jun 1914 Rosaceae WTU-135979, WTU-V-045471 Rubus lasiococcus A. Gray U.S.A., WASHINGTON, KITTITAS COUNTY: Scatter Creek Trail, 2 miles from junction with Fish Creek Road. Rosaceae Elev. 4200 ft. 47.507151°, -121.059494°; WGS 84, uncertainty: 10000 m., Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Source: GeoLocate, Georef'd by WTU Staff Under conifers near stream. Phenology: Flowers. Origin: Native. U.S.A., MONTANA, BEAVERHEAD COUNTY: Above Torrey Lake, Pioneer Range. C. L. Hitchcock 8012 6 Jul 1942 Talus slope. Plant tomentose and glandular. Phenology: Fertile. WTU-75341 C. L. Hitchcock 15074 27 Jul 1946 with C. V. Muhlick Rosaceae WTU-107658 Rubus parviflorus Nutt. Rosaceae U.S.A., OREGON, HOOD RIVER COUNTY: Mount Hood. Bluegrass Ridge. Rubus parviflorus Nutt. Elev. 5000 ft. 45.364284°, -121.596463°; WGS 84, U.S.A., WASHINGTON, SNOHOMISH uncertainty: 5000 m., Source: GeoLocate, COUNTY: Georef'd by Joseph Stormer Mount Dickerman, Cascade Mountains. Along stream on south slope. Phenology: Elev. 3000 ft. Flowers. Origin: Native. 48.06829°, -121.47068°; WGS 84, uncertainty: 3000 m., Source: GeoLocate, J. William Thompson 3211 Georef'd by WTU Staff Open woods. Phenology: Flowers. Origin: 31 Jul 1927 Native. WTU-23001JWT, WTU-V-045475 J. William Thompson 14742 15 Jun 1940 Rosaceae WTU-21917JWT, WTU-V-045469 Rubus lasiococcus A. Gray Rosaceae U.S.A., WASHINGTON, CLALLAM COUNTY: Rubus parviflorus Nutt. Olympic National Park, Idaho Camp. Elev. 700 ft. U.S.A., MONTANA, GLACIER COUNTY: 48.26667°, -124.675°; WGS 84, uncertainty: 10000 m., Source: Glacier National Park, foot of Mount Wilbur; Swiftcurrent Valley. GeoLocate, Georef'd by WTU Staff Elev. 5200 ft. Phenology: Flowers. Origin: Native. Base of 20 ft. cliff. Phenology: Flowers. Origin: Native. Ruby L. C. Sovereign 565 11 Jul 1941 Daniel H. Brant 113 28 Jul 1948 WTU-316853 WTU-129560 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus idaeus L. var. sachalinensis (H. Lév.) Focke CANADA, BRITISH COLUMBIA: U.S.A., MONTANA COUNTY: U.B.C. campus. Glacier National Park, Sun Camp. 49° 15' N, 123° 15' W Slide rock on hillside. Phenology: Fertile. Open clearing in forest. Acc # v177508. Coll: JP. Luitjens & students. Students collection for Botany 311. Phenology: Vegetative. W. T. McLaughlin 2949 24 Jul 1933 J. P. Luitjens 690164 12 May 1976 WTU-22558 WTU-304356 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus parviflorus Nutt. U.S.A., OREGON, TILLAMOOK COUNTY: Near Bay City. U.S.A., ALASKA COUNTY: Phenology: Flowers. Origin: Introduced. North Tongass National Forest, 1 mile east of Juneau, Basin Road. G. E. Morrill 166 8 Jul 1933 Elev. 500 ft. 58.308074°, -134.389992°; WGS 84, WTU-15798JWT uncertainty: 1309 m., Source: Digital Map, Georef'd by Joseph Stormer Cutover forest. Petals white, shrub 3-4 feet. Rosaceae Phenology: Flowers. Origin: Native. Rubus laciniatus Willd. Elbert L. Little Jr. 18524 U.S.A., OREGON, JOSEPHINE COUNTY: 21 Jun 1961 Along Sucker Creek at Three Star Mine, Oregon Caves National Monument and Vicinity. WTU-250593, WTU-V-045473 Phenology: Flowers. Origin: Introduced. William H. Baker 967 21 Aug 1950 with George C. Ruhle Rosaceae WTU-274603 Rubus lasiococcus A. Gray U.S.A., WASHINGTON COUNTY: Rosaceae White Pass, Wenatchee National Forest. Elev. 6000 ft. Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Grass type; wet loam; no slope; Carex, Poa asssociation. Common locally. Speciman typical. Phenology: Flowers. Origin: Native. CANADA, BRITISH COLUMBIA: 4 miles north of Lower Post. F. G. Renner 738 1 Aug 1925 59° 59' N, 128° 33' W Canyon, along highway, brown sandy soil. White flower. Phenology: WTU-323565 Fertile. G. A. Corcoran 63 1 Jul 1951 Rosaceae WTU-143635 Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., IDAHO, LEMHI COUNTY: Rosaceae Along Panther Creek 5 miles above Cabin Creek. 44.88008°, -113.979338°; WGS 84, uncertainty: 1000 m., Source: Rubus idaeus var. sachalinensis GeoLocate, Georef'd by WTU Staff In talus. Phenology: Fertile. U.S.A., IDAHO, CUSTER COUNTY: Along trail between Farley Lake and Toxaway Lake; Sawtooth C. L. Hitchcock 14249 1 Jul 1946 Primitive Area, Sawtooth Mountains. with C. V. Muhlick 43.98°, -114.93139°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-104487 Characteristically in rocks at base of bouldery talus. Phenology: Fertile. Rosaceae A. R. Kruckeberg 4216 19 Jul 1957 Rubus laciniatus Willd. WTU-189794 U.S.A., OREGON, TILLAMOOK COUNTY: Near Bay City. Phenology: Flowers. Origin: Introduced. G. E. Morrill 166 8 Jul 1933 WTU-37691 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus laciniatus Willd. U.S.A., IDAHO, IDAHO COUNTY: U.S.A., WASHINGTON, KING COUNTY: Whitebird Ridge, Little Salmon River Canyon. Seattle, South Park. 45.416646°, -116.314538°; WGS 84, uncertainty: 10000 m., 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Dry hillsides. Phenology: Flowers. Origin: Introduced. Damp ground. Phenology: Vegetative. Origin: Introduced. William H. Baker 12150 27 Jul 1954 C. Grainger 181 30 Apr 1910 WTU-295864 WTU-9773

Rosaceae Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rubus laciniatus Willd. U.S.A., MONTANA, FLATHEAD COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Bad Rock Canyon, south side of Flathead River, 4 miles east of Marysville. Columbia Falls. 48.05194°, -122.17583°; WGS 84, uncertainty: 3570 m., Source: 48.387538°, -114.092051°; WGS 84, uncertainty: 3200 m., GeoLocate, Georef'd by Maria Yousoufian Source: Digital Map, Georef'd by Joseph Stormer [Something is written for the habitat but it is unreadable]. Phenology: Cliff base, beside highway. Phenology: Flowers. Flowers. Origin: Introduced. H. T. Rogers 1005 28 Jun 1942 J. M. Grant s.n. May 1930 with J. M. Rogers WTU-42894 WTU-240877 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus laciniatus Willd. U.S.A., WASHINGTON, JEFFERSON COUNTY: U.S.A., OREGON, POLK COUNTY: Kelly Ranch, Queets River. Olympic forest. Along State Highway 99W, 2 miles south of Monmouth. Elev. 250 ft. Elev. 270 ft. 47.62333°, -124.0166°; WGS 84, uncertainty: 2500 m., Source: T9S R4W S6; NAD 27, uncertainty: 805 m., Source: TRS2LL, GeoLocate, Georef'd by WTU Staff Georef'd by Ben Legler Openings in burned over timbered area; bracken, vine maple A large bramble in ditch; associated genera: Holcus, association; heavy use; sandy. Common; forage value: low - eaten Symphoricarpos, Vicia, Rosa, Daucus, Crataegus, Parentucellia. to a slight extent by cattle and elk. Phenology: Flowers. Origin: Flowers white to pale pink. Phenology: Flowers. Origin: Introduced. Introduced. Richard R. Halse 2869 29 Jun 1984 Edward P. Cliff 104-C 8 Jun 1934 WTU-305370 WTU-322834

Rosaceae Rosaceae Rubus laciniatus Willd. Rubus parviflorus Nutt. U.S.A., OREGON, DOUGLAS COUNTY: U.S.A., MONTANA, MISSOULA COUNTY: Roseburg Quadrant. Foot of Spring Creek Trail, Rattlesnake Drainage. Escape from cultivation. Phenology: Fertile. Origin: Introduced. Phenology: Flowers & Fruits. Origin: Native. W. C. Cusick 4086 28 May 1914 C. L. Hitchcock 14598 9 Jul 1946 with C. V. Muhlick WTU-185976 WTU-105331 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus laciniatus Willd. U.S.A., WASHINGTON, KING COUNTY: South King County. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: 47.39361°, -122.42361°; WGS 84, uncertainty: 10000 m., Source: Humptulips, Olympic Peninsula. GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Origin: Introduced. Roadside. Phenology: Fruits. Origin: Introduced. Geo. N. Jones 3711 Aug 1931 Anita Watson 3 9 Aug 1949 WTU-21688 WTU-150080 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KING COUNTY: U.S.A., MONTANA, PARK COUNTY: Baring. 1 mile west of Four Mile Ranger Station; Elev. 700 ft. Boulder Canyon. 47.77333°, -121.48417°; WGS 84, uncertainty: 10000 m., Source: 45.340453°, -110.253937°; WGS 84, GeoLocate, Georef'd by WTU Staff uncertainty: 1000 m., Source: Digital Map, Phenology: Fertile. Origin: Introduced. Georef'd by Joseph Stormer In shade. Phenology: Flowers. F. S. Hall s.n. 15 Jul 1928 C. L. Hitchcock 16342 WTU-44393 12 Jul 1947 Rosaceae WTU-114551, WTU-V-045474 Rubus laciniatus Willd. U.S.A., WASHINGTON, MASON COUNTY: Rosaceae Waketichie [Waketickeh] Creek. 47.57606°, -123.05841°; WGS 84, uncertainty: 5500 m., Source: Rubus parviflorus Nutt. GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Origin: Introduced. U.S.A., IDAHO, LATAH COUNTY: North slope of Tomer's Butte, 4 miles southeast George Neville Jones 8499 5 Sep 1935 of Moscow. 46.691538°, -116.939429°; WGS 84, WTU-31537 uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff Woods. Phenology: Flowers. Origin: Native. Rosaceae William H. Baker 9009 Rubus parviflorus Nutt. 3 Jun 1952 U.S.A., IDAHO, IDAHO COUNTY: North Fork of Lake Fork, 9 miles east of McCall, WTU-154725, WTU-V-045476 Payette National Forest. 45.698334°, -116.051988°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Rosaceae Moist loam soil in Pinus-Pseudotsuga-Larix forest. Frequent. Phenology: Flowers. Origin: Rubus spectabilis Pursh Native. U.S.A., WASHINGTON, KING COUNTY: Arthur H. Holmgren 9587 Slopes of Interlaken Park, Seattle. Elev. 131 ft. 13 Jul 1953 T25N R4E S21; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler WTU-160684, WTU-V-045472 Forming thickets, shade of Acer macrophyllum. Pink flowers, 4-5 cm diameter, spiny shrubs, brown stems to 2.5 meters tall. Phenology: Flowers. Origin: Introduced. Rosaceae P. F. Zika 15903 Rubus laciniatus Willd. with A. L. Jacobson U.S.A., WASHINGTON, KING COUNTY: 17 Apr 2001 Cougar Mountain. 47.54056°, -122.09806°; WGS 84, uncertainty: 10000 m., Source: WTU-347257, WTU-V-18347 GeoLocate, Georef'd by WTU Staff Phenology: Fertile. Origin: Introduced. David M. Chapin s.n. 9 Oct 1982 Rosaceae WTU-314231 Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., IDAHO, BONNER COUNTY: [No locality given on label] Roadside. Phenology: Flowers. R. F. Daubenmire 44258 7 Jun 1944 WTU-238272 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., OREGON, KLAMATH COUNTY: U.S.A., MONTANA, LAKE COUNTY: Annie Creek at the south entrance to Crater St. Mary's Lake. Lake National Park. 48.698544°, -113.511001°; WGS 84, uncertainty: 10000 m., 42.768197°, -122.061518°; WGS 84, Source: Digital Map, Georef'd by Joseph Stormer, Invalid uncertainty: 500 m., Source: Digital Map, coordinates (located outside county bounds). Georef'd by Joseph Stormer Phenology: Flowers & Fruits. Growing along creek. Phenology: Flowers. Origin: Native. G. N. Jones 5641 23 Jul 1934 William H. Baker 6917 WTU-101586 7 Jul 1950 Rosaceae WTU-155834, WTU-V-045477 Rubus idaeus var. sachalinensis U.S.A., IDAHO, SHOSHONE COUNTY: Rosaceae Little Baldy Lookout. T50N R5E S8; WGS 84, uncertainty: 10000 m., Source: Rubus spectabilis Pursh GeoLocate, Georef'd by WTU Staff Talus slopes. Phenology: Fertile. U.S.A., WASHINGTON, KITTITAS COUNTY: Along Thorp Lake Road 4312, west of Lake Cle Elum. Jean G. Witt 1101 12 Aug 1946 Elev. 4000 ft. 47° 21.938' N, 121° 11.023' W; T22N R13E S26 WTU-119158 North-facing 20 degree slope, dirt w/rocks. Other plants: Alnus sinuata, Mertensia paniculata, Arnica latifolia, Valeriana sitchensis. Flowers dark pink; uncommon. Phenology: Flowers. Origin: Native. Rosaceae Don Knoke 453 24 Jun 2004 Rubus idaeus L. var. sachalinensis (H. Lév.) Focke WTU-357769 CANADA, YUKON TERRITORY: Canol Road: Mile 132; Lower Lapie River Crossing. Elev. 2750 ft. Rosaceae 61.90854°, -132.594276°; uncertainty: 3200 m., Georef'd by Joseph Stormer Rubus spectabilis Pursh Rich poplar woods. Obs. some leaflets 7-nate. Phenology: Fertile. CANADA, BRITISH COLUMBIA: A. E. Porsild 9838 22 Jun 1944 Alliford Bay area, Moresby Island, Queen Charlotte Islands. with A. J. Breitung In shade of alder and spruce along creek bank. Common; height 6'9'; flowers crimson. Voucher specimen cited in Flora of The Queen WTU-128622 Charlotte Islands. I. Systematics of The Vascular Plants by James A. Calder and Roy L. Taylor. Quqeen's Printer, Ottawa, 1968. Phenology: Flowers. Rosaceae J. A. Calder 21059 31 May 1957 Rubus spectabilis Pursh with D. B. O. Savile, R. L. Taylor U.S.A., ALASKA COUNTY: WTU-240343 Ketchikan lakes, Revillagigedo Islands. The yellow salmon-berry. Cf. notes on red specimen of like data. Berriese light orange-yellow. Phenology: Fruits. Origin: Native. Rosaceae T. T. McCabe 8583 25 Jul 1941 Rubus parviflorus Nutt. WTU-73968 U.S.A., OREGON COUNTY: Bingham Station, Blue Mountains. 45.744495°, -118.233018°; uncertainty: 1800 Rosaceae m., Georef'd by Joseph Stormer A common shrub. Phenology: Flowers. Origin: Rubus spectabilis Pursh Native. U.S.A., ALASKA COUNTY: Wm. C. Cusick 3239d Wrangell. 56.468454°, -132.370992°; WGS 84, uncertainty: 3000 m., 10 Jun 1908 Source: Digital Map, Georef'd by Joseph Stormer Hillsides. Phenology: Flowers. Origin: Native. WTU-142117, WTU-V-045479 Mr. Ernest P. Walker 611 2 May 1915 with Mrs. Ernest P. Walker WTU-76673 Rosaceae Rosaceae Rubus sachalinensis H. Lév. var. sachalinensis Rubus spectabilis Pursh U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Hyder, along road at border. Hillside, Akutan, Aleutian Island. With Rosa nutkana, Rubus parviflorus. Abundant. Phenology: [No collection date given]. Phenology: Flowers. Origin: Native. Fertile. Origin: Native. G. N. Jones 8923 Maxcine Williams 3325 22 Jun 1975 WTU-101510 WTU-263566 Rosaceae Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rubus spectabilis Pursh U.S.A., IDAHO, OWYHEE COUNTY: U.S.A., ALASKA COUNTY: Along the West Fork of Jarbidge River three miles northwest of Ketchikan lakes, Revillagigedo Islands. Murphy Hot Springs. Red Salmonberry. About equal in abudnance and growing alonside 42.249904°, -115.54885°; WGS 84, uncertainty: 10000 m., the yellow but not intergrading. Phenology: Fruits. Origin: Native. Source: GeoLocate, Georef'd by WTU Staff Thickets, rock slides. Frequent. Phenology: Fertile. T. T. McCabe 8589 25 Jul 1941 William H. Baker 8615 17 Aug 1951 WTU-73969 WTU-154727 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., ALASKA COUNTY: Port San Juan, Evans island. U.S.A., ALASKA COUNTY: 60.050833°, -148.0675°; uncertainty: 1200 m. Chomley [Chomly]. In wet hollow. Phenology: Flowers. Origin: Native. 55.247376°, -132.322482°; WGS 84, uncertainty: 2500 m., Source: Digital Map, Georef'd by Joseph Stormer W. J. Eyerdam 5781 23 May 1948 Phenology: Vegetative. Origin: Native. WTU-175194 G. B. Rigg 113 6 Aug 1929 WTU-39232 Rosaceae Rubus spectabilis Pursh Rosaceae U.S.A., ALASKA, VALDEZ CORDOVA COUNTY: Rubus spectabilis Pursh Middleton Island is located in the Gulf of Alaska about 115 miles south of Valdez. U.S.A., OREGON, WALLOWA COUNTY: 59° 25' N, 146° 20' W Near Forest Grove. Steep bluff, somewhat disurbed. Only a few flowers at this date. Fruit 45.52092°, -123.120308°; uncertainty: 3848 m., Georef'd by to 3 dm. long and to 2.5 cm. in diameter at base. Small shrubs to 4 Joseph Stormer, Invalid coordinates (located outside county feet tall. Phenology: Fruits. Origin: Native. bounds). Along creek bottoms. Phenology: Flowers. Origin: Native. John H. Thomas 6326 2 Aug 1956 J. William Thompson 639 29 Mar 1926 WTU-181777 WTU-23002JWT Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., ALASKA, ALEUTIANS WEST COUNTY: U.S.A., OREGON COUNTY: Unalaska. Chitwood. 53.583944°, -166.796107°; uncertainty: 68000 m. 44.65484°, -123.822612°; WGS 84, uncertainty: 3036 m., Source: Phenology: Fruits. Origin: Native. GeoLocate, Georef'd by Joseph Stormer Along streams. Phenology: Flowers. Origin: Native. Trevor Kincaid s.n. 18 Sep 1897 Helen Maw s.n. 12 May 1933 WTU-101450 WTU-72684 Rosaceae Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., IDAHO, OWYHEE COUNTY: U.S.A., IDAHO, BOISE COUNTY: West slope of Quicksilver Mountain, Silver City Range. Sawtooth Primitive Area; headwaters of South Fork Payette River 42.94778°, -116.63222°; WGS 84, uncertainty: 10000 m., Source: above Sacajawea Hot Springs; 5 miles north of Elk Lake. GeoLocate, Georef'd by WTU Staff 44.071634°, -115.750945°; WGS 84, uncertainty: 1000 m., Frequent, rock slides. Phenology: Fertile. Source: GeoLocate, Georef'd by WTU Staff Near stream. Branches pendent; plant glandular and tomentose. William H. Baker 8209 25 Jul 1951 Phenology: Fertile. WTU-154724 C. L. Hitchcock 9873 11 Jul 1944 with C. V. Muhlick Rosaceae WTU-91796 Rubus spectabilis Pursh Rosaceae U.S.A., ALASKA COUNTY: Port San Juan, Ewans Island. Rubus spectabilis Pursh 60.050833°, -148.0675°; uncertainty: 1000 m., Georef'd by Joseph Stormer U.S.A., ALASKA COUNTY: Forming thickets. Phenology: Flowers. Origin: Native. About 2 miles east of Sitka, near base of Mount Verstovia trail, in back of Kiksadi Club. W. J. Eyerdam 5793 Jun 1948 Phenology: Flowers. Origin: Native. WTU-175164 Mary Clay Muller 643 25 May 1975 WTU-274508 Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rosaceae U.S.A., OREGON, GRANT COUNTY: Rubus spectabilis Pursh On east slope of Fields Peak. Elev. 7100 ft. U.S.A., ALASKA COUNTY: T14S R29E S20, 21; NAD 27, uncertainty: 805 m., Source: Hydaburg. TRS2LL, Georef'd by Ben Legler Elev. 0 ft. Among diorite boulders on east slope of Fields Peak. Shrubs about 55.212366°, -132.806068°; WGS 84, uncertainty: 3500 m., 0.5 meters tall, petals white, shorter than the sepals. Phenology: Source: Digital Map, Georef'd by Joseph Stormer Flowers. Forest, muskeg, especially disturbed ground. A food plant. Phenology: Fruits. Origin: Native. Arthur Cronquist 7639 27 Jul 1953 Helen H. Norton 615-H 18 Feb 1979 WTU-157018 WTU-281171 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., OREGON, COOS COUNTY: Isthmus Slough. U.S.A., OREGON, COOS COUNTY: Elev. 25 ft. South of Charleston. T26S R13W S23; NAD 27, uncertainty: 805 m., Source: TRS2LL, T26S R14W; WGS 84, uncertainty: 7242 m., Source: Digital Map, Georef'd by Ben Legler Georef'd by Joseph Stormer Douglas fir forest; Pseudotsuga taxifolia, Tsuga heterophylla, Thuja Thickets and roadsides in logged and burned land. Weakly thorny plicata, Vaccinium parvifolium, V. ovatum; East hill slope, moderately shrub 1-3 meters tall; Flowers deep red-purple. Phenology: Flowers. moist clay-loam, in moderate shade. Phenology: Flowers. Origin: Origin: Native. Native. Arthur Cronquist 6851 15 Apr 1953 L. E. Detling 4099 21 Apr 1940 WTU-157001 WTU-74223 Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., IDAHO, OWYHEE COUNTY: Hot Hole, East Bruneau. 42.88056°, -115.79639°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Along the river. Phenology: Fertile. Aven Nelson 1907 3 Jul 1912 with J. Francis Macbride WTU-75184 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus pubescens Raf. U.S.A., OREGON, CLATSOP COUNTY: CANADA, BRITISH COLUMBIA: Astoria. Yale District: 10 miles east of Barriere. 46.182892°, -123.798981°; WGS 84, uncertainty: 5582 m., Plants trailing. Phenology: Fruits. Source: Digital Map, Georef'd by Joseph Stormer Phenology: Fruits. Origin: Native. C. L. Hitchcock 22062 30 Jun 1961 with C. V. Muhlick G. E. Morrill 36 17 Jun 1933 WTU-209303 WTU-26663 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus spectabilis Pursh U.S.A., IDAHO, LEMHI COUNTY: U.S.A., ALASKA COUNTY: Along Panther Creek, 10 miles north of Cabin North Tongass National Forest, 1 mile east of Juneau, Basin Road. Creek. Elev. 500 ft. 44.88008°, -113.979338°; WGS 84, 58.308074°, -134.389992°; WGS 84, uncertainty: 1309 m., uncertainty: 1000 m., Source: GeoLocate, Source: Digital Map, Georef'd by Joseph Stormer Georef'd by WTU Staff Cutover forest. Petals purplish; 3 foot high. Phenology: Fruits. Origin: Sandy bank. Phenology: Flowers. Origin: Native. Native. Elbert L. Little Jr. 18525 21 Jun 1961 C. L. Hitchcock 14276 with C. V. Muhlick WTU-250597 1 Jul 1946 Rosaceae WTU-105324, WTU-V-045478 Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rosaceae U.S.A., MONTANA, BEAVERHEAD COUNTY: At Pintlar Falls. Rubus pubescens Raf. On broken rocks. Phenology: Flowers. CANADA, BRITISH COLUMBIA: C. L. Hitchcock 12790 27 Jul 1945 Along a trail along the base of Sheep Mountain, west of the Fire with C. V. Muhlick Warden's Cabin in the Elk River Valley, 34 miles north of Natal. Clearings. Phenology: Fertile. WTU-98056 W. A. Weber 2341 7 Jul 1941 Rosaceae WTU-74512 Rubus pubescens Raf. Rosaceae U.S.A., WASHINGTON, STEVENS COUNTY: Pend Oreille Lakes. Rubus parviflorus Nutt. 48.6306°, -117.53388°; WGS 84, uncertainty: 3000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., IDAHO, IDAHO COUNTY: Phenology: Vegetative. Origin: Native. Lowell-Kooskia Road, 3 miles west of Lowell. T32N R6E S12; WGS 84, uncertainty: 1000 H. St. John 3277 26 Jun 1924 m., Source: GeoLocate, Georef'd by WTU Staff WTU-112980 In moist ravines in Thuja-Pseudotsuga association. Shrub to 2 meters tall. Phenology: Flowers. Origin: Native. Rosaceae Quentin Jones 53 Rubus idaeus L. var. sachalinensis (H. Lév.) Focke 17 Jun 1950 U.S.A., MONTANA, MEAGHER COUNTY: Along Checkerboard Creek, northeast base of Castle Mountains. WTU-142794, WTU-V-045480 46.533425°, -110.651042°; uncertainty: 9000 m. In shade of lodgepole pine. Phenology: Flowers. C. L. Hitchcock 12135 8 Jul 1945 WTU-102083 Rosaceae Rosaceae Rubus pubescens Raf. Rubus pedatus Sm. CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, OKANOGAN COUNTY: Ten miles on road to Horsefly from 153 Mile House. West of Salmon Meadows, Tiffany Range. Elev. 3300 ft. Elev. 4000 ft. On wet mossy hummocks in spruce swamp. Common; flowers white. 48.65556°, -119.83806°; WGS 84, uncertainty: 10000 m., Source: Phenology: Fertile. GeoLocate, Georef'd by WTU Staff In timber where small spring kept ground wet. Phenology: Flowers. J. A. Calder 19037 17 Jul 1956 Origin: Native. with J. A. Parmelee, R. L. Taylor Chas. B. Fiker 211 4 Jul 1931 WTU-195937 WTU-23013JWT Rosaceae Rosaceae Rubus pubescens Raf. Rubus pedatus Sm. CANADA, BRITISH COLUMBIA: Germansen Landing, Omineca River. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: 55.783333°, -124.7°; WGS 84, uncertainty: 3036 m., Source: Mount Colonel Bob Lookout, Olympic Mountains. GeoLocate, Georef'd by Joseph Stormer Elev. 4000 ft. Thin aspen, spruce, pine woods, moderately dry. Flowers white. 47.48889°, -123.74556°; WGS 84, uncertainty: 10000 m., Source: Phenology: Flowers. GeoLocate, Georef'd by WTU Staff Moist wooded slopes. Phenology: Flowers. Origin: Native. T. T. McCabe 7788 7 Jul 1940 J. William Thompson 7269 9 Jul 1931 WTU-72187 WTU-23008JWT Rosaceae Rosaceae Rubus pubescens Raf. Rubus parviflorus Nutt. CANADA, BRITISH COLUMBIA: Kinbasket Lake (Big Bend). U.S.A., OREGON, GRANT COUNTY: White silt flats under open small Pinus. Phenology: Flowers. [Digital specimen data says "Washington", but this is in Oregon] Near the bottom of Black T. T. McCabe 6292 24 May 1938 Canyon, 12 miles south of Dayville. Elev. 3100 ft. WTU-71792 T14S R26E S22 or 27; WGS 84, uncertainty: 2100 m., Source: Digital Map, Georef'd by Joseph Stormer Rosaceae Among conifers. 0.5-1 meters tall. Phenology: Flowers. Origin: Native. Rubus parviflorus Nutt. Arthur Cronquist 7601 U.S.A., MONTANA, LINCOLN COUNTY: Libby, roadside east of Libby. 24 Jul 1953 Phenology: Flowers. Origin: Native. WTU-157016, WTU-V-045434 F. B. Cotner s.n. with W. E. Booth 20 Jul 1952 Rosaceae WTU-146578, WTU-V-045481 Rubus pedatus Sm. U.S.A., WASHINGTON, JEFFERSON COUNTY: North Fork Quinault, .5 miles above north fork R.S. Elev. 1000 ft. 47.57571°, -123.64797°; WGS 84, uncertainty: 500 m., Source: Rosaceae GeoLocate, Georef'd by WTU Staff Side of road; moist thin pumus on rock, Salmonberry, huckleberry, Rubus pubescens Raf. weed association; deep forests, rocks and logs. Common; practically worthless. Phenology: Flowers. Origin: Native. CANADA, BRITISH COLUMBIA: Clearwater River Road, 20 miles from North Thompson River Road. Edward P. Cliff 126C 19 Jul 1925 Burn--open. Phenology: Flowers. WTU-322836 T. T. McCabe 1034 11 May 1934 WTU-36213 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus pedatus Sm. U.S.A., IDAHO, IDAHO COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Clearwater National Forest, Badger Creek, Lochsa River Canyon, Canyon Creek. about 40 miles southwest of Missoula. Elev. 3000 ft. Elev. 3900 ft. 48.00752°, -124.093917°; WGS 84, uncertainty: 10000 m., 46.504525°, -114.822919°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Source: GeoLocate, Georef'd by WTU Staff Hemlock forest. Phenology: Vegetative. Origin: Native. Along logging road in clear-cut. Infrequent. Phenology: Fruits. Origin: Native. G. N. Jones 5907 10 Aug 1934 Peter F. Stickney 853 26 Jul 1963 WTU-36099 WTU-234555 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus pedatus Sm. CANADA, BRITISH COLUMBIA: Goldstream, Vancouver Island. U.S.A., WASHINGTON, CHELAN COUNTY: Phenology: Flowers. Wenatchee Forest. Elev. 3950 ft. C. F. Newcombe s.n. 8 Jun 1974 T26N R15E S34; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler WTU-168983 Moist decomposed vegetation; heavy timber type. Fairly plentiful in immediate vicinity; June-July flowering period; Forage value not known. Phenology: Flowers. Origin: Native. Rosaceae Frank B. Lenzie 370 28 Jun 1918 Rubus pedatus Sm. WTU-322293 U.S.A., WASHINGTON, WHATCOM COUNTY: Baker Lake region. 48.72917°, -121.62778°; WGS 84, uncertainty: 10000 m., Source: Rosaceae GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Origin: Native. Rubus parviflorus Nutt. H. L. Mason 7421 24 Jul 1933 U.S.A., IDAHO, ADAMS COUNTY: Along Middle Fork of Weiser River Road. WTU-72740 Elev. 4592 ft. 44° 8' N, 116° 18' W; T15N R1E S11; WGS 84, uncertainty: 3200 m., Source: Nat. Geog. Rosaceae TOPO!, Georef'd by John Bassett Hills north of river, steep slope with clay and Rubus parviflorus Nutt. rock, by stream. To 1 meter. Phenology: Flowers. Origin: Native. U.S.A., ALASKA COUNTY: Tongas Village. James F. Smith 2802 Clearing at beach. Phenology: Fruits. Origin: Native. 22 May 1994 Mr. Ernest P. Walker 891 WTU-330735, WTU-V-045436 3 Aug 1915 WTU-76669, WTU-V-045439 Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., IDAHO, IDAHO COUNTY: Along middle fork of Sheep Creek, above the forks, Snake River Rosaceae Canyon. Elev. 2500 ft. Rubus leucodermis Douglas ex Torr. & A. Gray T25N R2W; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., OREGON, KLAMATH COUNTY: Trail, in moist copses. Phenology: Flowers. Origin: Native. Southwest corner, north wall, Red Blanket Canyon. Crater Lake National Park. Robt. Beaubier 63 16 May 1936 42.784045°, -122.367427°; WGS 84, uncertainty: 2500 m., Source: Digital Map, Georef'd by Joseph Stormer, Invalid WTU-9008JWT coordinates (located outside county bounds). Phenology: Flowers. Origin: Native. William H. Baker 7069 29 Jul 1950 WTU-155833 Rosaceae Rosaceae Rubus pedatus Sm. Rubus parviflorus Nutt. U.S.A., WASHINGTON, JEFFERSON COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Near Bogachiel State Park. Mount Dickerman, Cascade Mountains. 47.88°, -124.24167°; WGS 84, uncertainty: 5000 m., Source: Elev. 4500 ft. GeoLocate, Georef'd by WTU Staff 48.06829°, -121.47068°; WGS 84, uncertainty: 3000 m., Source: In swampy area in deep shade; woods. Pollen removed for GeoLocate, Georef'd by WTU Staff reference collection Linda Brubaker UW College of Forest Moist banks. Phenology: Flowers. Origin: Native. Resources February 1995. Phenology: Flowers. Origin: Native. J. William Thompson 8871 17 Jul 1932 C. L. Hitchcock 3527 29 May 1938 with J. S. Martin WTU-23010JWT-B WTU-45037 Rosaceae Rosaceae Rubus pedatus Sm. Rubus nigerrimus (Greene) Rydb. U.S.A., WASHINGTON, JEFFERSON COUNTY: On bench below Jackson G.S., Hoh River. U.S.A., WASHINGTON, WHITMAN COUNTY: Elev. 600 ft. [locality and coordinates withheld; status in Washington: G2/S2 47.814011°, -124.176698°; WGS 84, uncertainty: 10000 m., (Threatened; WANHP)] Source: GeoLocate, Georef'd by WTU Staff Along small stream. Plants erect then the branches arching or Slope south 15%, gravel soil; Oxalis Tiarella association; On clambering, up to 8 feet long, new shoots glaucous and only slightly bottoms and slopes. Plant trailing; July and August flowering period. paler on the lower, than on the upper surface, spines flattened and Phenology: Flowers. Origin: Native. mostly hooked, petals white elliptic, flared, 5, but many flowers with 6, and occasional flowers with up to 10 to 13 petals, in the whole not J. E. Schwartz 3A 1 Aug 1935 quuite so glaucous, especially on protected part of plant. Phenology: Flowers. WTU-94647 C. L. Hitchcock 21861 11 Jun 1959 with C. V. Muhlick Rosaceae WTU-197466 Rubus nivalis Douglas ex Hook. U.S.A., WASHINGTON, PIERCE COUNTY: Rosaceae Mishell Mountain. 47.00675°, -122.20434°; WGS 84, Rubus pedatus Sm. uncertainty: 62000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., WASHINGTON, CLALLAM COUNTY: Phenology: Flowers & Fruits. Origin: Native. Boulder Creek. 47.98167°, -123.60139°; WGS 84, uncertainty: 10000 m., Source: C. V. Piper 710 GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. 7 Aug 1889 George Neville Jones 8480 6 Sep 1935 WTU-9557, WTU-V-18513 WTU-31539 Rosaceae Rosaceae Rubus nivalis Douglas ex Hook. Rubus pedatus Sm. U.S.A., WASHINGTON, LEWIS COUNTY: U.S.A., WASHINGTON COUNTY: Storm King Mountain. North fork Brindger Creek. 46.64861°, -122.15722°; WGS 84, Phenology: Flowers. Origin: Native. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff C. V. Piper 697 Aug 1897 Woods. Phenology: Flowers. Origin: Native. WTU-20546 Harold E. Helmrich 40 28 Jun 1932 Rosaceae WTU-306035, WTU-V-18502 Rubus pedatus Sm. U.S.A., WASHINGTON COUNTY: Elwha Valley, Olympic Mountains. Phenology: Flowers. Origin: Native. J. B. Flett s.n. 20 Aug 1907 WTU-32287 Rosaceae Rosaceae Rubus pedatus Sm. Rubus nivalis Douglas ex Hook. U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Moranie Boulder Glacier, Mount Baker. Longmire Springs, Mount Rainier National Park. 48.761552°, -121.781368°; WGS 84, uncertainty: 2000 m., Elev. 2000 ft. Source: Digital Map, Georef'd by Ibro Polovina 46.75024°, -121.812778°; WGS 84, Phenology: Flowers. Origin: Native. uncertainty: 1500 m., Source: GeoLocate, Georef'd by Joseph Stormer W. Bailey s.n. 28 Jul 1908 By trail thru timber. Phenology: Flowers. Origin: Native. WTU-272414 A. S. Foster 1096 Rosaceae 12 Aug 1909 Rubus nivalis Douglas ex Hook. WTU-9785, WTU-V-18503 U.S.A., WASHINGTON, WHATCOM COUNTY: Skyline Ridge, Mount Baker. Elev. 3000 ft. Rosaceae 48.83065°, -121.85005°; WGS 84, uncertainty: 750 m., Source: Nat. Geog. Rubus pedatus Sm. TOPO!, Georef'd by John Bassett Lower wooded slopes. Phenology: Flowers. U.S.A., WASHINGTON, KING COUNTY: Origin: Native. Goldmeyer Hot Springs. Elev. 1800 ft. J. William Thompson 10972 47.205°, -121.54528°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff 11 Jul 1934 Deep woods. Phenology: Flowers. Origin: Native. WTU-15700JWT, WTU-V-18504 John Broadbent s.n. 23 Jun 1935 WTU-35117 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, ISLAND COUNTY: Mount Dickerman, Cascade Mountains. Deception Pass State Park, Whidbey Island. Elev. 4500 ft. 48.39389°, -122.6475°; WGS 84, uncertainty: 48.06829°, -121.47068°; WGS 84, 10000 m., Source: GeoLocate, Georef'd by uncertainty: 3000 m., Source: GeoLocate, WTU Staff Georef'd by WTU Staff [4-10-35 written on collection sheet in pencil]. Moist talus slopes. Phenology: Flowers. Origin: Phenology: Flowers. Origin: Native. Native. Harold W. Smith 52 J. William Thompson 8862 25 Mar 1935 17 Jul 1932 WTU-38889, WTU-V-044985 WTU-23012JWT, WTU-V-044973

Rosaceae Rosaceae Rubus pedatus Sm. Rubus nivalis Douglas ex Hook. U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: 1 mile out from Big 4 Inn. Near Longmire, Mount Rainier. Elev. 1800 ft. 46.749831°, -121.812597°; WGS 84, 48.06765°, -121.515°; WGS 84, uncertainty: 1800 m., Source: uncertainty: 3000 m., Source: GeoLocate, GeoLocate, Georef'd by WTU Staff Georef'd by Joseph Stormer Semi-shade, base of old stump. Phenology: Flowers. Origin: Native. Phenology: Vegetative. Origin: Native. Ruby L. C. Sovereign 121 26 May 1940 G. N. Jones 9660 WTU-315751 27 Sep 1936 WTU-101587, WTU-V-18506 Rosaceae Rosaceae Rubus pedatus Sm. Rubus nivalis Douglas ex Hook. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: In Snoqualmie Pass, Cascade Mountains. Mount Angeles. Elev. 3100 ft. 48.00365°, -123.445867°; WGS 84, 47.61028°, -121.42556°; WGS 84, uncertainty: 10000 m., Source: uncertainty: 5330 m., Source: Digital Map, GeoLocate, Georef'd by WTU Staff Georef'd by Joseph Stormer Moist woods. Phenology: Fertile. Origin: Native. Phenology: Fertile. Origin: Native. J. William Thompson 9696 7 Aug 1933 E. B. Webster 1622 WTU-23014JWT Aug 1911 WTU-32207, WTU-V-18501 Rosaceae Rubus pedatus Sm. Rosaceae U.S.A., WASHINGTON, KING COUNTY: Tye River valley below Scenic. Rubus pedatus Sm. Elev. 1700 ft. 47.710726°, -121.153856°; WGS 84, uncertainty: 10000 m., U.S.A., WASHINGTON, PIERCE COUNTY: Source: GeoLocate, Georef'd by WTU Staff Seattle, Mount Rainier. Phenology: Flowers. Origin: Native. Phenology: Fertile. Origin: Native. I. C. Otis 919 12 Jun 1920 C.V.P. s.n. Aug 1889 WTU-15721JWT WTU-9560

Rosaceae Rosaceae Rubus nivalis Douglas ex Hook. Rubus spectabilis Pursh U.S.A., WASHINGTON, CLALLAM COUNTY: U.S.A., WASHINGTON, WHATCOM COUNTY: Mount Angeles. Above Bagley Lake, Baker Lodge. 48.00365°, -123.445867°; WGS 84, Elev. 4400 ft. uncertainty: 5330 m., Source: Digital Map, 48.85675°, -121.68675°; WGS 84, Georef'd by Joseph Stormer uncertainty: 575 m., Source: Nat. Geog. Phenology: Fertile. Origin: Native. TOPO!, Georef'd by John Bassett Moist slopes. Phenology: Flowers & Fruits. E. B. Webster 1621 Origin: Native. 1910 J. William Thompson 5362 WTU-32285, WTU-V-18505 25 Jul 1930 WTU-23017JWT, WTU-V-044984 Rosaceae Rubus pedatus Sm. Rosaceae U.S.A., WASHINGTON, PIERCE COUNTY: Rubus nivalis Douglas ex Hook. Mount Rainier. 46.852827°, -121.760441°; WGS 84, uncertainty: 17500 m., U.S.A., OREGON, WASHINGTON COUNTY: Source: GeoLocate, Georef'd by Joseph Stormer Eastern slope of Coast Mountain, west of Phenology: Fertile. Origin: Native. Banks. Eastern slope. Phenology: Vegetative. Origin: C.V.P. s.n. Aug 1888 Native. WTU-9559 J. William Thompson 3068 20 Jul 1927 WTU-22956JWT-A, WTU-V-18500 Rosaceae Rosaceae Rubus pedatus Sm. Rubus spectabilis Pursh U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, JEFFERSON Silverton. COUNTY: 48.07861°, -121.56583°; WGS 84, uncertainty: 10000 m., Source: Hoods Canal near Brinnon. GeoLocate, Georef'd by WTU Staff 47.67944°, -122.89694°; WGS 84, Phenology: Flowers. Origin: Native. uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff L. S. Bouck s.n. 18 May 1895 Moist slopes facing. Phenology: Flowers. Origin: Native. WTU-9848 J. William Thompson 5937 Rosaceae 31 Mar 1931 Rubus nivalis Douglas ex Hook. WTU-23018JWT, WTU-V-044983 U.S.A., OREGON COUNTY: Coast Mountains near Roseburg. Rosaceae Phenology: Fertile. Origin: Native. Rubus pedatus Sm. Thomas Howell s.n. 23 Jun 1887 U.S.A., WASHINGTON, JEFFERSON COUNTY: WTU-22956JWT-B Below Jackson Guard Station, Hoh River, Olympic National Forest. 47.814011°, -124.176698°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Rosaceae Phenology: Flowers. Origin: Native. Rubus ursinus Cham. & Schltdl. J. E. Schwartz 3A 1 Aug 1935 U.S.A., WASHINGTON, CLALLAM COUNTY: WTU-33227 La Push. Wild Blackberry. No collection date given. Phenology: Flowers. Origin: Native. Rosaceae R. Ernest E2135 Rubus ursinus Cham. & Schltdl. WTU-316470 U.S.A., WASHINGTON, KING COUNTY: Three Tree Point, Puget Sound. 47.45056°, -122.38111°; WGS 84, uncertainty: 10000 m., Source: Rosaceae GeoLocate, Georef'd by WTU Staff Cliffside. Phenology: Flowers. Origin: Native. Rubus pedatus Sm. Lyman Benson 1330 10 May 1929 U.S.A., WASHINGTON COUNTY: Along Elwha River one mile below mountaineer's Camp. WTU-39425 On ground in woods. Phenology: Fruits. Origin: Native. T. C. Frye s.n. 18 Aug 1907 Rosaceae WTU-9802 Rubus nivalis Douglas ex Hook. U.S.A., OREGON, KLAMATH COUNTY: Rosaceae 20 miles southwest of Headquarters, at Crater Lake. Rubus nivalis Douglas ex Hook. Elev. 4500 ft. In Pseudotsuga forest, open areas. Phenology: U.S.A., OREGON, JOSEPHINE COUNTY: Flowers. Origin: Native. Oregon Caves National Monument and vicinity. Along the Grayback trail. C. L. Hitchcock 4961 Phenology: Flowers. Origin: Native. with J. S. Martin William H. Baker 125 27 Jun 1939 with George C. Ruhle WTU-53481, WTU-V-18499 29 Jun 1949 WTU-274594, WTU-V-18508 Rosaceae Rosaceae Rubus pedatus Sm. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., OREGON, JACKSON COUNTY: Mount Baker. Mount Ashland. 48.777343°, -121.813201°; WGS 84, uncertainty: 10000 m., 42.080833°, -122.717222°; uncertainty: 2400 m., Georef'd by Source: GeoLocate, Georef'd by Ibro Polovina Joseph Stormer Mountan Meadow. Phenology: Flowers. Origin: Native. Middle slope. Phenology: Flowers. Origin: Native. H. W. Bailey s.n. 28 Jul 1908 Morton E. Peck 7049 19 Jul 1913 WTU-9831 WTU-22954JWT-A

Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus pedatus Sm. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: McCleary. Steven's Pass. 47.05333°, -123.26417°; WGS 84, uncertainty: 10000 m., Source: Elev. 4000-6000 ft. GeoLocate, Georef'd by WTU Staff 47.744438°, -121.090111°; WGS84, uncertainty: 600 m., Source: Phenology: Flowers. Origin: Native. Georeferenced, Georef'd by WTU Staff Herbarium of the Grand Rapids Public Museum Deposited in the G. N. Jones 4592 7 Apr 1934 Herbarium of the University of Michigan, 1974. Catalogue Number: 50810. In 1867 the Grand Rapids Lyceum of Natural History merged WTU-15804JWT-A with the Grand Rapids Scientific Cluf to form the Kent Scientific Institute. In 1902 the name was changed to Kent Scientific Musum, and in 1936 to Grand Rapids Publis Museum. Catalogue numbers Rosaceae were assigned to the botanical collections during the period when the insitution was known as the Kent Scientific Museum. Phenology: Rubus spectabilis Pursh Flowers. Origin: Native. U.S.A., WASHINGTON, ISLAND COUNTY: J. H. Sandberg 2042 Aug 1893 Near Langley, Whidby [Whidbey] Island. with J. B. Leiberg 48.04028°, -122.405°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU-269488 WTU Staff Wooded stream. Phenology: Flowers. Origin: Native. Rosaceae Lyman Benson 1346 Rubus ursinus Cham. & Schltdl. 11 May 1929 U.S.A., WASHINGTON, SNOHOMISH COUNTY: On Mount Dickerman, Cascade Mountains. WTU-40682, WTU-V-044980 Elev. 5000 ft. 48.06829°, -121.47068°; WGS 84, uncertainty: 3000 m., Source: GeoLocate, Georef'd by WTU Staff Rosaceae Moist talus slopes. Phenology: Flowers. Origin: Native. Rubus ursinus Cham. & Schltdl. J. William Thompson 8869 17 Jul 1922 U.S.A., OREGON, CLACKAMAS COUNTY: WTU-22954JWT-B Banks of Eagle Creek near Estacada. Trailing on logs. Phenology: Flowers. Origin: Native. Rosaceae J. William Thompson 4251 20 May 1928 Rubus ursinus Cham. & Schltdl. WTU-15804JWT-B U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Vicinity Burnt Hill. Rosaceae Elev. 550 ft. 47.32583°, -123.865°; WGS 84, uncertainty: 10000 m., Source: Rubus pedatus Sm. GeoLocate, Georef'd by WTU Staff Level slope; cutover fir. Common; fair palatability; grazed by elk and U.S.A., WASHINGTON COUNTY: deer during winter and spring. The words on the label were blurred. Olympic Mountains, between mouth of Bodbrin Creek and Phenology: Flowers. Origin: Native. mountaineer's Camp, in valley of Elwha River. Phenology: Fruits. Origin: Native. J. E. Schwartz 48A 3 Jun J. B. Flett s.n. 20 Aug 1907 WTU-94653 WTU-9709 Rosaceae Rosaceae Rubus pedatus Sm. Rubus pedatus Sm. U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Steven's Pass. Mount Tacoma. Elev. 4000-6000 ft. Elev. 4000 ft. 47.744438°, -121.090111°; WGS84, uncertainty: 600 m., Source: Indefinite loan by Tacoma High School to University of Washington. Georeferenced, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. J. B. Flett s.n. 22 Aug 1895 J. H. Sandberg s.n. Aug 1893 with J. B. Leiberg WTU-33511 WTU-9520 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus spectabilis Pursh U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Olympic Forest; Humptulips C.C.C. Camp. U.S.A., WASHINGTON, SNOHOMISH Elev. 400 ft. COUNTY: 47.2329°, -123.95902°; WGS 84, uncertainty: 1500 m., Source: King (205th and Alaska Way)/Snohomish GeoLocate, Georef'd by WTU Staff Counties. Undeveloped property on the north Flat slope; deep sandy loam soil; cutover Douglas fir; associated border of LFP, 10 yards in from 205th. with Huckleberry, grass; cutover areas and timber. Common; good 47.91306°, -122.09694°; WGS 84, for cattle and sheep; eaten readily by deer and elk; big gaze uncertainty: 10000 m., Source: GeoLocate, animals; bear, deer, elk and gume; birds seek out the berries. Georef'd by WTU Staff Phenology: Flowers & Fruits. Origin: Native. Along side trail with other Rubus, Veronica hederaefolia and Circaea alpina. Phenology: Edward P. Cliff 109-c 21 Sep 1954 Flowers. Origin: Native. WTU-91412 Christie S. Isler 1-4-98 26 Apr 1998 Rosaceae WTU-335298, WTU-V-044982 Rubus pedatus Sm. U.S.A., WASHINGTON, WHATCOM COUNTY: Rosaceae Razor Hone Creek, Mount Baker National Forest. 48.87161°, -121.65965°; WGS 84, uncertainty: 2600 m., Source: Rubus ursinus Cham. & Schltdl. Nat. Geog. TOPO!, Georef'd by John Bassett Woodland banks. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: West of Scenic. Bassett Maguire 17273 14 Jul 1939 Elev. 1900 ft. 47.71167°, -121.15444°; WGS 84, uncertainty: 10000 m., Source: WTU-22035JWT GeoLocate, Georef'd by WTU Staff Along highway. Phenology: Flowers. Origin: Native. Rosaceae I. C. Otis 922 16 Jun 1920 Rubus spectabilis Pursh WTU-15920JWT U.S.A., WASHINGTON, SNOHOMISH COUNTY: Rosaceae Big Four Inn, Cascade Mountains. Elev. 1900 ft. Rubus pedatus Sm. 48.06611°, -121.50778°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, U.S.A., WASHINGTON, SKAMANIA COUNTY: Georef'd by WTU Staff Yie Creek. Deep woods. Phenology: Flowers. Origin: 45.96185°, -121.63356°; WGS 84, uncertainty: 500 m., Source: Native. GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Origin: Native. J. William Thompson 14534 Wilhelm N. Suksdorf s.n. 15 Aug 1900 18 May 1940 WTU-197549 WTU-21630JWT, WTU-V-044981 Rosaceae Rosaceae Rubus pedatus Sm. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., WASHINGTON, ISLAND COUNTY: Nevada Falls, Mount Rainier. Coupeville, Whidbey Island. 46.774879°, -121.746179°; uncertainty: 500 m., Georef'd by 48.219821°, -122.68628°; WGS 84, uncertainty: 3000 m., Source: Joseph Stormer, Invalid coordinates (located outside county GeoLocate, Georef'd by Joseph Stormer bounds). Woods. Phenology: Flowers. Origin: Native. Coniferous woods. Indefinite loan by Tacoma High School to University of Washington. Phenology: Flowers. Origin: Native. George Neville Jones 4835 19 May 1934 O. D. Allen 280 25 Aug 1897 WTU-42559 WTU-33512 Rosaceae Rosaceae Rubus pedatus Sm. Rubus ursinus Cham. & Schltdl. CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Wapta. Marysville. Not found in Herb. MBG. Phenology: Fertile. 48.05194°, -122.17583°; WGS 84, uncertainty: 3570 m., Source: GeoLocate, Georef'd by Maria Yousoufian unknown G-3 Jul 1927 Phenology: Flowers. Origin: Native. WTU-329457 J. M. Grant s.n. May 1929 WTU-42903 Rosaceae Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, SNOHOMISH COUNTY: Rubus arcticus L. Silverton. 48.07844°, -121.567062°; WGS 84, uncertainty: 5000 m., Source: CANADA, BRITISH COLUMBIA: GeoLocate, Georef'd by Joseph Stormer Little Yoho River. Running blackberry. Phenology: Flowers & Fruits. Origin: Native. Field. Herb MBG. Gray's Bot. Phenology: Fruits. L. S. Bouch s.n. 1 Jun 1895 unknown 1720 7 Aug 1957 WTU-9849 WTU-329147 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, CLALLAM COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR Canyon Creek. COUNTY: Elev. 3500 ft. Near Taholah. 48.00752°, -124.093917°; WGS 84, 47.3475°, -124.29194°; WGS 84, uncertainty: uncertainty: 10000 m., Source: GeoLocate, 5000 m., Source: GeoLocate, Georef'd by Georef'd by WTU Staff WTU Staff Along stream. Phenology: Fruits. Origin: Native. On bluffs along the ocean. Abundant. Phenology: Flowers & Fruits. Origin: Native. G. N. Jones 5832 George Neville Jones 6488 10 Aug 1934 11 May 1935 WTU-36211, WTU-V-044979 WTU-29195, WTU-V-044977 Rosaceae Rosaceae Rubus pedatus Sm. Rubus ursinus Cham. & Schltdl. CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, SAN JUAN COUNTY: Little Yoho River. San Juan Island. Field. Herb. MBG. Phenology: Flowers. 48.536386°, -123.071596°; WGS 84, uncertainty: 13000 m., Source: Digital Map, Georef'd by Joseph Stormer unknown 1720 7 Aug 1957 Northeast. Phenology: Flowers. Origin: Native. WTU-329402 A. S. Pope s.n. 5 Jul 1904 WTU-9538 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Ocean City. Near American Lake. 47.07111°, -124.16472°; WGS 84, uncertainty: 10000 m., Source: 47.1325°, -122.56361°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Woods. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. G. N. Jones 3898 27 Jun 1933 George Neville Jones 4602 29 Apr 1934 WTU-36217 WTU-15867JWT

Rosaceae Rosaceae Rubus pedatus Sm. Rubus ursinus Cham. & Schltdl. CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, ISLAND COUNTY: 21 Mile Creek Trail, Alta Lake. Cranberry lake, Deception Pass State Park, Whidbey Island. Phenology: Flowers. 48.14389°, -122.49111°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff unknown 5498 6 Aug 1967 Phenology: Flowers. Origin: Native. WTU-329146 Harold W. Smith 482 11 May 1936 WTU-40542 Rosaceae Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, KLICKITAT COUNTY: Rubus pedatus Sm. Bingen. 45.71495°, -121.46462°; WGS 84, uncertainty: 500 m., Source: U.S.A., ALASKA, JUNEAU COUNTY: GeoLocate, Georef'd by WTU Staff Juneau. Garden. Phenology: Flowers & Fruits. Origin: Native. Cultivated. 58.301944°, -134.419722°; uncertainty: 10000 m. Phenology: Flowers. Origin: Native. Wilhelm N. Suksdorf 8407 1 May 1905 Emma A. Shumway s.n. 24 Jul 1891 WTU-196417 WTU-112693 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus pedatus Sm. U.S.A., WASHINGTON, KING COUNTY: Paradise Lake. CANADA, BRITISH COLUMBIA: 47.77361°, -122.0675°; WGS 84, uncertainty: 9 - Mile Mountain (northeast of Hazelton). 10000 m., Source: GeoLocate, Georef'd by 55.334948°, -127.491326°; WGS 84, uncertainty: 5000 m., WTU Staff Source: Digital Map, Georef'd by Joseph Stormer Erect shrub, flowers purplish red Paradise Lake. High alpine pastures; often under Abies scrub and through (Puget Sound Botanical Club number 174.). Empetrum. Flowers white. Phenology: Flowers. Phenology: Flowers & Fruits. Origin: Native. T. T. McCabe 8210 5 Aug 1940 Ora P. Quense 74 WTU-8210 25 May 1940 WTU-315750, WTU-V-044978 Rosaceae Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, ISLAND COUNTY: Near Coupeville, Whidbey Island. Rubus pedatus Sm. 48.22°, -122.685°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff CANADA, BRITISH COLUMBIA: Woods. Phenology: Flowers. Origin: Native. 3 miles WestNorthwest of Tyee along road between Prince Rupert and Terrace. George Neville Jones 4835 19 May 1934 On mossy banks in dense coniferous corest; fruit red. Common. Phenology: Fertile. WTU-34937 J. A. Calder 15041 26 Aug 1954 with D. B. O. Savile, J. M. Ferguson WTU-172932 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus pedatus Sm. U.S.A., WASHINGTON, SNOHOMISH CANADA, BRITISH COLUMBIA: COUNTY: Ten miles north of Prince George along Hart Highway. Silverton. Very common in dense coniferous woods. Flowers white. 48.07844°, -121.567062°; WGS 84, Phenology: Flowers. uncertainty: 5000 m., Source: GeoLocate, Georef'd by Joseph Stormer J. A. Calder 12415 3 Jul 1954 Phenology: Flowers. Origin: Native. with D. B. O. Savile, J. M. Ferguson L. S. Bouck s.n. WTU-172933 7 May 1895 Rosaceae WTU-9846, WTU-V-18345 Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON COUNTY: Olympic Peninsula; Elwha River. Rubus ursinus Cham. & Schltdl. Elev. 1000 ft. Woods. Phenology: Flowers & Fruits. Origin: Native. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Near Ocean City. Geo. N. Jones 3242 Jul 1931 47.07111°, -124.16472°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-21311 Woods. Phenology: Flowers. Origin: Native. G. N. Jones 3935 28 Jun 1933 Rosaceae WTU-36218 Rubus pedatus Sm. CANADA, BRITISH COLUMBIA: Rosaceae Bennett, south end of Lake Bennett. 59.84931°, -134.985393°; uncertainty: 3000 m. Rubus pedatus Sm. Phenology: Fruits. CANADA, BRITISH COLUMBIA: A. A. Heller s.n. 8 Jul 1929 Takakkaw Falls, Yoho National Park. Wooded bank. Phenology: Flowers. WTU-87307 W. C. McCalla 7696 24 Jul 1943 Rosaceae WTU-182011 Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, COWLITZ COUNTY: Small valley; 5 miles west of Kelso. Rubus ursinus Cham. & Schltdl. 46.146892°, -123.01171°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Georef'd by WTU Staff Ocean City. In ravine under Pseudotsugs [Pseudotsuga]. 47.07111°, -124.16472°; WGS 84, uncertainty: 10000 m., Source: Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Woods. Phenology: Flowers. Origin: Native. C. L. Hitchcock 3276 G. N. Jones 3898 27 Jun 1933 24 Apr 1938 WTU-25418 WTU-45517, WTU-V-044972

Rosaceae Rosaceae Rubus spectabilis Pursh Rubus pedatus Sm. U.S.A., WASHINGTON, SAN JUAN COUNTY: Friday Harbor. U.S.A., ALASKA COUNTY: 48.534266°, -123.017124°; WGS 84, Hydaburg, Prince of Wales Island. uncertainty: 5000 m., Source: GeoLocate, Elev. 0 ft. Georef'd by Joseph Stormer Muskeg and its edges. A food plant. Phenology: Flowers. Origin: [No collection date given on label]. Phenology: Native. Fruits. Origin: Native. Helen H. Norton 601-H 18 Jun 1979 A. S. Pope s.n. WTU-281071 WTU-9539, WTU-V-044976 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus pedatus Sm. U.S.A., WASHINGTON COUNTY: U.S.A., IDAHO, KOOTENAI COUNTY: Olympic Peninsula; Elwha River. [No locality given on label] Elev. 1000 ft. 47.676278°, -116.686856°; uncertainty: 44000 m. Moist woods. Phenology: Flowers & Fruits. Origin: Native. Moist Canyons. Phenology: Flowers. Origin: Native. Geo. N. Jones 3242 Jul 1931 J. H. Sandberg s.n. Aug 1892 WTU-21450 WTU-97037

Rosaceae Rosaceae Rubus pedatus Sm. Rubus pedatus Sm. U.S.A., OREGON, CLATSOP COUNTY: U.S.A., ALASKA COUNTY: Onion Peak. Chomley [Chomly]. Elev. 3064 ft. 55.247376°, -132.322482°; WGS 84, uncertainty: 2500 m., T4N R10W S22; NAD 27, uncertainty: 805 m., Source: TRS2LL, Source: Digital Map, Georef'd by Joseph Stormer Georef'd by Ben Legler Phenology: Fertile. Origin: Native. Rocky slopes and pinnacles form the summit, where a small remnant stand of Abies amabilis and Tsuga heterophylla occurs (all G. B. Rigg 115 6 Aug 1929 the surrounding foreset are logged-off). Steep open slopes at the summit have an herbaceous flora with relic xeric elements. Shaded WTU-39244 places in the forest remnant, in rich humus. Flowers white. Phenology: Flowers. Origin: Native. Rosaceae K. L. Chambers 3131 29 Jun 1971 Rubus pedatus Sm. WTU-260940 U.S.A., ALASKA COUNTY: Hyder. Rosaceae 55.922087°, -130.037299°; WGS 84, uncertainty: 9639 m., Source: Digital Map, Georef'd by Joseph Stormer Rubus spectabilis Pursh Semi-open, forest edge on river flats, around old stumps; on moss over rocks etc. Common; fruit sub-translucent red. Phenology: U.S.A., WASHINGTON, SNOHOMISH Flowers. Origin: Native. COUNTY: Just northeast of the fork in Chennault Beach T. T. McCabe 8456 19 Jul 1941 road, above west facing steep grade, overlooking Puget Sound, about 8 miles south WTU-73611 of Mukilteo. (Mukilteo Quad. 7.5 Min. Topog. U.S.G.S.). Elev. 457 ft. Rosaceae T28N R4E S21 NW 1/4 SW 1/4; NAD 27, uncertainty: 805 m., Source: TRS2LL, Rubus ursinus Cham. & Schltdl. Georef'd by Ben Legler Alder woodlands, dispersed with evergreens, U.S.A., WASHINGTON, KLICKITAT COUNTY: shrub and grasslike vegetation. Phenology: Bingen Mountain, Bingen. Flowers. Origin: Native. 45.715°, -121.46333°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Lisa Westgard 8 Phenology: Flowers. Origin: Native. 9 Apr 1981 Wilhelm N. Suksdorf 1757 8 Jun 1901 WTU-316628, WTU-V-044970 WTU-197548 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus spectabilis Pursh U.S.A., WASHINGTON, PIERCE COUNTY: West of North-South alligned powerline. U.S.A., WASHINGTON COUNTY: Elev. 393 ft. Stevens Pass. 543800E, 5211500N 47.746222°, -121.085933°; WGS 84, Cytisus scoparius dominated prairie; slope flat; gravel loam; uncertainty: 3873 m., Source: GeoLocate, associated with Poa pratensis, Cytisus scoparius, Agrostis tenuis; Georef'd by Joseph Stormer few individuals growing in rock pile in prairie. Pacific blackberry. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. J. H. Sandberg 2986 David Thysell 518 23 Jun 1993 with J. B. Leiberg WTU-330001 Aug 1893 WTU-9521, WTU-V-18344 Rosaceae Rosaceae Rubus pedatus Sm. Rubus pedatus Sm. CANADA, BRITISH COLUMBIA: CANADA, BRITISH COLUMBIA: Glaicer. Columbia River, 9.7 miles south of Big Bend Bridge. [No collection date given]. Phenology: Flowers. Base of stupi-opening in dripping cedar-helock forest. Flowers white. Phenology: Fertile. Dr. C. Schaffer s.n. with Edwin B. Bartram T. T. McCabe 5326 25 Sep 1937 WTU-28604 WTU-78259

Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Cougar Mountain. San Juan Island, Friday Harbor Marine Laboratory grounds. 47.54056°, -122.09806°; WGS 84, uncertainty: 10000 m., Source: Elev. 10-23 ft. GeoLocate, Georef'd by WTU Staff 48° 32' 39" N, 123° 51" W; T35N R4W S12; Source: Specimen Phenology: Flowers. Origin: Native. label Mossy bald west of Fernald Laboratory and adjacent to parking lot. David M. Chapin s.n. 9 Oct 1982 Growing amidst grasses and over rocks; male and female plants present; flowers white. Phenology: Flowers. Origin: Native. WTU-314236 Sarah Gage SG-2813 7 May 1997 Rosaceae WTU-336248 Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, SNOHOMISH COUNTY: Rubus pedatus Sm. King (205th and Alaska Wy)/ Snohomish Undeveloped property of the north border of LFP, 10 yards in from 205th along the trail. U.S.A., ALASKA COUNTY: 47.91306°, -122.09694°; WGS 84, uncertainty: 10000 m., Source: Sitka. GeoLocate, Georef'd by WTU Staff 57.094858°, -135.272322°; WGS 84, uncertainty: 10000 m., Growing with various Rubus species. Phenology: Flowers. Origin: Source: Digital Map, Georef'd by Joseph Stormer Native. Phenology: Fertile. Origin: Native. Christie S. Isler s.n. 2 Apr 1998 T. Kincaid s.n. 16 Jun 1899 WTU-335294 WTU-101449

Rosaceae Rosaceae Rubus pedatus Sm. Rubus spectabilis Pursh CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, KING COUNTY: Five miles north of Takla Landing. Near Issaquah. Bare wet gravelly slide, open. Petals just fallen. Phenology: Fertile. 47.53028°, -122.03139°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, T. T. McCabe 7877 17 Jul 1940 Georef'd by WTU Staff Wet woods. Phenology: Flowers. Origin: Native. WTU-72629 G. N. Jones 6392 Rosaceae 20 Apr 1935 Rubus spectabilis Pursh WTU-28741, WTU-V-044975 U.S.A., WASHINGTON, SNOHOMISH COUNTY: Above Monte Cristo. Rosaceae Elev. 3500 ft. 47.98583°, -121.39278°; WGS 84, Rubus pedatus Sm. uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., ALASKA, WRANGELL PETERSBURG COUNTY: Moist slopes. Phenology: Flowers. Origin: Washington Bay, Kuiu Island. Native. 56.71885°, -134.364815°; WGS 84, uncertainty: 2000 m., Source: Digital Map, Georef'd by Joseph Stormer J. William Thompson 17412 On moss in spruce forest. Phenology: Flowers. Origin: Native. 30 Jun 1953 W. J. Eyerdam 5453 20 Aug 1947 WTU-171126, WTU-V-044974 WTU-118038 Rosaceae Rosaceae Rubus pedatus Sm. Rubus spectabilis Pursh U.S.A., IDAHO, BOUNDARY COUNTY: CANADA, BRITISH COLUMBIA: Near summit of Fisher Mountain. Near Fraser River, 3 miles southwest of Hope, B.C. Elev. 7000 ft. Alnus dominant. Swamp. Phenology: Flowers & Fruits. 48.866609°, -116.537156°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff James Hatter 10 8 May 1949 Dry coniferous woods, rocky soil. Phenology: Fertile. Origin: Native. WTU-143017 William H. Baker 13850 9 Aug 1955 WTU-295676 Rosaceae Rubus pedatus Sm. Rosaceae U.S.A., WASHINGTON COUNTY: Rubus ursinus Cham. & Schltdl. Base of Mount Hood. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, SAN JUAN COUNTY: Yellow Island. Thomas J. Howell s.n. Jul 1881 48.54361°, -123.16333°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-22953JWT-A Slopes above west spit; dry meadow. Trailing blackberry. Phenology: Flowers. Origin: Native. Rosaceae Dennis Phillips s.n. 3 May 1994 Rubus ursinus Cham. & Schltdl. WTU-332799 U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Island, Friday Harbor Marine Laboratory grounds; Rosaceae southwest of Fernald Lab. Elev. 9-22 ft. Rubus pedatus Sm. 48° 32' 39" N, 123° 51" W; T35N R4W S12; Source: Specimen label U.S.A., ALASKA COUNTY: Adjacent to proposed building site between parking lot and water's Kodiak Island. Olga Bay. Cannery Station. edge; mossy knoll and open meadow. With immature fruits. Elev. 500 ft. Phenology: Flowers. Origin: Native. T35S R31W S10; WGS 84, uncertainty: 2750 m., Source: Digital Map, Georef'd by Joseph Stormer Sarah Gage SG-2849 2 Jun 1997 Valley tundra, near Akalura cabin. Flowers pink. Phenology: Flowers. Origin: Native. WTU-336252 Ethel H. Looff 156 12 Jul 1981 Rosaceae WTU-314310 Rubus pedatus Sm. U.S.A., OREGON, CLACKAMAS COUNTY: Rosaceae Near Government Camp, Mount Hood. Elev. 5000 ft. Rubus ursinus Cham. & Schltdl. 45.304007°, -121.7548°; uncertainty: 3500 m. Alpine slopes. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, SAN JUAN COUNTY: Yellow Island, south side. J. William Thompson s.n. 17 Jul 1925 48.591111°, -123.033056°; WGS 84, uncertainty: 250 m., Source: Georeferenced, Georef'd by WTU Staff WTU-22953JWT-B Rocky slope above west spit; open rocky areas. Dewberry, trailing blackberry. Phenology: Flowers. Origin: Native. Rosaceae E. Habegger s.n. 1 May 1996 Rubus pedatus Sm. WTU-332798 CANADA, BRITISH COLUMBIA: Mountains 10-15 miles south of Telegraph Creek. Rosaceae Dry bank in high Abies timber slightly below timberline. Berries translucent red. Trailing or prone. Phenology: Fruits. Rubus pedatus Sm. T. T. McCabe 8865 15 Aug 1941 U.S.A., ALASKA COUNTY: Thum Bay, Knight Island, Prince William Sound. WTU-73658 60.206643°, -147.811189°; uncertainty: 2400 m. Amongst moss under Vaccinium bushes. Phenology: Fruits. Origin: Native. W. J. Eyerdam 3547 10 Sep 1940 WTU-292192 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus pedatus Sm. U.S.A., OREGON, CLATSOP COUNTY: U.S.A., ALASKA, WRANGELL PETERSBURG COUNTY: Near junction of Saddle Mountain Road & Highway 2, 10 miles Washington Bay, Kuiu Island. southeast of Seaside junction. 56.71885°, -134.364815°; WGS 84, uncertainty: 2000 m., Source: Elev. 800 ft. Digital Map, Georef'd by Joseph Stormer T5N R9W S24 N 1/2; NAD 27, uncertainty: 805 m., Source: In moss in spruce forest. Phenology: Fruits. Origin: Native. TRS2LL, Georef'd by Ben Legler At edge of coniferous forest; moist humus soil; with Menziesia, Alnus W. J. Eyerdam 5453 20 Aug 1947 & Pteridium. Common. Shrub 1-2.5 meters tall; fruit green. Phenology: Fruits. Origin: Native. WTU-292223 Albert N. Steward 7092 30 May 1956 with Celia B. Steward Rosaceae WTU-207193 Rubus pedatus Sm. U.S.A., ALASKA COUNTY: Rosaceae Chickamin Glacier, Texas Creek Summit. Open, above timberline. Abundant, frequently in and under Rubus ursinus Cham. & Schltdl. Phyllodoce, Cassiope; flowers white; just passing. Phenology: Flowers. Origin: Native. U.S.A., OREGON, JOSEPHINE COUNTY: Forest Road 176 off State Route 46; 1 mile west of Oregon Caves T. T. McCabe 8925 16 Jul 1941 National Monument; Siskiyou National Forest. Elev. 3650 ft. WTU-73612 42° 6' 35" N, 123° 24' 37" W; T40S R6W S4 S9 North-facing slope, mixed fir/ oak forest; along 176 road bank. Long stiff stems. Phenology: Flowers. Origin: Native. Rosaceae Jessie Johanson 07 26 Jun 1998 Rubus ursinus Cham. & Schltdl. with Jessie Johanson U.S.A., OREGON COUNTY: WTU-342633 Along the Umatilla River at Bingham Springs. 45.740935°, 0°; Georef'd by Joseph Stormer Dew Berry. Phenology: Flowers. Origin: Native. Rosaceae Wm. C. Cusick 3267a 16 Jul 1908 Rubus pedatus Sm. WTU-139203 U.S.A., ALASKA COUNTY: Bell Island, Sulphur Springs. Trailing over stumps in woods. Phenology: Fertile. Origin: Native. Rosaceae Mr. Ernest P. Walker 956 22 Aug 1915 Rubus spectabilis Pursh with Mrs. Ernest P. Walker U.S.A., ALASKA, KODIAK ISLAND COUNTY: WTU-76671 Port Vita, Raspberry Strait, Raspberry Island, Kodiak group. Elev. 1968 ft. 58.065556°, -153.076792°; uncertainty: 3000 m., Georef'd by Rosaceae Joseph Stormer Around alder brush. Forming thickets. Phenology: Flowers. Origin: Rubus pedatus Sm. Native. U.S.A., WASHINGTON COUNTY: W. J. Eyerdam 3700 1945 Washington Bay. In humus in hemlock-spruce forest. Phenology: Fertile. Origin: WTU-190880 Native. W. J. Eyerdam 8051 25 Jun 1955 Rosaceae WTU-292712 Rubus pedatus Sm. Rosaceae CANADA, BRITISH COLUMBIA: Moat Lake. Rubus ursinus Cham. & Schltdl. Elev. 4000 ft. 49° 41' N, 125° 23' W U.S.A., OREGON, JOSEPHINE COUNTY: Forest floor. Phenology: Flowers. Siskiyou National Forest; on 3905, 14.8 miles south of intersection of highways 3905 and 46 or [circa] 33 miles southeast of Cave J. E. Underhill 487 25 Jul 1957 Junction. Elev. 4080 ft. WTU-213996 Pseudotsuga-Quercus woods with wet area near stream, the upper slopes drier and with abundant Cornus. Abundant at margins of woods; petals white. Phenology: Flowers. Origin: Native. Melinda F. Denton 2678 25 Jun 1973 WTU-259010 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., OREGON, KLAMATH COUNTY: U.S.A., IDAHO, CLEARWATER COUNTY: Courtade ranch, near head of Denny Creek. Near Headquarters. Along wooded stream. Phenology: Flowers. Origin: Native. 46.63°, -115.80833°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Elmer I. Applegate 3711 26 Jul 1923 Creeping near-climax forest of Abies grandis and Thuja plicata. Berry black, 1.5 x 2 centimeters. Phenology: Flowers. Origin: Native. WTU-8828JWT R. F. Daubenmire s.n. 23 Aug 1942 Rosaceae WTU-271342 Rubus pedatus Sm. Rosaceae CANADA, BRITISH COLUMBIA: Near Yakoun River delta, Masset Inlet, Graham Island. Rubus spectabilis Pursh 53.646767°, -132.203746°; uncertainty: 2000 m. Trailing over logs and stumps in dense coniferous woods; flowers U.S.A., ALASKA COUNTY: white. Common; flowers white. Voucher specimen cited in Flora of Middleton Island is located in the Gulf of Alaska about 115 miles The Queen Charlotte Islands. 1. Systematics of the Vascular Plants. south of Valdez. by James A. Calder and Roy L. Taylor. Queen's Printer, Ottawa, 59° 25' N, 146° 20' W 1968. Phenology: Flowers. Steep west faing [facing?] bluff. Other species: Fritellaria camtchatcensis, Urtica lyallii, maiantheum dilatum, Rubus J. A. Calder 21559 15 Jun 1957 spectabilis, heracleum maximum, montia ibirica, Ranunculus with D. B. O. Savile, R. L. Taylor occidentalis, Cardamine umbelata, and Dryopteris austriaca. Petals rose-lavender. Spines quickly deciduous. Small shrubs to 1 meter WTU-240941 tall. At this date in flower only in more protected areas and on lower branches. Phenology: Flowers. Origin: Native. Rosaceae John H. Thomas 5811 6 Jun 1956 Rubus ursinus Cham. & Schltdl. WTU-181776 U.S.A., OREGON, MULTNOMAH COUNTY: Willamette River near Portland. Rosaceae Banks. Phenology: Flowers. Origin: Native. Rubus ursinus Cham. & Schltdl. J. W. McFarland s.n. 14 Apr 1941 U.S.A., OREGON, KLAMATH COUNTY: WTU-97657 Southwest corner of Crater Lake National Park, north wall of Redblanket Canyon. 42.805602°, -122.313802°; WGS 84, uncertainty: 7216 m., Rosaceae Source: Digital Map, Georef'd by Joseph Stormer, Invalid coordinates (located outside county bounds). Rubus spectabilis Pursh Phenology: Flowers. Origin: Native. U.S.A., ALASKA, KODIAK ISLAND COUNTY: William H. Baker 7036 29 Jul 1950 Port Vita, Raspberry Strait, Raspberry Island, Kodiak group. Elev. 1968 ft. WTU-155835 58.065556°, -153.076792°; uncertainty: 3000 m., Georef'd by Joseph Stormer One meter high; forming thickets. Phenology: Flowers. Origin: Rosaceae Native. Rubus ursinus Cham. & Schltdl. W. J. Eyerdam 3689 1945 CANADA, BRITISH COLUMBIA: WTU-192568 1 mile south of Sooke, Vancouver Island. Climbing over mossy glaciated boulders, open. Flowers white. Rosaceae Phenology: Flowers. Rubus pedatus Sm. T. T. McCabe 5665 27 Apr 1938 U.S.A., ALASKA COUNTY: WTU-72807 Port Armstrong, Baranof Island. Phenology: Vegetative. Origin: Native. Rosaceae A. A. Heller s.n. 25 Jul 1929 Rubus ursinus Cham. & Schltdl. WTU-87313 U.S.A., OREGON, DOUGLAS COUNTY: About Diamond Lake. Open woods. Phenology: Flowers. Origin: Native. Morton E. Peck 19220 7 Jul 1936 WTU-9438JWT Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., OREGON, TILLAMOOK COUNTY: U.S.A., WASHINGTON, KING COUNTY: Hebo. Seattle. Phenology: Flowers & Fruits. Origin: Native. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer G. E. Morrill 219 23 Jul 1933 Phenology: Flowers. Origin: Native. WTU-26662 Emma A. Shumway 2993 May 1892 WTU-9529 Rosaceae Rubus spectabilis Pursh Rosaceae CANADA, BRITISH COLUMBIA: Rubus ursinus Cham. & Schltdl. Glacier Gulch at base of Hudson Bay Mountain, Smithers. Elev. 3000 ft. U.S.A., WASHINGTON, KING COUNTY: Dominant on talus slope at base of cliff. Up to 5 feet tall; flowers red. Seattle. Phenology: Flowers. 47.60621°, -122.332071°; WGS 84, uncertainty: 15000 m., Source: GeoLocate, Georef'd by Ibro Polovina J. A. Calder 12897 13 Jul 1954 Found in woody thickets, in open. Phenology: Flowers. Origin: with D. B. O. Savile, J. M. Ferguson Native. WTU-172940 D.A.H. s.n. May 1909 WTU-9758 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: Seattle. U.S.A., OREGON, JOSEPHINE COUNTY: 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Siskiyou National Forest; on 3941, 3.2 miles east of interesection of Source: Digital Map, Georef'd by Joseph Stormer 3897 and 3941 or ca. [circa] 25 miles east of Cave Junction. Dew berry. Phenology: Flowers & Fruits. Origin: Native. Elev. 5120 ft. 42.116115°, -123.384297°; WGS 84, uncertainty: 11637 m., C. V. Piper 70 1888 Source: Digital Map, Georef'd by Joseph Stormer Pseudotsuga woods. On slopes. Occasional; petals deep WTU-9589 pinkish-purple. Phenology: Flowers. Origin: Native. Melinda F. Denton 2589 23 Jun 1973 Rosaceae WTU-258987 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, KING COUNTY: Rosaceae Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Rubus ursinus Cham. & Schltdl. Source: Digital Map, Georef'd by Joseph Stormer Blackberry. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: Seattle. C. V. Piper 70 1888 47.60621°, -122.332071°; WGS 84, uncertainty: 15000 m., Source: GeoLocate, Georef'd by Ibro Polovina WTU-9590 Open woods. Flowers perfect. Phenology: Flowers. Origin: Native. George Neville Jones s.n. 30 May 1933 Rosaceae WTU-36219 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, KING COUNTY: Rosaceae Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Rubus ursinus Cham. & Schltdl. Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers & Fruits. Origin: Native. U.S.A., WASHINGTON, PIERCE COUNTY: Tacoma. Emma A. Shumway s.n. May 1891 47.25306°, -122.44306°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-112688 Waste places among bogs. Phenology: Flowers. Origin: Native. J. B. Flett s.n. 27 May 1896 WTU-33514 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus vestitus Weihe & Nees U.S.A., OREGON, LANE COUNTY: U.S.A., OREGON, LANE COUNTY: Elk Creek. In The Nature Conservancy Willow Creek natural area, 0.3 Elev. 425 ft. kilometers west of Bertleson Drive; West Eugene. T17S R6W S30; NAD 27, uncertainty: 805 m., Source: TRS2LL, Elev. 420 ft. Georef'd by Ben Legler T18S R4W S4 NW 1/4 of SW 1/4; NAD 27, uncertainty: 805 m., Douglas Fir forest (Lower Montane). Pseudotsuga taxifolia- Thuja Source: TRS2LL, Georef'd by Ben Legler plicata- Acer macrophyllum - Alnus oregana. Creek bottom, coarse Partial shade, bird-sown adventive, with Rubus armeniacus, saturated loam in partial shade. Phenology: Flowers. Origin: Native. Pseudotsuga. Pink petals and filaments, reflexed sepals, 3-foliate, erect, arching fertile canes. Phenology: Flowers. Origin: Introduced. L. E. Detling 2475 8 Apr 1938 P. F. Zika 13229 8 Jul 1997 WTU-74386 WTU-352843 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus spectabilis Pursh U.S.A., WASHINGTON, CLALLAM COUNTY: Seqium; Fort Hills. U.S.A., ALASKA, KODIAK ISLAND COUNTY: Phenology: Flowers. Origin: Native. Port Vita, Raspberry Strait, Raspberry Island, Kodiak group. Elev. 1968 ft. J. M. Grant s.n. Jun 1910 58.065556°, -153.076792°; uncertainty: 3000 m., Georef'd by Joseph Stormer WTU-9714 In spruce forest. Phenology: Flowers. Origin: Native. W. J. Eyerdam 4083 1945 Rosaceae WTU-292224 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, ISLAND COUNTY: Rosaceae Whidbey Island. 48.146201°, -122.552685°; WGS 84, uncertainty: 30000 m., Rubus vestitus Weihe & Nees Source: Digital Map, Georef'd by Joseph Stormer Woods. Phenology: Flowers. Origin: Native. U.S.A., OREGON, LANE COUNTY: In The Nature Conservancy Willow Creek natural area, 0.3 N. L. Gardner s.n. 26 May 1897 kilometers west of Bertleson Drive; West Eugene. Elev. 420 ft. WTU-9740 T18S R4W S4 NW 1/4 of SW 1/4; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Partial shade, bird-sown adventive, with Rubus armeniacus, Rosaceae Pseudotsuga. Pink petals and filaments, reflexed sepals, 3-foliate, erect, arching fertile canes. Phenology: Flowers. Origin: Introduced. Rubus bifrons Vest P. F. Zika 13229 8 Jul 1997 U.S.A., WASHINGTON, YAKIMA COUNTY: West bank of Yakima River, north of West Birchfield Road. WTU-352842 Elev. 984 ft. 46° 35.3' N, 120° 28' W Adventive, brushy margins of Iris pseudacorus marsh, between Rosaceae arboretum. Arching 2 metres shrub, white flowers 32 milimeters diameter, occasional. Phenology: Flowers. Origin: Introduced. Rubus spectabilis Pursh Peter F. Zika 19771 31 May 2004 U.S.A., ALASKA COUNTY: Hinchinbrook Island. WTU-358809 In swamps. Phenology: Flowers. Origin: Native. Ingv. L. Norberg s.n. 28 Jun 1937 Rosaceae WTU-292194 Rubus spectabilis Pursh U.S.A., OREGON, CURRY COUNTY: Rosaceae 3.9 miles east of Highway 101, along north bank Pistol River. 42.277689°, -124.353225°; uncertainty: 600 m., Georef'd by Rubus ortivus (Bailey) Bailey × Rubus stellatus Joseph Stormer Abundant shrubs to 1.5 meters tall, red, drooping flowers, in U.S.A., ALASKA COUNTY: Pseudotsuga, Rhododendron. Phenology: Flowers. Origin: Native. Hope. 60.870848°, -149.640278°; WGS 84, uncertainty: 10000 m., E. K. Balls 16451 5 May 1951 Source: Digital Map, Georef'd by Joseph Stormer Deep shade on gravelly creek bank. Phenology: Flowers. WTU-141630 J. P. Anderson 6533 3 Jun 1941 WTU-292746 Rosaceae Rosaceae Rubus alpinus × Rubus stellatus Rubus parviflorus Nutt. U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Seward. Washington Bay, Kuiu Island. 60.104167°, -149.442222°; WGS 84, uncertainty: 8146 m., 56.721111°, -134.356907°; WGS 84, uncertainty: 2400 m., Source: GeoLocate, Georef'd by Joseph Stormer Source: Digital Map, Georef'd by Joseph Stormer Roadside. Phenology: Flowers. On border of spruce forest. Phenology: Fruits. Origin: Native. J. P. Anderson 6741 12 Jun 1941 W. J. Eyerdam 5445 20 Aug 1947 WTU-292747 WTU-122909

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., ALASKA COUNTY: U.S.A., OREGON, DOUGLAS COUNTY: Southeastern Alaska; Tongass National Forest; Chichagof Island, Beatty Creek RNA, Roseburg BLM, 10 air miles southwest of town of Tenakee Inlet. Head of Saltery Bay, east. Myrtle Creek. Elev. 25 ft. Elev. 950 ft. Brush thicket at flt [flat] edge. Phenology: Flowers. Origin: Native. T30S R6W S31 NW 1/4; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Jim Downs 74 19 Jun 1980 Margins of Umbellularia riparian zone, serpentine soil. White flowers. Phenology: Flowers. Origin: Native. WTU-282653 P. F. Zika 10264 11 May 1987 Rosaceae WTU-336853 Rubus parviflorus Nutt. Rosaceae U.S.A., ALASKA COUNTY: Southeastern. Tongass National Forest. Catherine Island (just north Rubus parviflorus Nutt. west of Baranof Island). East side about 2 miles south of Pt. Thatcher. U.S.A., OREGON, JOSEPHINE COUNTY: Small rocky bay at Local Creek, just southwest of a rocky spit. In Siskiyou National Forest. On Road 3941, 14.2 miles east of Cave large banks at the beach-forest ecotone. Common. Phenology: Junction or 1.4 miles east of intersection of 46 and 3941. Flowers. Origin: Native. Elev. 2160 ft. 42.127345°, -123.455933°; uncertainty: 5000 m., Georef'd by Mary Clay Muller 3540 18 Jun 1980 Joseph Stormer Stream near road with Alnus, Quercus, Acer macrophyllum, WTU-282572 Pseudotsuga, Salix, some Arbutus and Chamaecyparis. Frequent; clambering in brush; flowers white. Phenology: Flowers & Fruits. Rosaceae Origin: Native. Rubus parviflorus Nutt. Melinda F. Denton 2515 22 Jun 1973 U.S.A., ALASKA COUNTY: WTU-259622 Hydaburg. Elev. 0 ft. 55.212366°, -132.806068°; WGS 84, uncertainty: 3500 m., Rosaceae Source: Digital Map, Georef'd by Joseph Stormer Sea level. Forest, muskeg and especially distur ground. A food plant. Rubus parviflorus Nutt. Phenology: Flowers & Fruits. Origin: Native. U.S.A., OREGON, JOSEPHINE COUNTY: Helen H. Norton 616-H 29 Jun 1979 Siskiyou National Forest; on 3905, 14.8 miles south of intersection of highways 3905 and 46 or [circa] 33 miles southeast of Cave WTU-281168 Junction. Elev. 4080 ft. Pseudotsuga-Quercus woods with wet area near stream, the upper Rosaceae slopes drier and with abundant Cornus. Common in gulch; small perennial shrub; petals white. Phenology: Flowers. Origin: Native. Rubus parviflorus Nutt. Melinda F. Denton 2677 25 Jun 1973 U.S.A., ALASKA COUNTY: Baranof Island. About 5 miles north of Sitka at Old Sitka ruins. WTU-258801 Growing on fill area near the ocean. Phenology: Flowers. Origin: Native. Rosaceae Mary Clay Muller 709 14 Jul 1975 Rubus parviflorus Nutt. WTU-274555 U.S.A., MONTANA COUNTY: Glacier National Park. Many glacier. Phenology: Flowers. Origin: Native. G. N. Jones 5333 Jul 1935 WTU-36227 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus idaeus var. sachalinensis U.S.A., MONTANA COUNTY: CANADA, BRITISH COLUMBIA: Glacier National Park. Fernie. Many Glacier. Phenology: Flowers. Origin: Native. 49.5°, -115.066667°; uncertainty: 4000 m. Grey's Bot. 821. Phenology: Fertile. G. N. Jones 5333 Jul 1935 F. Comte F-3 Jul 1927 WTU-36224 WTU-329414 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., IDAHO COUNTY: White Bird Grade. U.S.A., IDAHO COUNTY: In ravine. Phenology: Fruits. Origin: Native. White Bird Creek. In ravine. Phenology: Fruits. Origin: Native. G. N. Jones 4994 Jun 1935 G. N. Jones 4994 Jun 1935 WTU-36226 WTU-36225 Rosaceae Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., IDAHO, CUSTER COUNTY: Mount Heyburn, Sawtooth Range. CANADA, BRITISH COLUMBIA: Elev. 8500 ft. Emerald Lake. Rocky slopes. Phenology: Fertile. Field. Gray's Bot. 821. Phenology: Flowers & Fruits. J. William Thompson 13670 4 Aug 1936 F. Comte 1754 10 Aug 1957 WTU-3994JWT WTU-329400

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. CANADA, BRITISH COLUMBIA: CANADA, BRITISH COLUMBIA: Subalpine slopes near Lake Bootahnie [Botanie Lake], Marble Base of Mt. Loveway about 6 miles north of Little Fort along North Mountains. Thompson River. Elev. 5000 ft. Elev. 1500 ft. 50.383684°, -121.541882°; uncertainty: 1464 m., Georef'd by Along edge of cliff at roadside. Common, flowers white. Phenology: Joseph Stormer Flowers. Origin: Native. Phenology: Flowers. J. A. Calder 8626 3 Jun 1953 J. William Thompson 190 20 Jun 1938 with D. B. O. Savile with Emily M. Thompson WTU-175458 WTU-48864 Rosaceae Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., MONTANA, LAKE COUNTY: U.S.A., ALASKA, KENAI PENINSULA COUNTY: On east shore of Flathead Lake, 1 mile south of Yellow Bay. Mount Marathon area, Seward. At edge of woods. Phenology: Fertile. 60° 6' N, 149° 27' W Occasional on grassy slope in ravine on south slope of mountain. C. L. Hitchcock 17735 27 Jun 1948 Phenology: Flowers. WTU-125083 J. A. Calder 5894 13 Jul 1951 WTU-173255 Rosaceae Rosaceae Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., IDAHO, VALLEY COUNTY: U.S.A., OREGON, WHEELER COUNTY: At base of Thunderbolt Mountain; Payette National Forest. Along Pine Creek 7 miles above of mouth. Elev. 7000 ft. Thicket. Phenology: Fertile. 44.7325°, -115.63861°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Morton E. Peck 10034 4 Jul 1921 Slopes. Phenology: Flowers. WTU-22955JWT J. William Thompson 13919 19 Jul 1937 WTU-10559JWT Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rosaceae U.S.A., ALASKA COUNTY: Rubus idaeus var. sachalinensis About 15 miles southeast of Fairbanks. 64.684242°, -147.356299°; WGS 84, uncertainty: 15000 m., U.S.A., MONTANA, MISSOULA COUNTY: Source: GeoLocate, Georef'd by Joseph Stormer Talus slide 5 miles above Bonner. Open area. Phenology: Flowers. Granitic rock. Phenology: Fertile. R. John Taylor 2876 25 Jun 1965 C. L. Hitchcock 11423 25 Jul 1945 with C. V. Muhlick WTU-226533 WTU-102081 Rosaceae Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rubus idaeus var. sachalinensis CANADA, BRITISH COLUMBIA: Head of Dease Lake. U.S.A., IDAHO, VALLEY COUNTY: Open dry brulee. Height 3 feet. Fertile. Phenology: Fruits. At base of Thunderbolt Mountain; Payette National Forest. Elev. 7000 ft. T. T. McCabe 8794 3 Aug 1941 44.7325°, -115.63861°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-72511 Slopes. Phenology: Fertile. J. William Thompson 13919 19 Jul 1937 Rosaceae WTU-38342 Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., ALASKA COUNTY: Rosaceae Near Park Headquarters, Mount McKinley National Park. Open forest slope. Phenology: Flowers. Rubus idaeus L. var. sachalinensis (H. Lév.) Focke Aven Nelson 3994 28 Jul 1939 U.S.A., MONTANA, LEWIS AND CLARK COUNTY: with Ruth A. Nelson Just below Gibson Dam circa 25 miles northwest of Augusta. Gravel bench. Phenology: Fertile. WTU-75145 C. L. Hitchcock 18075 6 Jul 1948 Rosaceae WTU-125084 Rubus idaeus L. ssp. strigosus (Michx.) Focke Rosaceae U.S.A., MONTANA, GRANITE COUNTY: Rock Creek 10 miles from mouth. Rubus idaeus L. ssp. strigosus (Michx.) Focke 46.622569°, -113.653333°; WGS 84, uncertainty: 1315 m., Source: Digital Map, Georef'd by Maria Yousoufian U.S.A., WASHINGTON, SPOKANE COUNTY: Dry bank. Fruits red. Phenology: Flowers & Fruits. Latah Creek, east of Spangle. 47.28167°, -117.15444°; WGS 84, uncertainty: 1000 m., Source: C. L. Hitchcock 13701 19 Aug 1945 GeoLocate, Georef'd by WTU Staff with C. V. Muhlick On a talus slope. Phenology: Fertile. WTU-100868 Wilhelm N. Suksdorf 8680 27 Jun 1916 WTU-197559 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus lasiococcus A. Gray U.S.A., OREGON, COLUMBIA COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Old Highway 30. 1/8th miles from intersection of Washington Way. Near Big Four Inn. Elev. 100 ft. Elev. 3500 ft. In loam. Northwest slope. With Acer macrophyllum, ferns, Vaccinium 48.06611°, -121.50778°; WGS 84, uncertainty: 5000 m., Source: parvilolium, Thuja picata. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Deep woods. Phenology: Flowers. Origin: Native. S. L. Strong 53 17 May 1982 J. William Thompson 14720 15 Jun 1940 WTU-361679 WTU-361685 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus lasiococcus A. Gray U.S.A., WASHINGTON, KING COUNTY: 10 miles from Renton. U.S.A., WASHINGTON, YAKIMA COUNTY: 47.48306°, -122.21583°; WGS 84, Chinook Pass, Cascade Mountains. uncertainty: 10000 m., Source: GeoLocate, Elev. 5000 ft. Georef'd by WTU Staff 46.87222°, -121.51444°; WGS 84, uncertainty: 10000 m., Source: Moist woods. Phenology: Flowers. Origin: GeoLocate, Georef'd by WTU Staff Native. Alpine slopes. Phenology: Flowers. Origin: Native. G. L. Wittrock s.n. J. William Thompson 15132 1 Aug 1940 6 Apr 1928 WTU-361686 WTU-361680, WTU-V-18374 Rosaceae Rosaceae Rubus bifrons Vest Rubus bifrons Vest U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Near Astoria Lane, Route 105, Ocosta. U.S.A., WASHINGTON, GRAYS HARBOR Elev. 49 ft. COUNTY: 46° 53.5' N, 124° 0.6' W; T16N R11W S3 Humptulips River. Sunny roadbanks, with Picea sitchensis, Tsuga heterophylla, 47.22956°, -123.956466°; WGS 84, Gaultheria shallon. Common arching shrub. Phenology: Flowers & uncertainty: 10000 m., Source: GeoLocate, Fruits. Georef'd by WTU Staff Climbing among shrubs along bank. Phenology: Peter F. Zika 19785 3 Jun 2004 Fertile. Origin: Introduced. WTU-361697 G. N. Jones 4593 8 Apr 1934 Rosaceae WTU-361683, WTU-V-060715 Rubus spectabilis Pursh U.S.A., OREGON, LANE COUNTY: Rosaceae Along the Siltcoos outlet, a mile from the ocean. T19S R12W; WGS 84, uncertainty: 6506 m., Source: Digital Map, Rubus lasiococcus A. Gray Georef'd by Joseph Stormer In open second growth. Shrub 2-3 meters tall; flowers deep pink, U.S.A., WASHINGTON, KITTITAS COUNTY: almost red. Phenology: Flowers & Fruits. Bald Mountain, Snoqualmie National Forest. Elev. 5000 ft. Arthur Cronquist 6872 21 Apr 1953 46.976389°, -120.965833°; NAD 27, uncertainty: 5000 m., Source: Georeferenced, Georef'd by WTU Staff WTU-157017 Alpine slopes. Phenology: Flowers. Origin: Native. J. William Thompson 14854 24 Jun 1940 Rosaceae WTU-361684 Rubus spectabilis Pursh CANADA, BRITISH COLUMBIA: Sarita River. Elev. 200 ft. Drain, in salal at roadside; Exposure northeast, Slope: 0-2. Flowers pink. Distributed by the Herbarium, University of Victory, Canada. Phenology: Fruits. Origin: Native. A.Y. 291 Dec 1963 with W. H. WTU-220552 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh CANADA, BRITISH COLUMBIA: U.S.A., ALASKA COUNTY: Drainage Kennedy River area, near Ucluelet, Vancouver Island. . Mist Harbor, Shumagin Islands. Elev. 810 ft. 55.127447°, -159.850674°; uncertainty: 2000 m., Georef'd by Exposure-shade; Slope-10. Along edge of river. Flowers red. Joseph Stormer Distributed by the Herbarium, University of Victory, Canada [No Hillside. Phenology: Flowers. Origin: Native. collection date given on label. However, Young and Hubbard collected together in May and June of 1963]. Phenology: Flowers & D. Martel 108 30 May 1935 Fruits. Origin: Native. WTU-101659 A. Young 90 with W. Hubbard Rosaceae WTU-218842 Rubus spectabilis Pursh Rosaceae U.S.A., ALASKA, ALEUTIANS EAST COUNTY: Squaw Harbor, Alaska Peninsula. Rubus spectabilis Pursh 55.240593°, -160.53857°; uncertainty: 3200 m. Phenology: Fruits. Origin: Native. CANADA, BRITISH COLUMBIA: Mount Douglas Park, Bower Creek. G. N. Jones 8922 14 Jul 1936 Elev. 200 ft. Among ferns under hemlock. Exposure-none; Slope-zero. WTU-101446 Distributed by the Herbarium, University of Victory, Canada . Phenology: Flowers. Origin: Native. Rosaceae W. Hubbard 28 6 May 1963 Rubus spectabilis Pursh WTU-221633 U.S.A., ALASKA, ALEUTIANS EAST COUNTY: Akutan, Aleutian Islands. Rosaceae 54.135556°, -165.773056°; WGS 84, uncertainty: 2500 m., Source: GeoLocate, Georef'd by Joseph Stormer Rubus spectabilis Pursh Phenology: Flowers. Origin: Native. U.S.A., ALASKA COUNTY: John Rudd s.n. Jul 1935 Hyder, Texas Lake. Forest edges etc. Abundant here at Texas Lake berries all red but WTU-101571 many be [are]yellow at intetrmediate altitudes. Height 3-4 feet. Phenology: Fruits. Origin: Native. Rosaceae T. T. McCabe 8473 19 Jul 1941 Rubus spectabilis Pursh WTU-73967 U.S.A., ALASKA COUNTY: Seward, Kenai Peninsula. Rosaceae 60° 6' N, 149° 27' W Along bank of small creek. Quite common; Only a few flowers in Rubus spectabilis Pursh bloom; flowers pink. Phenology: Flowers. CANADA, BRITISH COLUMBIA: J. A. Calder 5132 8 Jun 1951 Beach 4 miles northwest of Jordan River, Vancouver Island. Also seen on roadside, up to 5 miles south of here, probably passed WTU-173256 further south. Shrub up to 10 feet tall; flowers cold red, berries not yet colored. In flower. Phenology: Flowers. Origin: Native. Rosaceae T. T. McCabe 5670 27 Apr 1938 Rubus spectabilis Pursh WTU-72809 CANADA, BRITISH COLUMBIA: 21 Mile Creek, Alta Lake. Rosaceae Elev. 2500 ft. Petals: rose colored, faint, sweet, red or yellow. Herbarium MBG Rubus spectabilis Pursh 1016118 (Washington). Phenology: Flowers & Fruits. Origin: Native. U.S.A., ALASKA COUNTY: Unknown 5495 6 Aug 1967 Port Chalmers, Montague Island, Prince William Sound. Elev. 10 ft. WTU-329031 Along stream banks, north exposure. Common. Corolla rose. Phenology: Flowers. Origin: Native. J. David Solf 8 27 May 1959 WTU-232465 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Mineral Creek road, Valdez. Kodiak. Roadside. Phenology: Flowers. Origin: Native. Common; plant average 3-5 feet. [No collection date given on Label]. Phenology: Flowers. Origin: Native. Maxcine Williams 2190 21 Jun 1968 Jeannette Stewart 2 WTU-242865 WTU-169451 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., ALASKA COUNTY: Mainland, Le Conte Bay. U.S.A., ALASKA COUNTY: Wet woods. Phenology: Fruits. Origin: Native. South Tongass National Forest; mouth of Harris River, 12 mile Arm of Kasaan Bay Prince of Wales Island. Mr. Ernest P. Walker 883 30 Jul 1915 Open cutover Tsuga-Picea forest with shrub vegetation. Common in with Mrs. Ernest P. Walker openings, flowers pink. Height; 5-7 feet. Pollen removed for reference collection Linda Brubaker UW College of Forest WTU-76672 Resources February 1995. Phenology: Flowers. Origin: Native. Elbert L. Little Jr. 18004 3 May 1961 Rosaceae WTU-249782 Rubus spectabilis Pursh CANADA, BRITISH COLUMBIA: Rosaceae Whytecliffe, twelve miles west of north Vancouver. Along roadsides, open or semi-open. Generally in flower; flowers Rubus spectabilis Pursh strong crimson. Phenology: Flowers. Origin: Native. U.S.A., ALASKA COUNTY: T. T. McCabe 3278 24 Apr 1936 Entrance Island in Hobart Bay, 90 miles southeast of Juneau, North Tongass National Forest. WTU-45276 Elev. 10 ft. Tsuga-Picea forest, open area at edge of shore. Scattered in open area. Shrub,pink purple flowers. Stems light brown, smooth except Rosaceae for spines. Phenology: Flowers. Origin: Native. Rubus spectabilis Pursh Elbert L. Little Jr. 18070 10 May 1961 CANADA, BRITISH COLUMBIA: WTU-250250 East shore of Graham Reach, two miles north of Swanson Bay. Small slide, at edge of rock beach. Bush, four feet tall; flowers cold red. Only one yet seen in flower. Phenology: Flowers. Origin: Native. Rosaceae T. T. McCabe 3335 9 May 1936 Rubus spectabilis Pursh WTU-45275 CANADA, BRITISH COLUMBIA: Haines Road (km 86), Klehini River Valley. Elev. 1344 ft. Rosaceae 59° 33' N, 136° 31' W In openings of Picea sitchensis forest with Cornus canadensis. Rubus spectabilis Pursh Phenology: Flowers & Fruits. Origin: Native. CANADA, BRITISH COLUMBIA: G. W. Douglas 11019 25 Jun 1979 Upper Hat Creek, Marble Mountains. with M. J. Ratcliffe Elev. 3000 ft. 50.817329°, -121.577703°; uncertainty: 16000 m. WTU-282005 Arid slopes. Phenology: Fruits. Origin: Native. J. William Thompson 507 18 Jul 1938 with Emily M. Thompson WTU-17798JWT Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Seattle W.T. Seattle. 47.60639°, -122.33083°; WGS 84, 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., uncertainty: 10000 m., Source: GeoLocate, Source: Digital Map, Georef'd by Joseph Stormer Georef'd by WTU Staff Phenology: Flowers & Fruits. Origin: Native. Herbarium, University of Washington Collection by C.V. Piper Presented by Young Naturalists' Emma A. Shumway s.n. May 1892 Society, Seattle. Phenology: Flowers. Origin: Native. WTU-112692 E. S. Meany s.n. Rosaceae Apr 1885 Rubus spectabilis Pursh WTU-9641, WTU-V-18343 U.S.A., WASHINGTON, KING COUNTY: Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Rosaceae Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Rubus spectabilis Pursh E. S. Meany s.n. 17 Apr 1898 U.S.A., WASHINGTON, KING COUNTY: Seattle. WTU-36212 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer Rosaceae Phenology: Flowers. Origin: Native. Rubus spectabilis Pursh C. V. Piper s.n. U.S.A., WASHINGTON, KING COUNTY: 29 Apr 1888 Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., WTU-9600, WTU-V-18389 Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. E. S. Meany s.n. 17 Apr 1898 Rosaceae WTU-36209 Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: Rosaceae Seattle. 47.613617°, -122.310098°; WGS 84, Rubus spectabilis Pursh uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer U.S.A., WASHINGTON, KING COUNTY: Phenology: Flowers. Origin: Native. Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., C. E. Burglehaus s.n. Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. May 1891 E. S. Meany s.n. 17 Apr 1898 WTU-9812, WTU-V-18387 WTU-36210

Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Seattle. University Campus. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., 47.657643°, -122.305815°; WGS 84, uncertainty: 629 m., Source: Source: Digital Map, Georef'd by Joseph Stormer Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. [No collection year is given on label]. Phenology: Flowers. Origin: Native. Emma A. Shumway 2986 May 1892 Sueila W. Lutyse s.n. May WTU-9528 WTU-9636 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: University of Washington campus, Seattle. Fort Lawton, Seattle. 47.658007°, -122.306385°; WGS 84, uncertainty: 547 m., Source: 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: Digital Map, Georef'd by Joseph Stormer GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Thicket. Fruit yellow. Phenology: Fruits. Origin: Native. Sueila W. Lutyse s.n. Apr 1905 G. N. Jones 4850 19 May 1934 WTU-9635 WTU-25303

Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Seattle. Vashon Island. 47.60621°, -122.332071°; WGS 84, uncertainty: 16873 m., 47.4475°, -122.45861°; WGS 84, uncertainty: 10000 m., Source: Source: GeoLocate, Georef'd by Joseph Stormer GeoLocate, Georef'd by WTU Staff Moist woods. Abundant. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. D.A.H. s.n. May 1909 J. W. McFarland s.n. 18 Apr 1941 WTU-9757 WTU-89690

Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Fort Lawton, Seattle. 1/2 mile north of Seattle. 47.660003°, -122.418938°; WGS 84, uncertainty: 1382 m., 47.635339°, -122.33083°; WGS 84, uncertainty: 1000 m., Source: Source: Digital Map, Georef'd by Joseph Stormer GeoLocate, Georef'd by WTU Staff West slope. Phenology: Flowers. Origin: Native. In wooded swamp. Phenology: Flowers. Origin: Native. G. L. Wittrock s.n. 14 Apr 1928 W. J. Eyerdam 6493 10 May 1949 WTU-23251 WTU-290986

Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Bellevue. Snoqualmie Pass. Cascade Mountains. 47.61056°, -122.19944°; WGS 84, uncertainty: 10000 m., Source: Elev. 3000 ft. GeoLocate, Georef'd by WTU Staff 47.426389°, -121.406667°; NAD 27, uncertainty: 1000 m., Source: Phenology: Flowers. Origin: Native. Georeferenced, Georef'd by WTU Staff Alpine slopes. Phenology: Flowers. Origin: Native. I. L. Norberg s.n. 21 Mar 1934 J. William Thompson 9699 7 Aug 1933 WTU-26208 WTU-23015JWT Rosaceae Rosaceae Rubus spectabilis Pursh Rubus arcticus L. U.S.A., WASHINGTON, KING COUNTY: Fort Lawton, Seattle. U.S.A., ALASKA COUNTY: 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: Alaska, southeastern. Tongass National Forest mainland, Port GeoLocate, Georef'd by WTU Staff Houghton extreme eastern end at head of salt chuck. Thicket. Fruit red. Phenology: Fruits. Origin: Native. T52S R77E S9 N 1/4 Upper beach meadows, sea level. These are very lanky G. N. Jones 4850 19 May 1934 late-flowering specimens. Phenology: Flowers. WTU-25258 Mary Clay Muller 4377 18 Jan 1981 WTU-289092 Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Tongass National Forest. About 30 miles north of Juneau at Berners Port San Juan, Evans Island. Bay. Head of Echo Cove. 60.050833°, -148.0675°; uncertainty: 1200 m. Growing in upper beach meadow. Phenology: Flowers. In damp meadow. Phenology: Flowers. Mary Clay Muller 4177 16 Sep 1980 W. J. Eyerdam 5828 12 Jul 1948 WTU-288956 WTU-174829

Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Tongass National Forest East side of Baranof Island. Cosmos Cove, Glacier Highway. on the south shore 2/3 the way up the bay. Just west of moraine. 57.148867°, -134.115756°; WGS 84, uncertainty: 438323 m., T53S R66E S26 Source: Digital Map, Georef'd by Joseph Stormer Common in upper part of beach meadow. Sea level. Phenology: Phenology: Flowers. Flowers. Amy Rude s.n. 3 Jun 1945 Mary Clay Muller 3556 18 Jan 1980 WTU-110878 WTU-282574 Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA COUNTY: Aleutian Islands. Attu Island; Peaceful Valley. Tongass National Forest. Mainland. Head of Port Houghton 52.834262°, 173.152835°; uncertainty: 3000 m. southwest shore of Salt Chuck about 200 ft south of outlet canal. In moist loam at sea level. South slope. Phenology: Flowers. Elev. 0 ft. T52S R77E S20; uncertainty: 1200 m., Georef'd by Joseph Gilmore W. Soule M.D. 101 4 Jan 1945 Stormer Herbaceous, grassy meadow (50 feet wide) between high tide line WTU-155474 and spruce-hemlock forest. Much Empetrum nigrum, Vaccinium vitis-idea & Rubus Stellatus just below shrubs at forest ecotone. Puccinellia, Deschampsia and halophytic herbs under water at high Rosaceae tide. Phenology: Fruits. Rubus arcticus L. Mary Clay Muller 3954 2 Aug 1980 U.S.A., ALASKA COUNTY: WTU-282265 Akutan, Aleutian Island. 54.121589°, -165.894741°; uncertainty: 17000 m., Georef'd by Rosaceae Joseph Stormer Phenology: Fruits. Rubus arcticus L. John Rudd s.n. 1935 U.S.A., ALASKA COUNTY: Southeast. Chichagof Island. Port Frederick area. Humpback Creek WTU-101570 drainage. Elev. 450 ft. T43S R59E S24 Rosaceae Aspect 180; Slope 0-5 percent; high sedge muskeg adjacent to creek; growing with Carex sitchensis, Luzula campestris, Fauria Rubus arcticus L. crista-galli. Common berries red. Phenology: Fruits. U.S.A., ALASKA COUNTY: Kitty LaBounty 127 31 Jul 1981 Port Chalmers, Montague Island, Prince William Sound. Elev. 14 ft. WTU-282294 On gravel bars, tide, north exposure. Common. Corolla rose. Phenology: Flowers. Rosaceae J. David Solf 9 7 Jun 1959 Rubus arcticus L. WTU-232466 U.S.A., ALASKA, VALDEZ CORDOVA COUNTY: Middleton Island is located in the Gulf of Alaska about 115 miles south of Valdez. Near northern end of Island. 59° 25' N, 146° 20' W Partly disturbed area. Soil sandy and gravelly. At edge of undisturbed vegetation. Berries red. Phenology: Flowers. John H. Thomas 6363 3 Aug 1956 WTU-181775 Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA COUNTY: CANADA, YUKON TERRITORY: Attu Island; Sarana Bay. Northeast shoulder of Mount Sheldon and glacial cirque. 52.880963°, 173.306033°; uncertainty: 4600 m. Elev. 5000 ft. Top of rock cliff amongst deep moss with Rhododendron Phenology: Vegetative. Kamtschaticum. Phenology: Flowers. A. E. Porsild 11081 13 Aug 1944 Robert M. Hardy 176 10 Jul 1945 with A. J. Breitung WTU-102224 WTU-254234

Rosaceae Rosaceae Rubus arcticus L. Rubus arcticus L. U.S.A., ALASKA COUNTY: CANADA, YUKON TERRITORY: Popoff Island, Shumagin Islands. East of upper Rose River, opposite mile 95. West slope of granite Phenology: Flowers. range. Elev. 5000 ft. T. Kincaid s.n. 14 Jul 1899 Steep slope. In a summer-dry creek bed. Phenology: Flowers. WTU-101447 A. E. Porsild 10513 17 Jul 1944 with A. J. Breitung Rosaceae WTU-127246 Rubus arcticus L. Rosaceae U.S.A., ALASKA COUNTY: Muir Glacier. Rubus arcticus L. Phenology: Flowers. CANADA, YUKON TERRITORY: T. Kincaid s.n. 10 Jun 1899 Granite Mountain Range west of Upper Rose River, opposite mile 95. Alpine slopes and ravines from timberline (5000 feet) to summit WTU-101448 (6000 feet). Elev. 5000 ft. In alpine herbmats; elevation 5000 feet. Very fragrant. [Verbatium Rosaceae collection date: July 15-16, 1944]. Phenology: Flowers. Rubus arcticus L. A. E. Porsild 11941 15 Jul 1944 with A. J. Breitung U.S.A., ALASKA, ALEUTIANS WEST COUNTY: Dutch Harbor. WTU-254240 Moist and dry places. Indefinite loan by Tacoma High School to University of Washington. Phenology: Flowers. Rosaceae J. B. Flett 1820 17 Jun 1900 Rubus arcticus L. WTU-101458 CANADA, YUKON TERRITORY: Granite Mountain Range west of Upper Rose River, opposite mile Rosaceae 95. Alpine slopes and ravines from timberline (5000 feet) to summit (6000 feet). Rubus arcticus L. Elev. 5000 ft. In alpine herbmats; elevation 5000 feet. Very fragrant. [Verbatium U.S.A., ALASKA COUNTY: collection date: July 15-16, 1944]. Phenology: Flowers. Alaska Peninsula. Squaw Harbor. 55.234336°, -160.548263°; WGS 84, uncertainty: 2000 m., A. E. Porsild 11941 15 Jul 1944 Source: Digital Map, Georef'd by Joseph Stormer with A. J. Breitung Phenology: Flowers. WTU-254235 G. N. Jones 8899 14 Jul 1936 WTU-101658 Rosaceae Rosaceae Rubus arcticus L. Rubus parviflorus Nutt. CANADA, YUKON TERRITORY: U.S.A., MONTANA, SWEET GRASS COUNTY: Granite Mountain Range west of Upper Rose River, opposite mile By spring Half Moon Camp, Big Timber Creek, Crazy Mountains. 95. Alpine slopes and ravines from timberline (5000 feet) to summit In deep shade by spring. Phenology: Flowers. Origin: Native. (6000 feet). Elev. 5000 ft. C. L. Hitchcock 13198 4 Aug 1945 In alpine herbmats; elevation 5000 feet. Very fragrant. [Verbatium with C. V. Muhlick collection date: July 15-16, 1944]. Phenology: Flowers. WTU-102082 A. E. Porsild 11941 15 Jul 1944 with A. J. Breitung Rosaceae WTU-254235 Rubus parviflorus Nutt. Rosaceae U.S.A., OREGON, BENTON COUNTY: East foot of Alsea Mountain. Rubus parviflorus Nutt. 44.468878°, -123.488983°; WGS 84, uncertainty: 5000 m., Source: Digital map, Georef'd by Joseph Stormer U.S.A., MONTANA, PARK COUNTY: Phenology: Flowers. Origin: Native. Rock Island Lake, about 7 miles east of Cooke City. 45.015472°, -109.809738°; WGS 84, uncertainty: 1228 m., Garland M. Powell s.n. 27 May 1939 Source: GeoLocate, Georef'd by Joseph Stormer Talus slope, shore. Foliage fragrant. Phenology: Flowers. Origin: WTU-72685 Native. Jean G. Witt 1764 15 Jul 1949 Rosaceae WTU-175438 Rubus parviflorus Nutt. CANADA, BRITISH COLUMBIA: Rosaceae Parson's Bridge, V.I. B. C. Provincial Museum Victoria Herbarium. Phenology: Flowers. Rubus parviflorus Nutt. Origin: Native. U.S.A., ALASKA COUNTY: J. R. Anderson 1896 31 May 1956 Southeastern. Tongass National Forest. Admiralty Island, East side. West shore of Seymour Canal. North shore of Pleasant Bay. South WTU-182007 side of small forested peninsula. T49S R71E S10 NE 1/4 Beach meadow- forest ecotone. Phenology: Fruits. Origin: Native. Rosaceae Mary Clay Muller 4464 30 Jul 1981 Rubus parviflorus Nutt. WTU-289084 U.S.A., IDAHO, IDAHO COUNTY: 14 miles east of Kooskia along U.S. 12. 46.144627°, -115.684378°; WGS 84, uncertainty: 1000 m., Rosaceae Source: GeoLocate, Georef'd by WTU Staff Steep hillside with ponderosa pine, hemlock, and Douglas fir. Rubus parviflorus Nutt. Phenology: Flowers. Origin: Native. CANADA, BRITISH COLUMBIA: Gerrit Davidse 590 6 Jun 1966 1 mile west of Hope, B. C. with A. W. Collotzi Roadside, swampy. Alnus dominant. Phenology: Flowers. Origin: Native. WTU-238347 James Hatter 9 8 May 1949 Rosaceae WTU-143016 Rubus parviflorus Nutt. Rosaceae CANADA, BRITISH COLUMBIA: Sandspit, Moresby Island; Queen Charlotte Islands. Rubus parviflorus Nutt. Airport area. In thickets at edge of woods. Common. Flowers white; just in flower. Phenology: Flowers. Origin: Native. U.S.A., IDAHO COUNTY: Moscow Mountains. J. A. Calder 21100 1 Jun 1957 46.815368°, -116.830321°; WGS 84, uncertainty: 15000 m., with D. B. O. Savile, R. L. Taylor Source: Digital Map, Georef'd by Joseph Stormer In clearings. Common. Phenology: Flowers. Origin: Native. WTU-240954 F. L. Pickett 1507 20 May 1934 WTU-65951 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., MONTANA, RAVALLI COUNTY: CANADA, BRITISH COLUMBIA: Fifteen miles west of Skalkaho Road summit. Vancouver Island, near Iron Mine Creek. Drainage Quinsam River. Phenology: Flowers. Origin: Native. Elev. 1200 ft. In reforested area, exp.-n. slope-2. Thimbleberry. Phenology: Fruits. C. L. Hitchcock 14516 7 Jul 1946 Origin: Native. with C. V. Muhlick A. Young 900 31 Jul 1963 WTU-105334 with W. Hubbard WTU-218768 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., IDAHO, IDAHO COUNTY: Rubus parviflorus Nutt. Roadside, Frenchman Butte. 46.31611°, -115.53972°; WGS 84, uncertainty: 10000 m., Source: CANADA, BRITISH COLUMBIA: GeoLocate, Georef'd by WTU Staff Interurban road near Victoria. Drainage Colquits Creek. Spruce-fir, near-climax, open forest. Dominants: Picea engelmannii, Elev. 260 ft. Abies grandis. Phenology: Flowers. Origin: Native. In grass under alder. Exp.-shade, slope-5 elev. Flowers white. Phenology: Flowers. Origin: Native. R. F. Daubenmire 4562 6 Jul 1945 W. Hubbard 115 20 May 1963 WTU-239886 WTU-221358 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., IDAHO, CLEARWATER COUNTY: Several miles west of Elk River. U.S.A., MONTANA COUNTY: T40N R2E; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Rocky Mountains; Echo Lake; in the Flathead National Forest. Georef'd by WTU Staff Elev. 3500 ft. Second-growth coniferous woods. Shrub 0.5-1 m. tall; flowers white. Flower white; common. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. C. H. Kauffman s.n. 23 Jun 1928 Arthur Cronquist 5885 22 Jun 1949 with G. B. Cummins WTU-180678 WTU-251031

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. CANADA, BRITISH COLUMBIA: CANADA, BRITISH COLUMBIA: 3 miles south of Fernie on Old Morrissey Road. Big Bend Highway. 49° 27' N, 115° 3' W On cliffs. Phenology: Flowers. Origin: Native. In semi-open mixed woods on hillside. Common. Phenology: Flowers. Origin: Native. A. E. Porsild 16923 23 Jun 1948 R. L. Taylor 872 28 May 1958 WTU-127314 with D. H. Ferguson WTU-215524 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., OREGON, WASHINGTON COUNTY: Rubus parviflorus Nutt. Near Forest Grove. 45.519836°, -123.110663°; WGS 84, uncertainty: 7500 m., CANADA, BRITISH COLUMBIA: Source: GeoLocate, Georef'd by Joseph Stormer Edgewood, Lower Arrow Lake. Along creek bottoms. Phenology: Flowers. Origin: Native. 49.782003°, -118.154276°; WGS 84, uncertainty: 4959 m., Source: Digital Map, Georef'd by Joseph Stormer J. William Thompson 569 29 Mar 1926 In woods. Common. Phenology: Vegetative. Origin: Native. WTU-23016JWT M. Ridewood 60 25 Jun 1960 WTU-196952 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., OREGON, CLATSOP COUNTY: U.S.A., IDAHO, BEAR LAKE COUNTY: Near Astoria. Cache National Forest, Highway 36 north up Strawberry Canyon, 46.187884°, -123.831253°; WGS 84, uncertainty: 7090 m., then south cira 0.7 mile on Copenhagen Basin Road. Source: GeoLocate, Georef'd by Joseph Stormer Elev. 7450 ft. Phenology: Flowers. Origin: Native. 42° 21' 27" N, 111° 33' 15" W; WGS 84, uncertainty: 500 m., Source: GPS, Georef'd by John Bassett G. E. Morrill 33 17 Jun 1933 Steep west-facing slope, with Douglas Fir and Aspen, rich loamy soil. Associated species: Ceanothus velutinus, Erysimum asperum, WTU-27743 Senecio streptanthifolius, Symphoricarpos oreophilus. . Phenology: Flowers. Origin: Native. Rosaceae Sandy Long 40 12 Jul 1999 Rubus parviflorus Nutt. WTU-340226 U.S.A., IDAHO, IDAHO COUNTY: Near Powell. Rosaceae Phenology: Flowers. Origin: Native. Rubus parvifolius L. C. L. Hitchcock 14695 13 Jul 1946 with C. V. Muhlick U.S.A., OREGON, JOSEPHINE COUNTY: Forest Service Road 4611 at Mossback Creek, Siskiyou National WTU-105332 Forest. Elev. 2548 ft. 42° 9' 8" N, 123° 24' 7" W; T39S R6W S27 Rosaceae Shady streamside. Phenology: Flowers. Rubus parviflorus Nutt. Jessie Johanson 18 26 Jun 1998 with Joe Johanson U.S.A., IDAHO, BOISE COUNTY: Summit of Boise-Lowman Highway. WTU-342666 Elev. 6300 ft. 44.08361°, -115.61972°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Rosaceae In fir forest. Phenology: Flowers. Origin: Native. Rubus parviflorus Nutt. C. L. Hitchcock 9980 16 Jul 1944 with C. V. Muhlick U.S.A., ALASKA COUNTY: Washington Bay, Kuiu Island. WTU-91507 56.721111°, -134.356907°; WGS 84, uncertainty: 2400 m., Source: Digital Map, Georef'd by Joseph Stormer On border of spruce forest. Phenology: Fruits. Origin: Native. Rosaceae W. J. Eyerdam 5445 20 Aug 1947 Rubus parviflorus Nutt. WTU-292225 U.S.A., IDAHO, FRANKLIN COUNTY: Caribou National Forest, Highway 36 north east of Preston to Copenhagen Basin Road approximately 3 miles up toward Rosaceae Copenhagen Basin. Elev. 8388 ft. Rubus parviflorus Nutt. 42° 20.43' N, 111° 34.28' W North facing slope with rich loam soil. Moist site with a heavy mixed U.S.A., IDAHO, CUSTER COUNTY: conifer forest overstory. Associated with Pseudotsuga menziesii, Along the Yankee fork of the Salmon River at Blind Creek Forest Vaccinium membranaceum, and Paxistima myrsinites. Phenology: Camp, 1 mile north of Sunbeam. Fruits. Origin: Native. 44.283026°, -114.735105°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff Allyson Davis 194 12 Sep 1997 Phenology: Flowers. Origin: Native. with Andreas Leidolf William H. Baker 10775 29 Jul 1953 WTU-336460 WTU-295927

Rosaceae Rubus parviflorus Nutt. U.S.A., IDAHO, IDAHO COUNTY: Along the Selway River at Rackliff Forest Camp. 46.124909°, -115.487754°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Moist shady wood. Phenology: Flowers. Origin: Native. William H. Baker 14821 30 May 1957 WTU-295814 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., IDAHO, SHOSHONE COUNTY: U.S.A., IDAHO, BONNER COUNTY: About 1 mile below summit of Bullion Pass. Luby Bay, Priest Lake. Stream bank. Phenology: Flowers. Origin: Native. 48.54222°, -116.91722°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff William H. Baker 13463 15 Jul 1955 Deep coniferous woods. Phenology: Flowers. Origin: Native. WTU-295861 William H. Baker 14953 30 Jun 1957 WTU-295839 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., IDAHO, BENEWAH COUNTY: Rubus parviflorus Nutt. Junction of Skyline Drive and Highway 95. T43N R4W S20; NAD 27, uncertainty: 805 m., Source: TRS2LL, U.S.A., IDAHO, BONNER COUNTY: Georef'd by Ben Legler Along Priest River, 3 miles south of outlet to Priest Lake. Woods. Phenology: Flowers. Origin: Native. 48.446432°, -116.905153°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff William H. Baker 15859 9 Jun 1960 Woods. Phenology: Flowers. Origin: Native. WTU-295863 William H. Baker 15625 5 Jul 1959 WTU-295840 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., IDAHO, BEAR LAKE COUNTY: Rubus sachalinensis H. Lév. var. sachalinensis Above the Saint Charles Creek, Porcupine Campgrounds. 42.11389°, -111.38806°; WGS 84, uncertainty: 10000 m., Source: CANADA, BRITISH COLUMBIA: GeoLocate, Georef'd by WTU Staff Upper Hat Creek, Marble Mountains. Moist, brushy slopes. Phenology: Flowers. Origin: Native. Elev. 3000 ft. 50.817329°, -121.577703°; uncertainty: 16000 m. E. J. Larrison 313 13 Jul 1950 Arid slopes. Phenology: Fruits. Origin: Native. WTU-295865 J. William Thompson 507 18 Jul 1938 with Emily M. Thompson Rosaceae WTU-47803JWT Rubus parviflorus Nutt. Rosaceae U.S.A., IDAHO, BENEWAH COUNTY: Rocky Point, Lake Chatcolet. Rubus parviflorus Nutt. 47.37222°, -116.7625°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., IDAHO, IDAHO COUNTY: Wooded shores. Phenology: Flowers. Origin: Native. Along Shingle Creek, Seven Devils Mountains. 45.371608°, -116.484724°; WGS 84, uncertainty: 10000 m., William H. Baker 15819 31 May 1960 Source: GeoLocate, Georef'd by WTU Staff Moist woods. Phenology: Flowers. Origin: Native. WTU-295868 William H. Baker 12257 31 Jul 1954 Rosaceae WTU-295674 Rubus parviflorus Nutt. Rosaceae U.S.A., IDAHO, IDAHO COUNTY: Fog Mountain, Selway Mountains. Rubus parviflorus Nutt. 46.09333°, -115.23556°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., IDAHO, LATAH COUNTY: Thickets. Phenology: Flowers. Origin: Native. North slope of Tomer's Butte, 4 miles southeast of Moscow. 46.691538°, -116.939429°; WGS 84, uncertainty: 500 m., Source: William H. Baker 15273 13 Jul 1958 GeoLocate, Georef'd by WTU Staff Woods. Phenology: Flowers. Origin: Native. WTU-295838 William H. Baker 9009 3 Jun 1952 WTU-295711 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., IDAHO, LATAH COUNTY: U.S.A., WASHINGTON, THURSTON COUNTY: Northeastern slope of Tomer's Butte. Four and a half miles Olympia. southeast of Moscow. 47.03806°, -122.89944°; WGS 84, uncertainty: 10000 m., Source: T38N R5W S23; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Open hillsides and fields. A fine berry, by many preferred to the Phenology: Flowers. Origin: Native. cultivated. Three specimens on one sheet from Olympia and from Orcas Island, Washington. Phenology: Flowers. Origin: Native. R. L. Nisbet 87 23 Jul 1955 Louis F. Henderson s.n. 24 May 1892 WTU-295867 WTU-9684A Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus leucodermis Douglas ex Torr. & A. Gray CANADA, BRITISH COLUMBIA: Taylor River area, Vancouver Island. Taylor River drainage. U.S.A., WASHINGTON, SAN JUAN COUNTY: Elev. 440 ft. East Sound, Orcas Island. In sandy area along river, exp.- total slope-0-5 elev.- 440 ft. Pollen 48.6925°, -122.904167°; WGS 84, uncertainty: 800 m., Source: Collected Q R C 18 Sept. 80 RN. Phenology: Flowers. Origin: Georeferenced, Georef'd by WTU Staff Native. Open hillsides and fields. A fine berry, by many preferred to the cultivated. Three specimens on one sheet from Olympia and from A. Young 220 30 May 1963 Orcas Island, Washington. Phenology: Fruits. Origin: Native. with W. Hubbard Louis F. Henderson s.n. 3 Jul 1892 WTU-218584 WTU-9684B Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., OREGON, CLATSOP COUNTY: Rubus parviflorus Nutt. Near seaside. 45.993164°, -123.922638°; WGS 84, uncertainty: 4780 m., U.S.A., WASHINGTON, THURSTON COUNTY: Source: GeoLocate, Georef'd by Joseph Stormer Olympia. Phenology: Flowers. Origin: Native. 47.03806°, -122.89944°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff G. E. Morrill 124 27 Jun 1933 Rich, sunny hillsides. A rich acid and agreeable berry. Several specimens on one sheet from Olympia and from Sumas City, WTU-27975 Washington. Phenology: Flowers. Origin: Native. Louis F. Henderson s.n. 24 May 1892 Rosaceae WTU-9686A Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: Rosaceae Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Rubus parviflorus Nutt. Source: Digital Map, Georef'd by Joseph Stormer Campus woods. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, WHATCOM COUNTY: Sumas City. G. L. Wittrock s.n. 1 Apr 1928 49.00028°, -122.26361°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-361804 Just on northern border of the United States. A rich acid and agreeable berry. Several specimens on one sheet from Olympia and from Sumas City, Washington. Phenology: Flowers. Origin: Native. Rosaceae Louis F. Henderson s.n. 2 Jul 1892 Rubus lasiococcus A. Gray WTU-9686B U.S.A., WASHINGTON, KITTITAS COUNTY: Above the tunnel on the Northern Pacific Railroad at Stampede Pass. Rosaceae 47.27694°, -121.3375°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Rubus pedatus Sm. Common with Rubus pedatus on shady hillsides, Cascade Mountains. Phenology: Flowers & Fruits. Origin: Native. U.S.A., WASHINGTON, KITTITAS COUNTY: Stampede Pass, above the tunnel for the Northern Pacific Railroad. Louis F. Henderson s.n. 20 Jun 1892 47.27694°, -121.3375°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-9674 Common in deep fir woods; Cascade Mountains. Phenology: Flowers & Fruits. Origin: Native. Louis F. Henderson s.n. 20 Jun 1892 WTU-9675 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, THURSTON COUNTY: U.S.A., OREGON, DOUGLAS COUNTY: Olympia. Roseburg quadrangle. 47.03806°, -122.89944°; WGS 84, uncertainty: 10000 m., Source: 43.25°, -123.5°; uncertainty: 49000 m., Georef'd by Joseph GeoLocate, Georef'd by WTU Staff Stormer Dark, rich woods along streams. Abundant throughout Western A trailing shrub. Phenology: Flowers. Origin: Native. Washington, especially toward the coast. The fruit large, but watery. Phenology: Flowers & Fruits. Origin: Native. W. C. Cusick 3876 9 Apr 1914 Louis F. Henderson s.n. 25 Mar 1892 WTU-183981 WTU-9685 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., WASHINGTON, LEWIS COUNTY: Rubus ursinus Cham. & Schltdl. Shoulder of Highway 7 about 200 feet south of junction with Pleasant Valley Road, about 2 miles south of county line. U.S.A., WASHINGTON, THURSTON COUNTY: Elev. 1280 ft. Olympia. 46° 43.986' N, 122° 12.497' W; WGS 84, uncertainty: 6 m., 47.03806°, -122.89944°; WGS 84, uncertainty: 10000 m., Source: Source: GPS GeoLocate, Georef'd by WTU Staff Gravelly weedy roadside pullout and road shoulders, with Phalaris Open, burnt ground. A fine berry, preferred to the cultivated arundinacea, Leucanthemum valgare. Petals white; shrubs to 1.5 blackberry for pies and preserves. Two specimens on one sheet meters tall; scattered. Phenology: Flowers. Origin: Native. from Olympia and from Lakeview, Washington. Phenology: Flowers. Origin: Native. Jessica Ni 16 16 Jun 2006 with Ben Legler Louis F. Henderson s.n. 20 May 1892 WTU-362886 WTU-9687A Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, LEWIS COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Logging road on west side of Highway 97 between Carlson and Lakeview. Divide, circa 10 road miles north of Morton. 47.15944°, -122.49694°; WGS 84, uncertainty: 10000 m., Source: 46° 40' 55" N, 122° 11' 54" W; WGS 84, uncertainty: 1000 m., GeoLocate, Georef'd by WTU Staff Source: Digital map A fine berry, preferred to the cultivated blackberry for pies and Petals white; stems trailing. Phenology: Flowers. Origin: Native. preserves. Two specimens on one sheet from Olympia and from Lakeview, Washington. Phenology: Fruits. Origin: Native. Jessica Ni 34 16 Jun 2006 with Ben Legler Louis F. Henderson s.n. 25 Jul 1892 WTU-362917 WTU-9687B Rosaceae Rosaceae Rubus bifrons Vest Rubus parviflorus Nutt. U.S.A., WASHINGTON, LEWIS COUNTY: U.S.A., WASHINGTON, PEND OREILLE Entry road to Kosmos Boat Launch on northeast corner of Riffe COUNTY: Lake; 5 miles southeast of Morton. 17 air miles northeast of Colville, along FS Road Elev. 800 ft. 4668 along Rocky Creek 0.6 miles east of 46° 30' 7" N, 122° 11' 31" W; WGS 84, uncertainty: 300 m., junction with Rocky Creek Road. Source: Digital map Elev. 3060 ft. Exposed dirt road banks with Rubus discolor, Holcus lanatus, and 48° 40' 35" N, 117° 36' 8" W; NAD 83 other weeds. Petals white with faint pink tinge; abundant Weedy roadside and gravel bars along creek. thicket-forming shrub. Phenology: Flowers. Origin: Introduced. Shrubs to 1.5 meters tall; berries turning red. Phenology: Fruits. Origin: Native. Jessica Ni 49 16 Jun 2006 with Ben Legler Ben Legler 2114 WTU-362898 24 Jul 2004 WTU-382257, WTU-V-045455 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, LEWIS COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Entry road to Kosmos Boat Launch on northeast corner of Riffe San Juan Islands. Pole Island. Tiny island in the middle of Mosquito Lake; 5 miles southeast of Morton. Pass. Elev. 800 ft. 48° 36.056' N, 123° 10.096' W 46° 30' 7" N, 122° 11' 31" W; WGS 84, uncertainty: 300 m., Rim of beach with Distichlis especially on west and south sides. Source: Digital map Occasional trailing in meadow. Phenology: Flowers. Origin: Native. Exposed dirt road banks with Rubus discolor, Holcus lanatus, and other weeds. Arching shrub to 1.5 meters tall; uncommon. Eliza Habegger EH-1018 26 Apr 2006 Phenology: Fruits. Origin: Native. WTU-385183 Jessica Ni 50 16 Jun 2006 with Ben Legler Rosaceae WTU-362890 Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, CHELAN COUNTY: Lake Chelan National Recreation Area; Rubus ursinus Cham. & Schltdl. Northeast side of Coon Lake where creek enters lake. One kilometer northeast of High Bridge in U.S.A., WASHINGTON, SAN JUAN COUNTY: Stehekin River Valley. San Juan Islands. Barren Island. Elev. 2170 ft. Elev. 30 ft. 48° 23' 26.8" N, 120° 49' 57.5" W; UTM Zone 48° 37.366' N, 123° 9.361' W 10, 660447E, 5362004N; WGS 84, Bedrock dome rising 30 feet above the water. Lushly covered with uncertainty: 8 m., Source: GPS Camas and Plectritis. Uncommon. Phenology: Flowers. Origin: Moist deciduous forest and brushy thickets, with Native. Acer macrophyllum, Alnus sp., Cornus sericea, Rubus parviflorus, Viola sp. Fruits orange. Peter Dunwiddie 06-12 26 Apr 2006 Shrubs to 2 meters tall. Uncommon. Leaf material removed for RADseq analysis by WTU-385599 Megan Ruffley at WTU, February 2016. Phenology: Fruits. Origin: Native. Rosaceae Barbara Smith 06-27 29 Jun 2006 Rubus parviflorus Nutt. WTU-366875, WTU-V-019100 U.S.A., WASHINGTON, CHELAN COUNTY: Lake Chelan National Recreational Area. Stehekin State Airstrip. Elev. 1196 ft. Rosaceae 48.34332° N, 120.71854° W; UTM Zone 10, 669049.268E, 5356975.232N; WGS 83, uncertainty: 5 m., Source: GPS, UTM Rubus bifrons Vest calculated from Lat-Lon. Flat mowed area approximately 3,000 feet long by 300 feet wide. U.S.A., WASHINGTON, SNOHOMISH COUNTY: Public airstrip cut out of valley floor forest. Abies grandis, Alnus Stillaquamish River; Washington Department of Fish and Wildlife rubra, Acer macrophylla, Pseudotsuga menziesii dominate forest boat launch on 236th Street N.W. About 0.1 miles west of Marine edge. Airstrip dominated by non-native grasses. Flowers white. View Drive Bridge over river. Common along forest edge. Phenology: Flowers. Origin: Native. 48.21167°, -122.33967°; WGS 84, uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff Dale Blum 06-16 27 Jun 2006 Levee; Populus and Alnus overstory. Forming monoculture on top of levee. Phenology: Fruits. Origin: Introduced. WTU-366588 David Giblin 273 23 Sep 2005 with Ben Legler Rosaceae WTU-364321 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, CHELAN COUNTY: Rosaceae Lake Chelan National Recreational Area. Stehekin State Airstrip. Elev. 1196 ft. Rubus lasiococcus A. Gray 48.34332° N, 120.71854° W; UTM Zone 10, 669049.268E, 5356975.232N; WGS 83, uncertainty: 5 m., Source: GPS, UTM U.S.A., WASHINGTON, CHELAN COUNTY: calculated from Lat-Lon. North Cascades National Park. North side of Hock Mountain, on the Flat mowed area approximately 3,000 feet long by 300 feet wide. east side of the large boulder strewn moraine/tongue; 0.75 mile Public airstrip cut out of valley floor forest. Abies grandis, Alnus northeast of summit. rubra, Acer macrophylla, Pseudotsuga menziesii dominate forest Elev. 5600-5800 ft. edge. Airstrip dominated by non-native grasses. Stems trailing. 48° 27' 45.87137" N, 120° 39' 43.66343" W; UTM Zone 10, Phenology: Fruits. Origin: Native. 672829.16E, 5370155.2N; WGS 84, Source: Calc. from UTM Moraine composed of huge boulders interspersed with pockets of Dale Blum 06-52 27 Jun 2006 soil supporting vegetation. Dominant: tree is Larix lyallii, shrub - Phyllodoce empetriformis, and graminoid - Carex rossii. Very dry; WTU-366717 lots of cold air is channeled down from high elevation to moraine. Aspect: 90°; slope: 0-15°. Common under Larix lyallii; plants are trailing and prostrate. Phenology: Flowers. Origin: Native. Ben Legler 3539 30 Jul 2006 WTU-366957 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Mount Rainier National Park. Ipsut Pass circa 1 mile north of Mowich San Juan Islands. Ripple Island. Lake. 48° 39.415' N, 123° 7.896' W Elev. 5150 ft. Oblong (north-south) island with extensive shrub cover and a stand 46° 57.039' N, 121° 52.09' W; UTM Zone 10, 586127.103E, of mostly dead Populus tremuloides. Large shell and gravel beach 5200301.74N; WGS 84, uncertainty: 7 m., Source: GPS, UTM on west side, with broad shelving bedrock bench draped with calculated from Lat-Lon. seaweed. Frequent among shrubs and on shoreline bluffs. Steep northeast-facing brushy cirque with Rubus, Sorbus, Valeriana, Phenology: Flowers. Origin: Native. Veratrum, topped by steep dry rock faces and subalpine conifer forest. Petals white; common shrubs to 1+ meter tall on open slope. Rod Gilbert 06-69 27 Apr 2006 Phenology: Flowers. Origin: Native. WTU-387763 Lucia Harrison 06-1 8 Aug 2006 WTU-366666 Rosaceae Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: Rubus spectabilis Pursh San Juan Islands. Gossip Island. Elev. 0-25 ft. U.S.A., WASHINGTON, PIERCE COUNTY: 48° 39.789' N, 123° 10.374' W Mount Rainier National Park. Ipsut Pass circa 1 mile north of Mowich Low conglomerate bedrock island with gently sloping bedrock on all Lake. sides. Scattered madrone, Douglas fir, and junipers across island Elev. 5150 ft. with lush camas on swales between outcrop. Occasional, trailing 46° 57.039' N, 121° 52.09' W; UTM Zone 10, 586127.103E, over other low shrubs. Phenology: Flowers. Origin: Native. 5200301.74N; WGS 84, uncertainty: 7 m., Source: GPS, UTM calculated from Lat-Lon. Steve Hahn 06-35 27 Apr 2006 Steep northeast-facing brushy cirque with Rubus, Sorbus, Valeriana, Veratrum, topped by steep dry rock faces and subalpine conifer WTU-383661 forest. Fruits orange; common shrub to 1.5 meters tall on brushy slope. Phenology: Fruits. Origin: Native. Rosaceae Lucia Harrison 06-16 8 Aug 2006 Rubus parviflorus Nutt. WTU-366698 U.S.A., IDAHO, CLEARWATER COUNTY: Clearwater National Forest. 0.2 miles northeast of Mush Saddle. Rosaceae Forest Service Road 711. Elev. 5287 ft. Rubus idaeus L. ssp. strigosus (Michx.) Focke 46.75606°, -115.3646°; WGS 84, uncertainty: 500 m., Source: Nat. Geog. TOPO!, Georef'd by John Bassett U.S.A., IDAHO, CLEARWATER COUNTY: Southeast facing rocky roadcut. Open forest and brush. Uncommon. Clearwater National Forest. Hidden Creek Camp. Phenology: Flowers. Origin: Native. Elev. 3410 ft. 46.83177°, -115.17851°; WGS 84, uncertainty: 500 m., Source: Wendy McClure 05-56 10 Jul 2005 Nat. Geog. TOPO!, Georef'd by John Bassett with Suzanne Bagshaw, Ben Legler, Don Knoke, Jessie Johanson, Level damp soil; few rocks. With Thuja plicata, Symphoricarpos alba, Joe Johanson, Mary Johanson Trillium ovatum, Acer glabrum. Young fruit. Uncommon. Phenology: Flowers. Origin: Native. WTU-365694 Don Knoke 880 9 Jul 2005 Rosaceae WTU-375262 Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Flattop Island. Weed Point on northeast end of Rubus ursinus Cham. & Schltdl. Flattop Island; northern part of San Juan Archipelago. Elev. 0-10 ft. U.S.A., WASHINGTON, SAN JUAN COUNTY: 48° 38.871' N, 123° 4.707' W; WGS 84, uncertainty: 2 m. San Juan Islands. Battleship Island. Rocky open point, treeless, with thin soil, with Rosa nutkana, Elev. 0-40 ft. Grindelia, Lathyrus japonicus, grasses. Stems trailing, in open areas. 48° 37.494' N, 123° 11.115' W Phenology: Flowers. Origin: Native. Five acre rock outcrop rising 40 feet above water, with shrub thickets, a few Pseudotsuga and Madrona trees, lush camas swales Carrie Marschner 06-22 31 May 2006 and bedrock outcrops. One plant; trailing; white flowers. Phenology: Flowers. Origin: Native. WTU-387836 Rod Gilbert 06-34 26 Apr 2006 WTU-383644 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus bifrons Vest U.S.A., IDAHO, CLEARWATER COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Clearwater National Forest. Toboggan Hill Road. 9.9 miles south of San Juan Islands. Skull Island. Cayuse Creek crossing on Forest Road 581. Elev. 40 ft. Elev. 5579 ft. 48° 38.343' N, 122° 59.168' W; WGS 84, T10N R12E S4; WGS 84, uncertainty: 1000 m., Source: Nat. Source: GPS Geog. TOPO!, Georef'd by John Bassett A 40 foot high island with many Juniperus trees, Lodgepole pine woodland with Vaccinium and Xerophyllum. White abundant Camassia leichtlinii and Plectritis flowered shrub to 1 meter. Phenology: Flowers. Origin: Native. congesta. Numerous bedrock outcrops with ledges. Large patch of shrubs on west side of Richard G. Olmstead 2005-15 8 Jul 2005 island. Phenology: Flowers. Origin: Introduced. WTU-365595 Pam Camp 06-12 30 May 2006 Rosaceae WTU-388705, WTU-V-060737 Rubus parviflorus Nutt. U.S.A., MONTANA, MINERAL COUNTY: Lolo National Forest. Heart Lake Trailhead off Forest Service Road Rosaceae 250. East side of road. T14N R27W S11 SW 1/4; WGS 84, uncertainty: 500 m., Source: Rubus ursinus Cham. & Schltdl. Digital map, Georef'd by John Bassett Open coniferous forest dominated by Picea. Associated with U.S.A., WASHINGTON, SAN JUAN COUNTY: Senecio, Castilleja, Rubus, Penstemon. Phenology: Flowers. Origin: San Juan Islands. Skull Island. Native. Elev. 40 ft. 48° 38.343' N, 122° 59.168' W; WGS 84, Source: GPS Richard Robohm 05-06 9 Jul 2005 A 40 foot high island with many Juniperus trees, abundant Camassia with David Giblin leichtlinii and Plectritis congesta. Numerous bedrock outcrops with ledges. Several plants trailing across open grassland. Phenology: WTU-369561 Flowers & Fruits. Origin: Native. Rosaceae Pam Camp 06-59 30 May 2006 Rubus bifrons Vest WTU-388745 U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Flattop Island. South side of Rosaceae Flattop Island; north end of San Juan Archipelago. Rubus ursinus Cham. & Schltdl. Elev. 60 ft. 48° 38.752' N, 123° 4.953' W; WGS 84, U.S.A., WASHINGTON, SAN JUAN COUNTY: uncertainty: 5 m. San Juan Islands. Oak Island. Dry open forest of Pseudotsuga and Arbutus, Elev. 10 ft. with dry smooth rock balds and small soil 48° 36.463' N, 122° 57.283' W; WGS 84, Source: GPS pockets. Abundant, in large patches, flowers Very small, 0.19 acre (measured) island with Holodiscus/Rosa white. Phenology: Flowers. Origin: Introduced. shrubs, Sanicula understory; one goose nest; otter evident abundantly; 3 Pseudotsuga menziesii, 2 Juniperus scopulorum, and Carrie Marschner 06-93 1 Quercus garryana. A couple stems clambering across shrubs. Phenology: Flowers. Origin: Native. 31 May 2006 Peter Dunwiddie 06-195 30 May 2006 WTU-388908, WTU-V-060736 WTU-389811

Rosaceae Rosaceae Rubus laciniatus Willd. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Flattop Island. South side of Flattop Island; north San Juan Islands. McConnell Island off southwest side of Orcas end of San Juan Archipelago. Island; north end of island near small wetland. Elev. 60 ft. Elev. 5 ft. 48° 38.752' N, 123° 4.953' W; WGS 84, uncertainty: 5 m. 48° 35.902' N, 123° 1.444' W; WGS 84, uncertainty: 5 m., Source: Dry open forest of Pseudotsuga and Arbutus, with dry smooth rock GPS balds and small soil pockets. Flowers white; common. Phenology: Small rocky bald by wetland; mostly grassy with scattered Flowers. Origin: Introduced. Pseudotsuga, Juniperus, and Quercus. Petals white; stems trailing; in moister areas. Phenology: Flowers. Origin: Native. Carrie Marschner 06-99 31 May 2006 Victoria Wyllie-Echeverria 06-36 30 May 2006 WTU-388914 WTU-390124 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Iceberg Island. San Juan Islands. Long Island National Wildlife Refuge. Unnamed Elev. 0-30 ft. Island #1, off the south shore of Long Island. 48° 25.625' N, 122° 53.354' W; WGS 84, Source: GPS Elev. 6-16 ft. Island is low, relatively flat, with extensive bedrock outcrops 48° 26.38' N, 122° 55.103' W; Source: GPS throughout. Sedum lanceolatum is abundant across the bedrock. A Smallest unnamed island by Long Island, connected by a gravel bar large swale in mid-island is dense Rosa nutkana thicket. A couple of at low tide; 0.76 acres (measured with GPS). A few plants trailing small vernal pools in the bedrock near the south side are over open ground with Grindelia stricta. Phenology: Vegetative. noteworthy. Occasional, trailing, with Solidago canadensis. Origin: Native. Phenology: Fruits. Origin: Native. Tara Martin 06-01 6 Sep 2006 Joe Bennett 06-50 6 Sep 2006 WTU-389223 WTU-392758 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: 0.1 miles up Swakane Canyon Road from San Juan Islands. Posey Island. Highway Alternate 97. Elev. 0-20 ft. Elev. 852 ft. 48° 37.104' N, 123° 10.052' W 47° 32.769' N, 120° 17.607' W; NAD 27, Low, one acre island with campsites, outhouse, fire pits. Heavily Source: GPS used, with deer grating. Mostly highly disturbed. Trailing in shoreline Dry steep loose slopes around brushy seasonal area; a couple of plants. Phenology: Vegetative. Origin: Native. creek. Petals white; shrub to 1 meter tall; in large patches by creek. Phenology: Flowers. John Floberg 06-19 7 Sep 2006 Origin: Native. WTU-390088 Ben Legler 2651 20 May 2005 Rosaceae WTU-374080, WTU-V-045437 Rubus bifrons Vest U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Battleship Island. Rosaceae Elev. 0-40 ft. 48° 37.494' N, 123° 11.115' W Rubus leucodermis Douglas ex Torr. & A. Gray Five acre rock outcrop rising 40 feet above water, with shrub thickets, a few Pseudotsuga and Madrona trees, lush camas swales U.S.A., WASHINGTON, CHELAN COUNTY: and bedrock outcrops. Several large plants among shrubs (Rosa, 0.1 miles up Swakane Canyon Road from Highway Alternate 97. Symphoricarpos). Phenology: Fruits. Origin: Introduced. Elev. 852 ft. 47° 32.769' N, 120° 17.607' W; NAD 27, Source: GPS Pam Camp 06-102B 7 Sep 2006 Dry steep loose slopes around brushy seasonal creek. Stems glaucous, arching shrub; uncommon by creek. Phenology: Flowers. WTU-389977 Origin: Native. Ben Legler 2652 20 May 2005 Rosaceae WTU-374079 Rubus bifrons Vest U.S.A., WASHINGTON, SAN JUAN COUNTY: Rosaceae San Juan Islands. Deadman Island. Elev. 9-32 ft. Rubus vestitus Weihe & Nees 48° 27.5' N, 122° 56.629' W Bedrock knob ca. [circa] 10 meters tall, with numerous crevices, U.S.A., WASHINGTON, GRANT COUNTY: several shrub thickets. One patch with Sambucus racemosa, Quincy Lakes Wildlife Area, about 0.15 miles north of Stan Coffin Crataegus monogyna. Phenology: Fruits. Origin: Introduced. Lake. Elev. 1210 ft. Phil Green 06-48 6 Sep 2006 47° 9.223' N, 119° 55.075' W; Source: Field notes Small draw, below basalt ledges. Smothering creek for 50 meters; WTU-394541 evergreen; glandular. Phenology: Fruits. Origin: Introduced. Ben Legler 2501 23 Apr 2005 with Peter F. Zika, David Giblin WTU-369701 Rosaceae Rosaceae Rubus bifrons Vest Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, GRANT COUNTY: U.S.A., WASHINGTON, SKAGIT COUNTY: Base of Vantage Road, east bank of the Padilla Bay National Estuarine Research Reserve. Saddlebag Columbia River. Below Babcock Bench. Island. Elev. 557 ft. Elev. 17 ft. 47° 0.009' N, 119° 59.339' W; Source: Field 48° 32.117' N, 122° 33.333' W; WGS 84, Source: GPS notes High island with rocky shoreline cliffs; Douglas fir/Madrone forests Full sun, riparian, and immediately adjacent with Snowberry, Mahonia. South bank of island, above beach. upland; basalt bedrock. Evergreen shrub to 2 Common trailing up bank. Phenology: Flowers. Origin: Native. meters tall. Phenology: Fertile. Origin: Introduced. Peter Dunwiddie 06-137 23 May 2006 Ben Legler 2507 WTU-371446 with Peter F. Zika, David Giblin Rosaceae 23 Apr 2005 Rubus parviflorus Nutt. WTU-369696, WTU-V-060718 U.S.A., WASHINGTON, SNOHOMISH COUNTY: At the Legler residence (4115 234th St. NE) ca. 1.7 air miles NW of Rosaceae Arlington. Elev. 290 ft. Rubus bifrons Vest 48° 12' 28" N, 122° 10' 25" W; WGS 84, uncertainty: 100 m., Source: Digital map U.S.A., WASHINGTON, PACIFIC COUNTY: Mixed conifer and deciduous forest with brushy understory. Shrubs North side of Pioneer Road, 1 air kilometer east to 1.5 meters tall; twigs brown. Phenology: Buds. Origin: Native. of Clear Lake. Elev. 16 ft. Ben Legler 3143 2 Feb 2006 46° 22.1' N, 124° 1.9' W; T10N R11W S10 Irrigation pond and adjacent agricultural fields of Herbarium: WTU research cranberry station, full sun, damp ground. Leaves green below; 0.5 meter; arching over cranberry field; adventive. Phenology: Rosaceae Fruits. Origin: Introduced. Rubus spectabilis Pursh Peter F. Zika 22272 with David Giblin, Cathy Maxwell, Ben Legler U.S.A., WASHINGTON, SNOHOMISH COUNTY: At the Legler residence (4115 234th St. NE) ca. 1.7 air miles NW of 30 Aug 2005 Arlington. Elev. 290 ft. WTU-373949, WTU-V-060717 48° 12' 28" N, 122° 10' 25" W; WGS 84, uncertainty: 100 m., Source: Digital map Mixed conifer and deciduous forest with brushy understory. Common shrubs to over 2 meters tall dominating wetter areas of forest Rosaceae understory; twigs and bark light brown, peeling. Phenology: Buds. Origin: Native. Rubus ursinus Cham. & Schltdl. Ben Legler 3144 2 Feb 2006 U.S.A., WASHINGTON, JEFFERSON COUNTY: Protection Island. Herbarium: WTU Elev. 178 ft. 48° 7.733' N, 122° 55.692' W A tiny remnant native prairie near the forest. Collections from both Rosaceae habitats. Occasional under shrubs at forest edge; petals white. Phenology: Flowers. Origin: Native. Rubus leucodermis Douglas ex Torr. & A. Gray Fred Weinmann 06-1003 3 May 2006 U.S.A., WASHINGTON, SNOHOMISH COUNTY: At the Legler residence (4115 234th St. NE) ca. 1.7 air miles NW of WTU-371087 Arlington. Elev. 290 ft. 48° 12' 28" N, 122° 10' 25" W; WGS 84, uncertainty: 100 m., Rosaceae Source: Digital map Mixed conifer and deciduous forest with brushy understory. Rubus parviflorus Nutt. Brambles with arching stems, the stems green with strong glaucous cast, thorny. Phenology: Buds. Origin: Native. U.S.A., WASHINGTON, SKAGIT COUNTY: Padilla Bay National Estuarine Research Reserve. Saddlebag Ben Legler 3146 2 Feb 2006 Island. Elev. 17 ft. Herbarium: WTU 48° 32.117' N, 122° 33.333' W; WGS 84, Source: GPS High island with rocky shoreline cliffs; Douglas fir/Madrone forests with Snowberry, Mahonia; forest and understory. Occasional in openings in forest and along shore. Phenology: Flowers. Origin: Native. Ben Legler 3358 23 May 2006 WTU-371865 Rosaceae Rosaceae Rubus bifrons Vest Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, SKAGIT COUNTY: At the Legler Residence (4115 234th St. NE) ca. 1.5 air miles Padilla Bay National Estuarine Research Reserve. Dot Island. northwest of Arlington. Elev. 69 ft. Elev. 290 ft. 48° 32.0412' N, 122° 33.2112' W; Source: Digital map 48° 12' 28" N, 122° 10' 25" W; WGS 84, uncertainty: 100 m., Sloping island with high end at north end. Rich slopes of Festuca Source: Digital map rubra, Plectritis, Cerastium, Camassia, and Zygadenus. Juniperus Mixed conifer and deciduous forest with open brushy area by road are common, very shaped and stunted by salt spray, little deer and adjacent small seasonal creek. Thicket-forming bramble, stems browse. No geese nesting; otter use on southwest side. Occasional arching, green to maroon. Phenology: Buds. Origin: Introduced. trailing across grassy patches; flowers white. Phenology: Flowers. Origin: Native. Ben Legler 3188 15 Feb 2006 Ben Legler 3755 1 May 2007 Herbarium: WTU WTU-377244 Rosaceae Rosaceae Rubus bifrons Vest Rubus pedatus Sm. U.S.A., WASHINGTON, SKAGIT COUNTY: Northwest shore of Grandy Creek, 3 air kilometers northeast of U.S.A., WASHINGTON, WHATCOM COUNTY: Birdsview Siding. North Cascades National Park. Chilliwack River trail, approximately Elev. 557 ft. 0.5 kilometers from junction with Copper Ridge Trail. 48° 33.5' N, 121° 50.5' W; T35N R7E S1 Elev. 4363 ft. Gravel streambank, partial shade, with Tanacetum, Thuja, Phalaris. 48° 53.329' N, 121° 31.236' W; NAD 83, uncertainty: 21 m. Black fruit, common adventive shrub, 1 meter tall, arching. Old growth forest dominated by Abies lasiocarpa with active stream. Phenology: Fruits. Origin: Introduced. Common in dry upland forest along trail. Phenology: Flowers & Fruits. Origin: Native. Peter F. Zika 22165 11 Aug 2005 Steve Walker 07-04 5 Aug 2007 WTU-363375 with Steve Hahn, Megan A. Jensen, Dale Blum, Lily Hickenbottom, David Giblin Rosaceae WTU-367596 Rubus idaeus L. ssp. strigosus (Michx.) Focke Rosaceae U.S.A., WASHINGTON, KING COUNTY: Montlake, Seattle; Near the intersection of East Louisa Street and Rubus parviflorus Nutt. 22nd Avenue East. Elev. 82 ft. U.S.A., WASHINGTON, KITTITAS COUNTY: 47° 39' N, 122° 18' W; T25N R4E S21 Wenatchee National Forest. Forest Service Shaded disturbed and neglected ground; with Prunella, Agrostis Road Number 9716. North-facing scree slope capillaris, Ilex aquifolium. Occasional self-sown seedlings; to 2 along road. meters tall; cultivated plants 25 distant are glandular. Phenology: Elev. 4969 ft. Vegetative. Origin: Introduced. 47° 18' 17" N, 120° 35' 50.5" W; WGS 84, uncertainty: 9 m. Peter F. Zika 20419 18 Oct 2004 Dry slope with large rocks; appears to be seasonally moist. Corollas white. Scrambling WTU-362308 shrub. Phenology: Flowers. Origin: Native. Ryan Miller 06-05 Rosaceae with David Giblin Rubus arcticus L. 8 Jul 2006 U.S.A., ALASKA, VALDEZ CORDOVA COUNTY: WTU-373675, WTU-V-045435 Central west shore of Kayak Island, near Cape St. Peter, Gulf of Alaska. Elev. 16 ft. 59° 56.2' N, 144° 25.3' W Rosaceae Margin of small fresh-water marsh, with Platanthera dilatata, Poa eminens, Ranunculus occidentalis, Equisetum arvense, E. Rubus parviflorus Nutt. Variegatum, Achillea millefolium, Carex lyngbyei, C. saxatilis. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, KITTITAS COUNTY: Wenatchee National Forest. Vicinity of Swauk Creek Camground. Peter F. Zika 17000 5 Jul 2002 Elev. 3257 ft. 47° 19.708' N, 120° 39.196' W WTU-365245 White flowers; shrub to 0.5 meters. In meadow near creek. Phenology: Flowers. Origin: Native. Sheila Olmstead 2006-06 8 Jul 2006 with Richard Olmstead, Richard Robohm WTU-370999 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, SKAGIT COUNTY: Esmeralda Basin. Padilla Bay National Estuarine Research Reserve. Hat Island. Elev. 4571 ft. Elev. 65 ft. 47° 26.475' N, 120° 56.387' W; WGS 84 48° 31.222' N, 122° 32.814' W; WGS 84 Wet meadow. 10 degree south facing slope. Saturated black soil. Douglas fir forested high island with steep grassy balds (Elymus Associated; Pinus monticola, Rhododendron albiflorum, Lonicera glaucus, Festuca rubra, Koeleria macrantha) on south side with involucrata, Alnus sinuata. Growing in mucky soil in forest mostly native bunch grasses and abundant Zigadenus venenosus. understory. Corolla white. Phenology: Flowers. Origin: Native. South grassy balds. Frequent in Pseudotsuga menziesii forest understory. Phenology: Flowers. Origin: Native. Barbara Williams 06-07 9 Jul 2006 with David Giblin Karen Reagan 07-35 1 May 2007 WTU-371871 WTU-377937

Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, YAKIMA COUNTY: Reef Island. Wenatchee National Forest, base of Devil's Table. Forest Service 48° 36.179' N, 123° 1.021' W Road 1503; 0.1 kilometers from junction with Forest Road 1501; Large island dominated by Pseudotsuga menziesii; open understory Little Rattlesnake Creek Canyon. with Elymus and Holodiscus. Several plants with Cirsium arvense. Elev. 3115 ft. Phenology: Vegetative. Origin: Native. 46° 46.215' N, 121° 1.005' W; WGS 84, Source: GPS Seasonal pond surrounded by Ponderosa pine upland and Peter Dunwiddie 07-100 19 Sep 2007 dominated by Carex aquatilis. Uncommon in forest understory. with Phil Green, Megan Jensen, David Giblin, Ted Thomas Phenology: Fruits. Origin: Native. WTU-375671 Megan A. Jensen 193 5 Jul 2007 with David Giblin, Don Knoke Rosaceae WTU-369867 Rubus laciniatus Willd. Rosaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands National Wildlife Refuge. Colville Island. Rubus armeniacus Focke × Rubus laciniatus Willd. ? 48.415278°, -122.82397°; WGS 84, uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., WASHINGTON, SAN JUAN COUNTY: Several extensive, dense patches. Phenology: Fruits. Origin: Puget Sound, San Juan Island. About 0.7 km south of Rosler Road Introduced. on Cattle Point Road. Elev. 164 ft. Ted Thomas 07-46 21 Sep 2007 48° 28.6' N, 123° 1.9' W with Peter Dunwiddie, Megan Jensen, Phil Green, David Giblin, Dry roadside thickets bordering cattle pasture, with: both putative Barbara French parents, Crataegus monogyna, C. suksdorfii, C. douglasii, Malus fusca. Possible hybrid, broad leaflets suggest influence of R. WTU-375538 armeniacus. Phenology: Fruits. Origin: Introduced. Peter F. Zika 23413 12 Aug 2007 Rosaceae WTU-367108 Rubus parviflorus Nutt. U.S.A., WASHINGTON, SKAGIT COUNTY: Padilla Bay National Estuarine Research Reserve. Hat Island. Rosaceae 48° 31.326' N, 122° 32.728' W; WGS 84 Douglas fir forested high island with steep grassy balds (Elymus Rubus idaeus L. ssp. strigosus (Michx.) Focke glaucus, Festuca rubra, and Koeleria macrantha) on south side, with mostly native bunch grasses and abundant Zigadenus venenosus. U.S.A., WASHINGTON, KING COUNTY: Understory of forest, interior of island to 325 feet. A couple of plants Montlake, Seattle. Near intersection of east Louisa Street and 22nd on east side of island. Phenology: Flowers. Origin: Native. Avenue east. Elev. 82 ft. David Giblin 862 30 Apr 2007 47° 39' N, 122° 18' W; T25N R4E S21 3 wild plants in shaded disturbed and neglected ground; (Cultivated WTU-377708 plants 25 meters distant); with Prunella, Agrostis capillaris, Ilex aquifolium. Occasional self-sown adventive arching glandular shrubs, fruit red, leaves white below. Phenology: Fruits. Origin: Introduced. Peter F. Zika 22154 19 Jul 2005 WTU-364964 Rosaceae Rosaceae Rubus bifrons Vest Rubus laciniatus Willd. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, COWLITZ COUNTY: North Finger Island. East bank of Carrolls Channel. Opposite south end of Cottonwood Elev. 0-65 ft. Island, Columbia River. 48° 45' 24.18" N, 122° 53' 27.64" W; WGS 84 Elev. 32 ft. Primarily Pseudotsuga/Arbutus forest with 46° 3.7' N, 122° 52' W; T7N R2W S30 exposed areas along the perimeter, especially Steep rocky bank. Flowers pinkish; 1 meter adventive evergreen at east end of island; sandstone substrate. shrub, arching; occasional. Phenology: Flowers & Fruits. Origin: Tissue removed by Gwen Ellis at University of Introduced. Washington for DNA analysis. February 4, 2020. Phenology: Fruits. Origin: Introduced. Peter F. Zika 19469 4 May 2004 David Giblin 1376 WTU-368550 27 Sep 2007 Rosaceae WTU-381067, WTU-V-060734 Rubus laciniatus Willd. U.S.A., WASHINGTON, SAN JUAN COUNTY: Rosaceae Puget Sound, San Juan Island. About 0.7 km south of Rosler Road on Cattle Point Road. Rubus laciniatus Willd. Elev. 164 ft. 48° 28.6' N, 123° 1.9' W U.S.A., WASHINGTON, SAN JUAN COUNTY: Dry roadside thickets bordering cattle pasture, with: Rubus North Finger Island. armeniacus and possible hybrids, Crataegus monogyna, Crataegus Elev. 0-65 ft. suksdorfii, Crataegus douglasii, Malus fusca. 1.5 meters tall arching 48° 45' 24.18" N, 122° 53' 27.64" W; WGS 84 adventive evergreen shrub. Phenology: Fruits. Origin: Introduced. Primarily Pseudotsuga/Arbutus forest with exposed areas along the perimeter, especially at east end of island; sandstone substrate. Peter F. Zika 23412 12 Aug 2007 Phenology: Fruits. Origin: Introduced. WTU-368592 David Giblin 1406 27 Sep 2007 WTU-381587 Rosaceae Rubus bifrons Vest Rosaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: Rubus pedatus Sm. Puget Sound, San Juan Island. About 0.7 km south of Rosler Road on Cattle Point Road. U.S.A., WASHINGTON, KITTITAS COUNTY: Elev. 164 ft. Wenatchee National Forest. Cottonwood Lake. 48° 28.6' N, 123° 1.9' W Elev. 3910 ft. Dry roadside thickets bordering cattle pasture, 47.34854° N, 121.42999° W; WGS 84, Source: GPS with: Rubus laciniatus and possible hybrids, Two hectare lake surrounded by old growth or mature second Crataegus monogyna, Crataegus suksdorfii, growth forest. South end of lake bordered by meadow of Carex sp. Crataegus douglasii, Malus fusca. 1.5 meter tall Uncommon in forest understory on south side of lake. Phenology: arching adventive evergreen shrub. Phenology: Fruits. Origin: Native. Fruits. Origin: Introduced. David Giblin 1199 13 Sep 2007 Peter F. Zika 23411 with Don Knoke, Barbara Williams, Megan A. Jensen 12 Aug 2007 WTU-374943 WTU-368593, WTU-V-060696 Rosaceae Rubus vestitus Weihe & Nees Rosaceae U.S.A., WASHINGTON, KLICKITAT COUNTY: Rubus bifrons Vest North side of Route 14, Bingen Gap, north shore of Columbia River. Elev. 82 ft. U.S.A., WASHINGTON, SNOHOMISH 45° 42' N, 121° 25.6' W; T3N R11E S34 COUNTY: Near pool at base of seeping cliff, roadcut, full sun, with Lactuca Kents Prairie near Arlington. serriola, Scrophularia. Black fruit, common evergreen shrub with 48.19889°, -122.12389°; WGS 84, scattered stipitate glands; common. Phenology: Remnants. Origin: uncertainty: 5000 m., Source: GeoLocate, Introduced. Georef'd by WTU Staff Roadside. Sprawly shrub 15 decimeters tall. Peter F. Zika 22615 13 Oct 2005 [Located in teaching collection]. Phenology: Flowers. Origin: Introduced. WTU-368317 M. Arnot 54 16 Jun 1987 WTU-314181, WTU-V-005623 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus pedatus Sm. U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., WASHINGTON, SNOHOMISH Along access road to Swinomish Channel. COUNTY: T34N R2E S11; NAD 27, uncertainty: 805 m., Mountain Loop Hiway [Highway] east of Red Source: TRS2LL, Georef'd by Ben Legler Bridge. Gravelly waste area. With other introduced T30N R9E S23; NAD 27, uncertainty: 805 m., weedy species. Large evergreen vinelike shrub. Source: TRS2LL, Georef'd by Ben Legler Stems arching, flowers pinkish. [Located in Roadside. Mossy bank with ferns and teaching collection]. Phenology: Flowers. Origin: Maianthemum dilatatum. Oplopanax horridum Introduced. and Polystichum munitum nearby. [Located in teaching collection]. Phenology: Flowers. Origin: M. Arnot 599 Native. 7 Jun 1993 M. Arnot 588 WTU-327222, WTU-V-005622 26 May 1993 WTU-327997, WTU-V-006611 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus spectabilis Pursh U.S.A., WASHINGTON, CHELAN COUNTY: Along trail at Basalt Pass. U.S.A., WASHINGTON, SNOHOMISH Elev. 4920 ft. COUNTY: T29N R17E S28; WGS84, uncertainty: 800 50 yards east of Stillaguamish River at m., Source: GeoLocate, Georef'd by WTU Arlington. Staff 48.233172°, -122.371907°; WGS 84, Conifer woods with understory of Vaccinium. uncertainty: 1000 m., Source: GeoLocate, [Located in teaching collection]. Phenology: Georef'd by WTU Staff Flowers. Origin: Native. Flood plain. With Acer macrophyllum, Populus trichocarpa, Salix spp. Prickly stemmed shrub 2 M. Arnot 155 meters tall. Flowers very deep pinkish purple. with E. Burnett [Located in teaching collection]. Phenology: Flowers. Origin: Native. 9 Jul 1989 M. Arnot 4 WTU-317074, WTU-V-006613 23 Mar 1987 Rosaceae WTU-314125, WTU-V-005606 Rubus parviflorus Nutt. Rosaceae U.S.A., WASHINGTON, SNOHOMISH Rubus ursinus Cham. & Schltdl. COUNTY: Stillaguamish River. 1.5 miles west of Arlington. U.S.A., OREGON, BENTON COUNTY: State Road 530. 2 miles east of State Highway 99W at the 48.201684°, -122.160147°; WGS 84, junction of Camp Adair Road and Independence uncertainty: 1000 m., Source: GeoLocate, Highway. Georef'd by WTU Staff Elev. 216 ft. Banks of river. Abundant. Shrub 1 meter tall. T10S R4W S16; NAD 27, uncertainty: 805 m., [Located in teaching collection]. Phenology: Source: TRS2LL, Georef'd by Ben Legler Flowers. Origin: Native. Associated genera: Cirsium, Hypericum, Festuca, Rhus, Daucus, Taraxacum, Lupinus. M. Arnot 103 Flowers white; prickly vine trailing on the ground. [Located in teaching collection]. 9 May 1988 Phenology: Flowers. Origin: Native. WTU-314672, WTU-V-006612 Richard R. Halse 2812 2 May 1984 WTU-305366, WTU-V-005605 Rosaceae Rosaceae Rubus bifrons Vest Rubus vestitus Weihe & Nees U.S.A., WASHINGTON, WHITMAN COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Near railroad and Wawawai River Road, west Northwest bank of Waatch River, southwest of town of Neah Bay, side of Snake River, 2 air kilometers northwest north of Hobuck Lake, north shore of Olympic Peninsula. of mouth of Steptoe Canyon. Elev. 6 ft. Elev. 738 ft. 48° 20.9' N, 124° 39.6' W; T33N R15W S21 46° 28.3' N, 117° 13.9' W; T12N R44E S36 Thickets. Pink flowers, evergreen, 1 meter tall, adventive. Thickets, with Rosa eglanteria, Carex Phenology: Flowers. Origin: Introduced. vulpinoidea, shore of backwater. Flowers pink; arching branches; adventive shrub. Phenology: P. F. Zika 22462 3 Oct 2005 Flowers. Origin: Introduced. with F. Weinmann Peter F. Zika 19758 WTU-368957 with R. Old 30 May 2004 Rosaceae WTU-368915, WTU-V-060719 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SAN JUAN COUNTY: Little Sucia Island, off the southwest corner of Sucia Island. Rosaceae 48° 45.283' N, 122° 55.198' W; Source: GPS Includes forested uplands (circa 80 feet high) with shrub understory, Rubus bifrons Vest steep sandstone cliffs, eroding dirt bluffs. Linked by a low, shrub-covered (Rosa nutkana) shell-midden berm. Common on U.S.A., WASHINGTON, CHELAN COUNTY: eroding south-facing bluffs. Phenology: Fertile. Origin: Native. West side of Route 97A, 6 air kilometers north-northeast of Rocky Reach Dam on Columbia River. Erica Nees 08-20 23 Apr 2008 Elev. 721 ft. 47° 34.7' N, 120° 15.3' W; T24N R21E S18 WTU-393525 Margin of pool, full sun. Common adventive arching shrub 3 meters tall, fruit black. Phenology: Fruits. Origin: Introduced. Rosaceae P. F. Zika 19914 10 Aug 2004 with F. Weinmann Rubus spectabilis Pursh WTU-369073 U.S.A., WASHINGTON, SAN JUAN COUNTY: Matia Island. 48.74791° N, 122.84196° W; Source: GPS Rosaceae Vicinity of campsites at west end of island. Large mixed conifer forest, with several open, disturbed campsites. Frequent around Rubus ursinus Cham. & Schltdl. campsite. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, PACIFIC COUNTY: Peter Dunwiddie 08-15 24 Apr 2008 Both sides of Route 101, 5 air kilometers east northeast of Stoney Point, Willapa Bay, Stuart Slough. WTU-377296 Elev. 16 ft. 46° 41.2' N, 123° 52.3' W; T14N R10W S24 Roadside thickets, full sun, with Rubus armeniacus, Rubus Rosaceae laciniatus, Rubus macrophyllus. Common shrub, canes glaucous purple, arching or trailing, to 1 meter high. Phenology: Vegetative. Rubus ursinus Cham. & Schltdl. Origin: Native. U.S.A., WASHINGTON, SAN JUAN COUNTY: Peter F. Zika 20354 29 Sep 2004 Matia Island. 48.74791° N, 122.84196° W; Source: GPS WTU-368976 Along the trail across to the east cove, and from bluffs, headlands, and shoreline habitats near the cove. Abundant understory trailer, Rosaceae forming dense patches extensively under Pseudotsuga and Thuja forest. Phenology: Fertile. Origin: Native. Rubus idaeus L. Peter Dunwiddie 08-54 24 Apr 2008 U.S.A., IDAHO, BONNER COUNTY: North end of town of Sandpoint. WTU-376200 48.27667°, -116.55222°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Along railroad track. Plants about 3 feet tall, the stamens very prominent even from a distance. Phenology: Flowers. Origin: Native. Rich Old s.n. 28 May 2005 WTU-369397 Rosaceae Rosaceae Rubus bifrons Vest Rubus laciniatus Willd. U.S.A., WASHINGTON, WALLA WALLA U.S.A., WASHINGTON, SAN JUAN COUNTY: COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. South side of Route 124, edge of McNary Turn Island. Large island 0.25 miles off the east shore of San Juan National Wildlife Refuge, west end of Burbank Island. Heights. 48° 31.98' N, 122° 58.22' W; WGS 84, uncertainty: 10 m., Source: Elev. 360 ft. GPS 46° 12.7' N, 119° 0.4' W; T8N R30E S1 On dry rocks. A few plants at southeast end of island, near Full sun, dry gravelly slope below road, with campsites. Phenology: Vegetative. Origin: Introduced. Melilotus albus, Bromus tectorum. Common adventive to 3 meters tall, arching, black fruit Eliza Habegger EH1124 29 May 2008 13-22 x 17-21 millimeters wide, shiny, juicy; field edges. Phenology: Fruits. Origin: Introduced. WTU-394387 Peter F. Zika 22330 Rosaceae 12 Sep 2005 Rubus parviflorus Nutt. WTU-370146, WTU-V-060699 U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. Turn Island. Large island 0.25 miles off the east shore of San Juan Rosaceae Island. 48° 31.98' N, 122° 58.22' W; WGS 84, uncertainty: 10 m., Source: Rubus parviflorus Nutt. GPS On dry rocks. A few plants at southeast end of island. Phenology: U.S.A., WASHINGTON, SAN JUAN COUNTY: Flowers. Origin: Native. Ewing Island. Long, narrow island off NE end of Sucia Island. Steep sandstone ridges run length of island. Eliza Habegger EH1141 29 May 2008 48.76455° N, 122.88093° W; WGS 84, Source: Digital map Mostly forested with PSME, ARME, and JUMA. Grassy balds along WTU-395000 south shore and at east end of island. Several plants with Rosa nutkana. Phenology: Flowers. Origin: Native. Rosaceae Janneke Hille-Ris Lambers 08-01 28 May 2008 Rubus ursinus Cham. & Schltdl. WTU-393270 U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. Small islet just off the southwest shore of Turn Island. Rosaceae 48° 31.87' N, 122° 58.22' W; NAD 83, Source: Digital map Several Juniperus, one Pseudotsuga menziesii; open, weedy Rubus ursinus Cham. & Schltdl. grassland understory. Several trailing plants. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, SAN JUAN COUNTY: Ewing Island. Long, narrow island off NE end of Sucia Island. Steep Thor Hanson TH89 29 May 2008 sandstone ridges run length of island. 48.76455° N, 122.88093° W; WGS 84, Source: Digital map WTU-394638 Mostly forested with PSME, ARME, and JUMA. Grassy balds along south shore and at east end of island. Frequent and widespread across island. Phenology: Flowers. Origin: Native. Rosaceae Janneke Hille-Ris Lambers JHRL 08-36 28 May 2008 Rubus bifrons Vest WTU-395989 CANADA, BRITISH COLUMBIA: Martin's Island. 50° 39' N, 126° 14' W Rosaceae Near shore on east side. Phenology: Fruits. Origin: Introduced. Rubus ursinus Cham. & Schltdl. Brian D. Compton 96 2 Aug 1987 U.S.A., WASHINGTON, SAN JUAN COUNTY: WTU-371244 San Juan Islands National Wildlife Refuge and Wilderness Area. Turn Island. Large island 0.25 miles off the east shore of San Juan Island. Rosaceae 48° 31.98' N, 122° 58.22' W; WGS 84, uncertainty: 10 m., Source: GPS Rubus spectabilis Pursh On dry rocks. Common and widespread under trees and shrubs. Phenology: Flowers. Origin: Native. CANADA, BRITISH COLUMBIA: Area in back of Bella Bella Community School, Campbell Island. Eliza Habegger EH1123 29 May 2008 52° 9' N, 128° 7' W Gravelly area with small marshy area. Phenology: Flowers. Origin: WTU-394386 Native. Brian D. Compton 59 30 Jun 1987 WTU-371298 Rosaceae Rosaceae Rubus parviflorus Nutt. ssp. parvifolorus Rubus leucodermis Douglas ex Torr. & A. Gray CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, SKAGIT COUNTY: Area in back of Bella Bella Community School, Campbell Island. Huckleberry Island State Park. 52° 9' N, 128° 7' W Elev. 0-28 m. Gravelly area with small marshy area. Phenology: Flowers. Origin: 48.53666° N, 122.56889° W; WGS 84, Source: Nat. Geog. Native. TOPO!, Georef'd by WTU Staff Pseudotsuga menziesii forest dominates interior island; grassy Brian D. Compton 61 30 Jun 1987 headlands ring almost the entire island perimeter. Uncommon in forest understory. Phenology: Flowers. Origin: Native. WTU-371338 David Giblin 1723 16 Jun 2008 Rosaceae WTU-380163 Rubus bifrons Vest Rosaceae CANADA, BRITISH COLUMBIA: Oweekeno Village. Rubus leucodermis Douglas ex Torr. & A. Gray Phenology: Buds. Origin: Introduced. U.S.A., WASHINGTON, YAKIMA COUNTY: Brian D. Compton 137 11 May 1989 Wenatchee National Forest. Junction of Forest Road 1400 and Forest Road 216. Oak Creek Canyon. WTU-371344 Elev. 2791 ft. 46.73701° N, 120.91067° W; WGS 84, uncertainty: 11 m. Pinus ponderosa and Pseudotsuga menziesii overstory; dry Rosaceae understory dominated by Pseudoroegneria spicata, Carex geyeri, and Lupinus sp. Growing on steep south-facing embankment; Rubus pedatus Sm. uncommon. Phenology: Flowers. Origin: Native. CANADA, BRITISH COLUMBIA: Don Knoke 1617 9 Jun 2008 Oweekeno Village. with David Giblin Phenology: Remnants. Origin: Native. WTU-379917 Brian D. Compton 139 11 May 1989 WTU-371345 Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rosaceae U.S.A., OREGON, DOUGLAS COUNTY: Rubus parviflorus Nutt. Beatty Creek RNA, Roseburg BLM, 10 air miles SW of town of Myrtle Creek. U.S.A., WASHINGTON, SAN JUAN COUNTY: Elev. 950 ft. North Finger Island. T30S R6W S31 NW1/4; NAD 27, uncertainty: 805 m., Source: Elev. 0-65 ft. TRS2LL, Georef'd by Ben Legler 48° 45' 24.18" N, 122° 53' 27.64" W Mesic slope under mixed conifers. White flowers, glaucous purple 0.8 mile-long island dominated by Pseudotsuga menziesii overstory. stem. Phenology: Fertile. Origin: Native. Coastal bluff habitat on south and extreme eastern edge of island. Occasional in forest understory. Phenology: Flowers. Origin: Native. P. F. Zika 10328 19 May 1987 David Giblin 1466 12 May 2008 WTU-372274 WTU-385810 Rosaceae Rosaceae Rubus bifrons Vest Rubus bifrons Vest U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. U.S.A., WASHINGTON, GRANT COUNTY: Flower Island, east of Lopez Island. Babcock Bench. Along lightly forested edge of Elev. 49 ft. Columbia River. 48.54536° N, 122.8539° W Elev. 540 ft. Several hectare, 50' high bedrock island. A mix of grassy ledges, 47° 9.9864' N, 120° 0.368' W shrub thickets, a few trees (Arbutus, Pseudotsuga, low Quercus). Riparian strip dominated by Ulmus pumila and Common and widespread across most of island. Phenology: Fruits. Morus alba. Phenology: Remnants. Origin: Origin: Introduced. Introduced. Phil Green 08-119 25 Sep 2008 Joe Arnett 08-49 WTU-396975 29 Apr 2008 WTU-380774, WTU-V-060697 Rosaceae Rosaceae Rubus hispidus L. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, PACIFIC COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Near Jim Street, circa 2 air kilometers northeast of Seaview. San Juan Islands National Wildlife Refuge and Wilderness Area. Elev. 16 ft. East Sucia #6. T10N R11W S22 NW 1/4; NAD 27, uncertainty: 805 m., Source: 48.76422° N, 122.88631° W TRS2LL, Georef'd by Ben Legler A high, cliffy sandstone island, cliffs all around, Juniperus over top of In cultivated cranberry fields, with Vaccinium macrocarpon. Trailing island. 0.55 acres. Island is predominantly old junipers, with dry weed, shiny leaves. Phenology: Flowers. Origin: Introduced. south east facing slope of Vulpia and Bromus, rich north-facing slope with Plectritis, Sedum spath, Collinsea. A couple of plants at edge of P. F. Zika 14184 31 Aug 1999 island, west side. Phenology: Flowers & Fruits. Origin: Native. WTU-340863 Cheryl Lowe 09-20 29 May 2009 WTU-394460 Rosaceae Rubus hispidus L. Rosaceae U.S.A., WASHINGTON, PACIFIC COUNTY: Rubus ursinus Cham. & Schltdl. Near Jim Street, circa 2 air kilometers northeast of Seaview. Elev. 16 ft. U.S.A., WASHINGTON, SAN JUAN COUNTY: T10N R11W S22 NW 1/4; NAD 27, uncertainty: 805 m., Source: San Juan Islands National Wildlife Refuge and Wilderness Area. TRS2LL, Georef'd by Ben Legler East Sucia #7. Near cultivated cranberry fields, sandy mowed slope. Trailing weed, Elev. 50 ft. shiny leaves. Phenology: Flowers. Origin: Introduced. 48.76398° N, 122.88579° W The largest island in the chain. Forested with Arbutus, Pseudotsuga, P. F. Zika 14184A 31 Aug 1999 with shrub understory. Densely forested with Juniperus. Height of 50 feet or so. Several trailing over shrubs on south side of island. WTU-342040 Flowers white. Phenology: Flowers. Origin: Native. Cheryl Lowe 09-50 29 May 2009 Rosaceae WTU-395097 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SAN JUAN COUNTY: Rosaceae San Juan Islands National Wildlife Refuge and Wilderness Area. Flower Island, east of Lopez Island. Rubus bifrons Vest Elev. 49 ft. 48.54536° N, 122.8539° W; WGS 84 U.S.A., WASHINGTON, SAN JUAN COUNTY: Several hectare, 50' high bedrock island. A mix of grassy ledges, San Juan Islands. Gossip Island. shrub thickets, a few trees (Arbutus, Pseudotsuga, low Quercus). Elev. 25 ft. Common throughout islands on rocky outcroppings. Phenology: 48° 39.789' N, 123° 10.374' W Flowers. Origin: Native. Low conglomerate and sandstone bedrock island with gently sloping bedrock on all sides. Scattered madrone, Douglas fir, and junipers Tad Dillhoff 09-22 22 Apr 2009 across island with lush camas on swales between outcrop. One shrub on steep south-facing bank. Phenology: Remnants. Origin: WTU-395185 Introduced. Ben Legler 11319 28 May 2009 Rosaceae Herbarium: WTU Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SAN JUAN COUNTY: Rosaceae San Juan Islands National Wildlife Refuge and Wilderness Area. Willow Island. A large, 100' high rocky island 1/3 mile west of Blakely Rubus parviflorus Nutt. Island. Elev. 100 ft. U.S.A., WASHINGTON, SAN JUAN COUNTY: 48.5402° N, 122.82242° W; WGS 84, Source: GPS Little Sucia Island, off the southwest corner of Sucia Island. Eastern Partially treed with Pseudotsuga and Arbutus. Open understory portion. dominated by Elymus glaucus and Symphoricarpos albus. East side Elev. 80 ft. has extensive cliffs and ledges and is particularly rich with ferns. 48° 45.283' N, 122° 55.198' W Considerable deer browse. Rock appears to be serpentine. Common Includes forested uplands (circa 80 feet high) with shrub understory, on island in rocky outcroppings. Phenology: Flowers. Origin: Native. steep sandstone cliffs, eroding dirt bluffs. Linked by a low, shrub-covered (Rosa nutkana) shell-midden berm. Psuedotsuga Rick Dillhoff 09-20 22 Apr 2009 menziesii and arbutus forest with dense shrub understory. Isthmus connects two higher upland areas. Several scattered plants in moist WTU-393207 ledges and gullys along north side of island. Phenology: Flowers. Origin: Native. Ben Legler 11331 29 May 2009 WTU-393616 Rosaceae Rosaceae Rubus bifrons Vest Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Gossip Island. San Juan Islands. Little Sucia Island, off the Elev. 25 ft. southwest corner of Sucia Island. 48° 39.789' N, 123° 10.374' W Elev. 80 ft. Low conglomerate and sandstone bedrock island with gently sloping 48° 45.283' N, 122° 55.198' W bedrock on all sides. Bedrock is conglomerate and sandstone. Includes forested uplands (circa 80 feet high) Scattered madrone, Douglas fir, and junipers across island with lush with shrub understory, steep sandstone cliffs, camas on swales between outcrop. One shrub on steep, eroding dirt bluffs. Linked by a low, south-facing bank. Phenology: Remnants. Origin: Introduced. shrub-covered (Rosa nutkana) shell-midden berm. Uncommon in understory at forest edge, Ben Legler 11319 28 May 2009 east end of island. Phenology: Vegetative. Origin: Native. WTU-393644 Amy Dearborn 09-51 Rosaceae 24 Sep 2009 Rubus ursinus Cham. & Schltdl. WTU-399069, WTU-V-005501 U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Little Sucia Island, off the southwest corner of Rosaceae Sucia Island. Elev. 80 ft. Rubus ursinus Cham. & Schltdl. 48° 45.283' N, 122° 55.198' W Includes forested uplands (circa 80 feet high) with shrub understory, U.S.A., WASHINGTON, SAN JUAN COUNTY: steep sandstone cliffs, eroding dirt bluffs. Linked by a low, Ewing Island. Long, narrow island off northeast shrub-covered (Rosa nutkana) shell-midden berm. Psuedotsuga end of Sucia Island. Steep sandstone ridges run menziesii and arbutus forest with dense shrub understory. Isthmus length of island. connects two higher upland areas. Frequent trailing over shrubs and Elev. 39 ft. ground on steep south-facing bank. Phenology: Fruits. Origin: 48.76455° N, 122.88093° W; WGS 84, Native. Source: Digital map Mostly forested with PSME, ARME, and JUMA. David Giblin 2857 28 May 2009 Grassy balds along south shore and at east end of island. Scattered in shrub patches. WTU-393564 Phenology: Vegetative. Origin: Native. Rosaceae Peter Dunwiddie 09-102 Rubus laciniatus Willd. 24 Sep 2009 U.S.A., WASHINGTON, SAN JUAN COUNTY: WTU-403756, WTU-V-012811 San Juan Islands National Wildlife Refuge and Wilderness Area. Flower Island, east of Lopez Island. Elev. 49 ft. 48.54536° N, 122.8539° W; WGS 84 Rosaceae Several hectare, 50' high bedrock island. A mix of grassy ledges, shrub thickets, a few trees (Arbutus, Pseudotsuga, low Quercus). Rubus lasiococcus A. Gray Several scattered patches, with Rubus armeniacus. Phenology: Vegetative. Origin: Introduced. U.S.A., WASHINGTON, KITTITAS COUNTY: Near Forest Road 3100 (Quartz Mountain Road) about 1/2 mile Phil Green 09-55 27 May 2009 above junction with Forest Road 230. Elev. 5910 ft. WTU-393445 47.07017° N, 121.06326° W; T18N R14E S3; WGS 84 Dry, gravelly soil in forest opening; 5º north-facing slope. Other plants: Abies lasicarpus, Pinus albicaulis, Ligusticum grayi, Erigeron Rosaceae peregrinus. Flowers white; common. Phenology: Flowers. Origin: Native. Rubus ursinus Cham. & Schltdl. Don Knoke 1918 29 Jul 2009 U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. West Matia #1. Large island WTU-379911 at north side of harbor, west end of Matia Island. 48.74946° N, 122.84378° W; WGS 84, Source: Digital map Cliffy sandstone island rising steeply out of cove at west end of Matia Island. Approximately 50 feet high, with Pseudotsuga menziesii and Arbutus menziesii forest across top. Common in forest understory, with Gaultheria shallon. Phenology: Vegetative. Origin: Native. Brianne Cohen 09-106 24 Sep 2009 WTU-398352, WTU-V-005549 Rosaceae Rosaceae Rubus bartonianus M. Peck Rubus vestitus Weihe & Nees U.S.A., IDAHO, ADAMS COUNTY: U.S.A., OREGON, LANE COUNTY: [locality and coordinates withheld; status in Idaho: S2 (Imperiled; South Eugene at intersection of Agate Street and Stony Ridge Road. IDNHP)] Elev. 700 ft. Elev. 1700 ft. UTM Zone 10, 494503E, 4874003N; NAD 27 Bunchgrass Zone; cliffside, in rocks and crevices. Associate: Edge of mature Douglas fir forest in part shade. With Pseudotsuga Philadelphus lewisii, Amelanchier alnifolia, Agropyron spicatum. menziesii, Galium aparine, Berberis aquifolium, Prunus avium. Slope: steep. Exposure: west. Soil: metamorphic sedimentary. Flowers pink to white. Phenology: Flowers. Origin: Introduced. Abundance (1-5): 2-3. Height: 3-7 feet. Mass collection from several different plants. A few in late flower. Several seedlings in disturbed R.E. Brainerd 853 20 Jun 2007 areas since Hell's Canyon Dam construction ceased--1968-1969. Duplicate sheet from University of Idaho Herbarium. Phenology: WTU-379745 Flowers & Fruits. Origin: Native. F. D. Johnson s.n. 24 Jun 1976 Rosaceae with J. R. Johnson Rubus bifrons Vest WTU-377009 U.S.A., WASHINGTON, WALLA WALLA COUNTY: Rosaceae Entrance to Hood Park campground and roadside edge of State Highway 124. Rubus flagellaris Willd. Elev. 357 ft. 46.21044° N, 119.01283° W; WGS 84, U.S.A., VERMONT, RUTLAND COUNTY: uncertainty: 5 m. Near Route 22A, Fair Haven. Mowed pasture and gravelly roadside edge. Railroad right-of-way, full sun, dry ground, with Lactuca canadensis, Locally common in roadside gravel forming Artemisia vulgaris, Panicum clandestinum. Prostrate. Phenology: large, dense stand. Phenology: Remnants. Fruits. Origin: Native. Origin: Introduced. Peter F. Zika 4411 22 Jul 1981 David Giblin 2574B with Don Knoke WTU-375193 16 Apr 2009 Rosaceae WTU-399964, WTU-V-006254 Rubus allegheniensis Porter Rosaceae CANADA, BRITISH COLUMBIA: Pitt Meadows. Rubus laciniatus Willd. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, WALLA WALLA V. C. Brink s.n. 5 Aug 1973 COUNTY: with S. McHale Ice Harbor Dam and Locks Complex, Snake River, Charbonneau Park, area around parking WTU-376111 lot. Elev. 442 ft. 46.24729° N, 118.87057° W; WGS 84, Rosaceae uncertainty: 5 m. Highly disturbed sand dune community with Rubus allegheniensis Porter Bromus tectorum and Artemisia tridentata. Forming dense, impenetrable stands along U.S.A., WASHINGTON, KING COUNTY: riverbank. Phenology: Remnants. Origin: Along trail at edge of Christmas Lake, south of North Bend. Introduced. Elev. 950 ft. UTM Zone 10, 593486E, 5254271N; WGS 84, uncertainty: 300 David Giblin 2585 m., Source: Digital Map, Georef'd by Ibro Polovina with Don Knoke Forming thicket on brushy trail side in sun by lake, with Rubus armeniacus (which has leaves white woolly beneath and less of a 16 Apr 2009 trip tip on the leaflets), and Rubus spectabilis. In zone of Alnus rubra between conifer forest and lake. Fruit black. Canes arching. WTU-400746, WTU-V-007264 Phenology: Fruits. Origin: Native. Barbara L. Wilson 15804 3 Aug 2009 with Nick Otting, Laura Potash Martin WTU-379069 Rosaceae Rosaceae Rubus bifrons Vest Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, BENTON COUNTY: U.S.A., MONTANA, GRANITE COUNTY: South side of Highway 240 along irrigation ditch Butte Creek, two miles up creek from Rock Creek. Trail ends at adjacent to Yakima River. 46.51519 N, 113.73225 W. Elev. 410 ft. Elev. 4453-4842 ft. 46.37389° N, 119.40719° W; WGS 84, 46.51036° N, 113.73225° W uncertainty: 5 m. Uneven area between creek and talus slopes. Soil moist, rocky. Highly degraded shrub-steppe habitat with Associated species: Picea engelmannii, Aster radulinus, sandy soil; Artemisia tridentata dominant with Philadelphus lewisii, Symphoricarpos albus, Physocarpus. White Poa bulbosa. Locally common along irrigation flowers; common. Phenology: Flowers. Origin: Native. ditch, forming impenetrable patches. Phenology: Remnants. Origin: Introduced. Hannah Marx 08-41 12 Jul 2008 Don Knoke 1772 WTU-395892 with David Giblin 17 Apr 2009 Rosaceae WTU-398370, WTU-V-005567 Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., MONTANA, GRANITE COUNTY: Lolo National Forest. Rock Creek Canyon. Turnout for Microburst Rosaceae interpretive sign. 46° 26' 39.03" N, 113° 45' 41.42" W; WGS 84, uncertainty: 10 m., Rubus parviflorus Nutt. Source: Digital map Mixture of mesic and dry meadows along Rock Creek, bordered by U.S.A., MONTANA, RAVALLI COUNTY: south-facing, dry rocky slope. Shrub to 2 meters tall near edge of Bitterroot National Forest. Skalkaho Falls. meadow. Phenology: Flowers. Origin: Native. Southeast slope of falls and along road. Elev. 6300 ft. Hanh-La Phan 08-44 12 Jul 2008 46.2874° N, 113.82658° W; WGS 84, uncertainty: 16 m., Source: GPS WTU-394726 Moist/wet/spray areas along falls and water course; edges of rock scree; rocky cliff along falls and opposite side of road; hot, dry road cut Rosaceae slope; downhill side road cut. Subalpine fir, Picea woods either side of water course. Border Rubus parviflorus Nutt. of rock scree adjacent to falls; white flowers; common. Phenology: Flowers. Origin: Native. U.S.A., MONTANA, GRANITE COUNTY: Lolo National Forest, Harry's Flat campground. Robert Goff 08-31 Elev. 4153 ft. 46.53459° N, 113.75323° W; WGS 84 11 Jul 2008 Mixed conifer forest (Pseudotsuga, Pinus, Picea) with opening, wet meadow, and riparian habitats. White flowers; shrub to 1 meter. WTU-394492, WTU-V-006787 Phenology: Flowers. Origin: Native. Richard G. Olmstead 2008-14 11 Jul 2008 Rosaceae WTU-396127 Rubus parviflorus Nutt. Rosaceae U.S.A., MONTANA, GRANITE COUNTY: Butte Creek, two miles up creek from Rock Creek. Trail ends at Rubus parviflorus Nutt. 46.51519 N, 113.73225 W. Elev. 4453-4842 ft. U.S.A., WASHINGTON, KITTITAS COUNTY: 46.51036° N, 113.73225° W Along Forest Road 33 about 1 mile west of end Uneven area between creek and talus slopes. Soil moist, rocky. of county road (Taneum Road). Associated species: Picea engelmannii, Aster radulinus, Elev. 2184 ft. Philadelphus lewisii, Symphoricarpos albus, Physocarpus. White; 47° 5.84' N, 120° 49' W; T19N R16E S34; abundant. Phenology: Flowers. Origin: Native. WGS 84 Dry, gravelly soil at roadside. Other plants: Rosa Hannah Marx 08-13 12 Jul 2008 woodsii, Populus trichocarpa, Clematis lingusticifolia, Purshia tridentata. Flowers white; WTU-395860 common. Phenology: Flowers. Origin: Native. Don Knoke 1844 19 Jun 2009 WTU-380808, WTU-V-045459 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus pedatus Sm. U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., IDAHO, KOOTENAI COUNTY: Along Forest Road 33 about 1 mile west of end of county road Coeur d'Alene Forest. Montford Creek Natural Area, Deception (Taneum Road). Creek Experimental Forest; 12 miles east-northeast of Coeur Elev. 2184 ft. d'Alene. 47° 5.84' N, 120° 49' W; T19N R16E S34; WGS 84 Elev. 3100 ft. Dry, gravelly soil at roadside. Other plants: Rosa woodsii, Populus T51N R1W S30; NAD 27, uncertainty: 805 m., Source: TRS2LL, trichocarpa, Clematis lingusticifolia, Purshia tridentata. Occasional. Georef'd by Ben Legler Phenology: Fruits. Origin: Native. Western hemlock forest. Stream valley bottom. Slope 10-30%. Northwest exposure. Perennial herb. Locally frequent. Fruit a red Don Knoke 1843 19 Jun 2009 aggregate. Rhizomatose. Phenology: Flowers & Fruits. Origin: Native. WTU-380811 Peter F. Stickney 1249 18 Jul 1965 Rosaceae WTU-380616 Rosaceae Rubus ursinus Cham. & Schltdl. Rubus parviflorus Nutt. U.S.A., IDAHO, BONNER COUNTY: Shepherd Lake. U.S.A., IDAHO, BONNER COUNTY: Elev. 2280 ft. Forests around Lee Lake, 1 mile east of Coolin along Lee Road T56N R2W S24; NAD 27, uncertainty: 805 m., Source: TRS2LL, (Forest Road 19). Georef'd by Ben Legler Elev. 2500 ft. Thuja plicata/Coptis occidentalis habitat type moderately dense T59N R4W S11 NE 1/4 of SW 1/4; NAD 27, uncertainty: 805 m., pole-size stand of Thuja; occasional with Vaccinium globulare. Source: TRS2LL, Georef'd by Ben Legler Phenology: Vegetative. Origin: Native. 10-15% slope, northern exposure, humus soil; moist shaded understory in Thuja plicata/Clintonia uniflora habitat type; common Fred Johnson 86106 6 Aug 1986 with Physocarpus malvaceus, Rosa woodsii, Cornus canadensis and Acer glabrum. White flowers; height up to 1 meter. Phenology: WTU-380620 Flowers. Origin: Native. Christine L. Johnson 1456 7 Jun 1986 Rosaceae WTU-380615 Rubus ursinus Cham. & Schltdl. U.S.A., IDAHO, IDAHO COUNTY: Rosaceae Clearwater National Forest. Middle Fork Clearwater River Valley, Highway US-12, 0.1 miles west of Three Devils; 15.8 miles east of Rubus pedatus Sm. Kooskia. Elev. 1300 ft. U.S.A., IDAHO, BONNER COUNTY: T33N R6E S1 SE; NAD 27, uncertainty: 805 m., Source: TRS2LL, About 5 miles north of Lionhead State Park (along north end of Georef'd by Ben Legler Priest Lake), Kaniksu National Forest. Overgrown roadway. Southeast exposure. Slope 10%. Western Elev. 2700 ft. redcedar-grand fir open forest. Appears associated with disturbance. T63N R4W S23; NAD 27, uncertainty: 805 m., Source: TRS2LL, Reclining to prostrate low woody plant. Frequent to locally common. Georef'd by Ben Legler Phenology: Flowers. Origin: Native. Gentle slope, south aspect, soil granitic p.m. very wet with rivulets; mature Tsuga heterophylla/Gymnocarpium dryopteris habitat type Peter F. Stickney 2638 29 May 1972 dominated by Thuja plicata and Tsuga heterophylla; occasional with dense herbaceous understory of Athyrium filix-femina and WTU-380617 Oplopanax horridum. Prostrage woody evergreen. Phenology: Vegetative. Origin: Native. Rosaceae Fred D. Johnson 87031 13 Aug 1987 Rubus ursinus Cham. & Schltdl. WTU-380648 U.S.A., IDAHO, BONNER COUNTY: Sandpoint Baldy Road (road to Bald Mountain), about 9 miles Rosaceae west-northwest of Sandpoint. Elev. 4000 ft. Rubus pedatus Sm. T57N R3W S4; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler U.S.A., IDAHO, BOUNDARY COUNTY: Tsuga heterophylla/Clintonia habitat type; 20% southwest slope; in Copper Lake. 15-year old clearcut with Rubus ideaus, Salix scouleriana and Acer T65N R3E S8; uncertainty: 587 m., Source: GeoLocate, Georef'd glabrum. Stems prostrate, trailing 6-8 feet long before rooting. by Melissa Bowers Phenology: Flowers. Origin: Native. In sphagnum around lake edge; surrounding forest of Abies lasiocarpa/Menziesia ferruginea habitat type; rare, procumbent F. Johnson 88049 8 Jul 1988 plants with Kalmia microphylla; no other woody plants present at the sphagnum fringe. Phenology: Vegetative. Origin: Native. WTU-380618 Fred Johnson 8673 3 Aug 1986 WTU-380649 Rosaceae Rosaceae Rubus pedatus Sm. Rubus nivalis Douglas ex Hook. U.S.A., IDAHO, BOUNDARY COUNTY: U.S.A., IDAHO, CLEARWATER COUNTY: Beaver Creek, 0.5 mile from main Smith Creek Road. St. Joe National Forest. 5 miles east of Elk Elev. 4300 ft. Road on old Breakfast Creek Road. T64N R3W S9; NAD 27, uncertainty: 805 m., Source: TRS2LL, Elev. 3425 ft. Georef'd by Ben Legler T40N R3E S7; NAD 27, uncertainty: 805 m., 0-10% slope, northern exposure; moist pockets within mature Tsuga Source: TRS2LL, Georef'd by Ben Legler heterophylla/Gymnocarpium dryopteris habitat type. Common Thuja/Pachistima habitat type. Pole stand. ground cover with Asarum caudatum, Vaccinum membranaceum Associates: Pachistima, Rubus parviflorus, and Coptis occidentalis. Phenology: Fruits. Origin: Native. Vaccinium membranaceum. Steep slope. Northeast exposure. Creeping shrub; rare. This Fred Johnson 8677 3 Aug 1986 plant is rare in Idaho. Survey 439A. Phenology: Vegetative. Origin: Native. WTU-380646 F. D. Johnson 6622 Rosaceae 28 Jun 1966 Rubus pedatus Sm. WTU-380642, WTU-V-18514 U.S.A., IDAHO, SHOSHONE COUNTY: 20 miles north of Prichard at Nowhere Flat, junction of Miners Creek with the Coeur d'Alene River, Coeur d'Alene National Forest. Rosaceae Elev. 2500 ft. T52N R3E S29 NW; NAD 27, uncertainty: 805 m., Source: Rubus parviflorus Nutt. TRS2LL, Georef'd by Ben Legler Dense mature Tsuga heterophylla/Athyrium filix-femina habitat type; U.S.A., IDAHO, BOUNDARY COUNTY: humusy soil, toe slope near river; scattered with Oplopanax 7.1 miles from junction of West Side Road and horridum, Clintonia uniflora, Trautvetteria caroliniensis, Coptis Boundary Creek Road, (10 airmiles southwest occidentalis and Dryopteris filix-mas. Phenology: Flowers. Origin: of Porthill) along Smith Creek, Kaniksu National Native. Forest. Elev. 3500 ft. F. Johnson 88004 10 Jun 1988 T64N R3W S1 SE of NE; WGS 84, uncertainty: 2000 m., Source: Digital Map, WTU-380647 Georef'd by Melissa Bowers 0% slope, soil decomposed granite, organic; Rosaceae openings within Tsuga heterophylla/Clintonia uniflora habitat type; occasional with Thuja Rubus pedatus Sm. plicata, Salix sitchensis, Linnaea borealis, Alnus sinuata, Menziesia ferruginea and Abies U.S.A., IDAHO, SHOSHONE COUNTY: lasiocarpa. Height to 1 meter. Phenology: Nowhere Flat along Coeur d'Alene River, 19 miles upstream from Flowers. Origin: Native. Shoshone Station. T52N R3E S19; NAD 27, uncertainty: 805 m., Source: TRS2LL, Christine L. Johnson 1693 10 Jul 1986 Georef'd by Ben Legler 0% slope, alluvial Belt Series; Tsuga heterophylla/Athyrium WTU-380637, WTU-V-045460 filix-femina and Streptopus amplexifolius habitat type mosaic, mature; occasional with Gymnocarpium dryopteris, Asarum caudatum and Coptis occidentalis. Trailing perennial. Phenology: Rosaceae Flowers & Fruits. Origin: Native. Rubus parviflorus Nutt. F. D. Johnson 8629 20 Jun 1986 with R. S. Barth U.S.A., IDAHO, CARIBOU COUNTY: Preuss Range, upper Deer Creek/Crow Creek WTU-380644 drainage. 20 miles east of Soda Springs. Elev. 7400 ft. Pole size lodgepole pine forest. Abies Rosaceae lasiocarpa/Vaccinium globulare habitat type. Associates: Pinus contorta, Abies lasiocarpa Rubus pedatus Sm. and Calamagrostis rubescens. 50% slope, north exposure. Soil type: Rocky, quartzite parent U.S.A., IDAHO, SHOSHONE COUNTY: material. Abundance 2 on a 1-5 scale. 8 miles northeast of Coeur d'Alene River on Shoshone Creek. Phenology: Fruits. Origin: Native. Elev. 2700 ft. 0-5% slope, soil deep, Belt Series; under mature Tsuga F. D. Johnson 2295 heterophylla/Clintonia uniflora habitat type, clambering over rotten with S. J. Brunsfeld logs. Occasional. Phenology: Vegetative. Origin: Native. 21 Aug 1984 Fred D. Johnson s.n. 13 Jun 1985 WTU-380635, WTU-V-045458 WTU-380645 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus bifrons Vest U.S.A., IDAHO, SHOSHONE COUNTY: U.S.A., OREGON, WHEELER COUNTY: Spion Kop Creek along Coeur d'Alene River, 10 Bear Creek, approximately 11 miles northwest of Mitchell. Bureau of miles northeast of Magee Ranger Station, Coeur Land Management land approximately 0.5 mile west of the Painted d'Alene National Forest. Hills Unit of the John Day Fossil Beds National Monument. Elev. 2900 ft. Elev. 1960 ft. T52N R3E S5; NAD 27, uncertainty: 805 m., 44.65418° N, 120.2972° W; NAD 27 Source: TRS2LL, Georef'd by Ben Legler Sandy, cobbly flood terrace near creek, in shade of Alnus 5% slope, southern exposure, stoney Belt rhombifolia. With Alnus rhombifolia, Salix lasiolepis, Salix Series soils; upper edge of riparian fringe in amygdaloides, Cornus sericea, Salix exigua. Stems arching, tips of shrubfield of Tsuga heterophylla/Clintonia primocanes rooting. Phenology: Fruits. Origin: Introduced. uniflora habitat type; common with Populus trichocarpa, Pinus contorta, Holodiscus discolor, R.E. Brainerd 972 14 Sep 2007 Salix scouleriana and Rosa nutkana. Height up to 1 meter. Phenology: Flowers. Origin: Native. WTU-382070 F. D. Johnson s.n. 22 Jun 1986 with C. L. Johnson Rosaceae WTU-380634, WTU-V-045461 Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SKAGIT COUNTY: Rosaceae Vendovi Island. Bluffs along southwest side of island. Rubus bifrons Vest Elev. 20 m. 48.608981°, -122.608264°; WGS 84, Source: U.S.A., WASHINGTON, WHATCOM COUNTY: Georeferenced, Georef'd by WTU Staff Growing along roadside at the end of Decatur Mix of open grassy meadows and Pseudotsuga Drive in Sandy Point Heights. menziesii groves. Common and widespread Elev. 10 ft. trailing on ground and across shrubs. UTM Zone 10, 523621E, 5405455N; T38N Phenology: Flowers. Origin: Native. R1E S10; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Peter W. Dunwiddie 2011-40 Other plants: Tanacetum vulgare, Daucus with Tanner Harris, Ben Legler, James carrota, Trifolium praetense, Leontodon Duemmel, John Floberg autumnalis. A few flowers pale pink, most white. Phenology: Flowers & Fruits. Origin: Introduced. 29 Apr 2011 James Duemmel 10-507 WTU-403916, WTU-V-012139 11 Aug 2010 WTU-381106, WTU-V-060738 Rosaceae Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, SKAGIT COUNTY: Vendovi Island. Near cove on north side of Rubus parviflorus Nutt. island. Elev. 5 m. : 48.6129°, -122.614°; WGS 84, uncertainty: 50 [No collection site is given on label]. m., Source: Georeferenced, Georef'd by WTU [No date given]. Phenology: Flowers. Origin: Staff Native. Open, mowed area around house and outbuildings. Common and widespread across unknown 37 island, mostly in wetter areas. Leaf material removed for RADseq analysis by Megan Ruffley WTU-381478, WTU-V-045457 at WTU, February 2016. Phenology: Flowers. Origin: Native. John Floberg 2011-11 with James Duemmel, Peter Dunwiddie, Tanner Harris, Ben Legler 1 May 2011 Rosaceae WTU-404055, WTU-V-012233 Rubus laciniatus Willd. U.S.A., WASHINGTON, KING COUNTY: Seattle, Discovery Park. 47.659354°, -122.417842°; WGS 84, uncertainty: 1400 m., Source: Digital Map, Georef'd by Ibro Polovina FN #91. Phenology: Flowers. Origin: Introduced. Cathy Bolles s.n. 6 Jun 1977 WTU-381698 Rosaceae Rosaceae Rubus arcticus L. Rubus idaeus L. RUSSIAN FEDERATION, SAKHALIN REGION: RUSSIAN FEDERATION, SAKHALIN REGION: Southern part of Sakhalin Island. West of Sokol. Approximately Northwestern tip of Sakhalin Island. Sakhalinsky Bay. Environs of halfway between Sea of Japan and Okhotsk Sea. Near Maly Takoy Uspenskoye Lake. River. Elev. 0 ft. Elev. 65 ft. 53° 27.471' N, 141° 58.368' E 47° 14.728' N, 142° 42.094' E Mixed forest of Picea dominant with Alnus. Open understory. Meadow with wet areas and small shaded irrigation slough. In Flowers past, fruit beginning. Frequent. Up to 60 centimeters tall. agricultural area. Pink flowers. Commonly scattered throughout taller Phenology: Fruits. Origin: Native. vegetation. Phenology: Flowers. Origin: Native. Suzanne Joneson 3267 12 Aug 2001 Suzanne Joneson 2758 20 Jul 2001 WTU-390628 WTU-391481 Rosaceae Rosaceae Rubus chamaemorus L. Rubus sachalinensis H. Lév. RUSSIAN FEDERATION, SAKHALIN REGION: RUSSIAN FEDERATION, SAKHALIN REGION: Northeast corner of Sakhalin Island. Sea of Okhotsk side. Environs Southern part of Sakhalin Island. Approximately 25 kilometers west of Longri River, south of Cape Longri. of Yuzhno-Sakhalinsk; north side of Tiobut River. Approximately 407 Elev. 0 ft. kilometers east of junction with Lyntoga River. 54° 1.273' N, 142° 57.162' E Elev. 656 ft. First small unnamed stream gulch, south of Longri River. Flowers 46° 58.639' N, 142° 14.876' E past. Common on slopes above stream. Phenology: Flowers. Origin: Hillside of dominantly Sasa, with occasional patches of Populus and Native. Betula. In flower (white.) Growing along road, as small shrub. Phenology: Flowers. Origin: Native. Suzanne Joneson 3554 16 Aug 2001 Suzanne Joneson 2927 26 Jul 2001 WTU-392027 WTU-390798 Rosaceae Rosaceae Rubus sachalinensis H. Lév. Rubus chamaemorus L. RUSSIAN FEDERATION, SAKHALIN REGION: Southeast corner of Sakhalin Island. West of Cape Menaputsy. RUSSIAN FEDERATION, SAKHALIN REGION: Elev. 196 ft. Northeast part of Sakhalin Island. Sea of Okhotsk side. North part of 46° 23.175' N, 143° 36.508' E Odoptu Lake, east of town of Tungor. Hillside and ravine at edge of Picea and Betula forest, with Avalia Elev. 39 ft. cordata growing in taller grasses. Flowers past (some still with white 53° 25.979' N, 143° 8.174' E petals), fruits beginning. Sporadic throughout the area, and growing Boggy area with ponds. Low vegetation with Myrica, Ledum and near the beach. Phenology: Fruits. Origin: Native. Juncus dominant. Iris and Pinus pumila occasional. Fruit turning red. One flower (white) present. Very common vegetatively. Growing Suzanne Joneson 3642 21 Aug 2001 throughout Empetrum, Ledum, and low herbs. Phenology: Fruits. Origin: Native. WTU-390590 Suzanne Joneson 3032 3 Aug 2001 Rosaceae WTU-390582 Rubus arcticus L. Rosaceae RUSSIAN FEDERATION, SAKHALIN REGION: Northeast part of Sakhalin Island. Sea of Okhotsk side. Environs of Rubus chamaemorus L. Taliki River. Elev. 0 ft. RUSSIAN FEDERATION, SAKHALIN REGION: 54° 16.483' N, 142° 49.111' E Northwestern tip of Sakhalin Island. Sakhalinsky Bay. Rocky beach shore meadow and cliffs. Northeastern exposure. Elev. 0 ft. Flowers magenta pink. Common vegetatively. Growing with Rosa. 53° 51.869' N, 142° 41.744' E Phenology: Flowers. Origin: Native. Hills above lake. Site of old forest fire. Ericaceous herbs and small shrubs dominant, with soil covered by lichens and mosses. In fruit, Suzanne Joneson 3351 14 Aug 2001 red turning to yellow. Common, covering large areas. Phenology: Fruits. Origin: Native. WTU-391434 Suzanne Joneson 3225 10 Aug 2001 WTU-390833 Rosaceae Rosaceae Rubus arcticus L. Rubus spectabilis Pursh RUSSIAN FEDERATION, SAKHALIN REGION: U.S.A., WASHINGTON, MASON COUNTY: Northeast part of Sakhalin Island. Sea of Okhotsk side. Environs of Olympic National Forest: along Fir Creek at Longri River, south of Cape Longri. junction of Forest Service Roads 23 and 2350. Elev. 0 ft. Elev. 284 m. 54° 1.728' N, 142° 58.052' E 47.353772° N, 123.321009° W; WGS 84, Flat meadow near beach, on old river plateau. Flowers pink, mostly uncertainty: 13 m., Source: GPS, Georef'd by past. Growing with Carex. Phenology: Flowers. Origin: Native. Ben Legler Moist forest of Pseudotsuga menziesii and Suzanne Joneson 3406 16 Aug 2001 Alnus sinuata along small stream; road shoulders with tall perennial grasses. Fruit WTU-391440 orange; shrubs to 2 meters tall under Alnus. Phenology: Fruits. Origin: Native. Rosaceae Ben Legler 12095 Rubus sachalinensis H. Lév. 24 Jul 2011 RUSSIAN FEDERATION, SAKHALIN REGION: WTU-409941, WTU-V-023998 Sakhalin Island; on southern part of island along the Nayba River circa 5 kilometers north-northwest of town of Bykov. Elev. 98 ft. 47° 59' 54.48" N, 142° 53' 56.52" E Rosaceae Along banks of river; substrate of gravel and silt; with Salix, Alnus, Crataegus, Carex, herbs, sedges. Flowers white, small; shrub under Rubus bifrons Vest trees. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, SKAGIT COUNTY: Ben Legler 705 17 Jul 2003 Vendovi Island. 48.6094° N, 122.6065° W; WGS 84, Herbarium: WTU uncertainty: 500 m., Source: Digital map Interior of island, under Pseudotsuga, Thuja, Acer macrophyllum canopy. Occasional, forming Rosaceae thickets, mostly along trails. Phenology: Fruits. Origin: Introduced. Rubus parviflorus Nutt. Peter Dunwiddie PD-2011-201 U.S.A., OREGON, GRANT COUNTY: Malheur National Forest. Trout Farm 10 Sep 2011 Campground. 15 miles southeast of Prairie City, Oregon. WTU-404028, WTU-V-012206 Elev. 4933 ft. 44.30439° N, 118.5518° W In wet areas along creek. Flowers white. Phenology: Flowers. Origin: Native. Rosaceae Richard G. Olmstead 2010-18 Rubus ursinus Cham. & Schltdl. 15 Jul 2010 U.S.A., WASHINGTON, KLICKITAT COUNTY: Mile point 1; BZ Corners-Glenwood Highway. WTU-400319, WTU-V-006824 45.86614°, -121.470663°; WGS 84, uncertainty: 5000 m., Source: Digital Map, Georef'd by Dan Post Road shoulder with annual forbs and grass. Pacific Blackberry. Phenology: Flowers. Origin: Native. Rosaceae David Gross 549 16 May 2010 Rubus ursinus Cham. & Schltdl. WTU-385539 U.S.A., WASHINGTON, MASON COUNTY: Olympic National Forest: shoulder of Forest Service Road 2352 ca. 1.8 road miles from its Rosaceae junction with Road 2350; 1.5 air miles SE of Spider Lake. Rubus leucodermis Douglas ex Torr. & A. Gray Elev. 620 m. 47.393554° N, 123.402209° W; WGS 84, U.S.A., WASHINGTON, JEFFERSON COUNTY: uncertainty: 13 m., Source: GPS, Georef'd by On Murhut Falls Road, Olympic National Forest, approximately 2 Ben Legler miles up from Falls trailhead. Moist road shoulder under mixed forest of Elev. 2000 ft. Pseudotsuga menziesii and Tsuga heterophylla. T25N R3W; WGS 84, uncertainty: 500 m., Source: Digital Map, Petals white; stems trailing on road side. Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Steep loamy soil. With Pseudotsuga menziesii, Acer glabrum, Cornus unalaschkensis. Common; fruits reddish-purple. Phenology: Ben Legler 12068 Fruits. Origin: Native. 23 Jul 2011 Regina Johnson 2011-027 9 Oct 2011 WTU-411998, WTU-V-026578 WTU-385621 Rosaceae Rosaceae Rubus vestitus Weihe & Nees Rubus spectabilis Pursh CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, SAN JUAN COUNTY: Greater Vancouver, Burnaby, railroad track 300 meters due east of Watmough Bay, southeast end of Lopez Island. Gaglardi Way, 270 meters due north of Highway 1. T34N R1W S21; WGS 84, uncertainty: 10000 m., Source: Elev. 75 ft. GeoLocate, Georef'd by WTU Staff 49° 14' 54" N, 122° 54' 30.8" W Edge of Pseudotsuga menziesii, Alnus rubra, Thuja plicata woods. Vegetated railroad track side at top of ravine. Semi shaded with Much branched shrub circa 3 meters tall. Flowers bright pink. Oemleria cerasiformis, polystichum munitum, Epilobium montanum, Phenology: Flowers & Fruits. Origin: Native. Rubus armeniacus, R. ursinus. Primocanes lax, horizontal arching,leaflets 5, but mostly 4 with lowest ones lobed. Phenology: Dorothy Naas 5755 19 Apr 1996 Flowers. Origin: Introduced. with Mildred Arnot Frank Lomer 6968 9 Jun 2009 WTU-387252 WTU-386001 Rosaceae Rosaceae Rubus pubescens Raf. Rubus lasiococcus A. Gray U.S.A., WASHINGTON, OKANOGAN COUNTY: Okanogan Highlands, Buckhorn Mountain, United States Forest U.S.A., WASHINGTON, WHATCOM COUNTY: Service. South Fork Bolster Creek. Mount Baker-Snoqualmie National Forest, Excelsior Pass Trail 2.75 Elev. 4000 ft. miles from the trailhead on State Route 542. T40N R30E S23 NW; NAD 27, uncertainty: 805 m., Source: Elev. 4100 ft. TRS2LL, Georef'd by Ben Legler UTM Zone 10, 587974E, 5419663N; T40N R8E S29; NAD 27, Plants growing along moist forested edge of creek. Associated uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler species: Pseudotsuga menziesii, Picea engelmannii, Cornus Growing beside the trail in a closed canopy Silver Fir/Western sericea, Lonicera involucrata, Symphoricarpos albus, Rubus Hemlock forest. Other plants: Vaccinium membranaceum, V. parviflorus, Rubus pedatus, Osmorhiza berteroi, Circaea alpina, ovalifolium, Rubus spectabilis, Tiarella trifoliata, Valeriana sitchensis, Cinna latifolia. Phenology: Flowers. Origin: Native. Abies amabilis, Tsuga heterophylla, Maianthemum stellatum, Athyrium filix-femina. Phenology: Flowers. Origin: Native. Kathryn A. Beck 201056 27 Jul 2010 James Duemmel 2011JD10 19 Aug 2011 WTU-387390 WTU-386556 Rosaceae Rosaceae Rubus pubescens Raf. Rubus parviflorus Nutt. U.S.A., WASHINGTON, OKANOGAN COUNTY: Okanogan Highlands, Buckhorn Mountain, United States Forest U.S.A., WASHINGTON, OKANOGAN COUNTY: Service. Lower south side of unnamed peak south of Joe Mills Mountain. Elev. 4100 ft. T40N R24E S14 NW of SW; NAD 27, uncertainty: 805 m., Source: T40N R31E S6 SW of NE; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler TRS2LL, Georef'd by Ben Legler In subalpine zone; 70% slope. Phenology: Flowers. Origin: Native. Plants growing in moist to wet seasonal, forested riparian area, in dark mucky soil. Aspect: 170 degrees. Slope: 2%. Associated P. Camp s.n. 24 Jun 1982 species: Picea engelmannii, Larix occidentalis, Acer douglasii, Symphoricarpos albus, Rubus parviflorus, Cornus canadensis, WTU-387191 Rubus pubescens, Viola glabella, Osmorhiza berteroi, Tiarella unifoliata, Carex disperma. Phenology: Flowers. Origin: Native. Rosaceae Kathryn A. Beck 201139 24 Jun 2011 Rubus ursinus Cham. & Schltdl. WTU-387374 U.S.A., WASHINGTON, SAN JUAN COUNTY: Lopez Island; Iceberg Point, 7 miles SSE of Lopez Village. Rosaceae T34N R2W S24; WGS 84, uncertainty: 500 m., Source: Georeferenced, Georef'd by WTU Staff Rubus idaeus L. ssp. strigosus (Michx.) Focke Edge of Pseudotsuga menziesii woods. Associates: Rosa nutkana, Sambucus racemosa, Poaceae spp. Trailing plant; petals white. U.S.A., OREGON, GRANT COUNTY: Phenology: Flowers. Origin: Native. Trout Farm Campground, Malheur National Forest. Elev. 4850 ft. Dorothy Naas 5717 17 May 1995 UTM Zone 11, 376152E, 4906938N; WGS 84, uncertainty: 7 m. Slope between pond and trail. Fruit red when ripe, separated from WTU-387255 receptacle. Stems of primocanes pinkish purple. Phenology: Fruits. Origin: Native. Barbara L. Wilson 16926 12 Sep 2011 with Nick Otting WTU-387464 Rosaceae Rosaceae Rubus bifrons Vest ex Tratt. × Rubus laciniatus Willd. Rubus allegheniensis Porter U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: North end of Lincoln Park, 0.75 air kilometers north-northeast of Near northeast shore of Christmas Lake; B. Wilson's station. Point Williams, Seattle. Elev. 967 ft. Elev. 13 ft. 47° 25.9' N, 121° 45.7' W 47° 32.2' N, 122° 23.7' W Dense partly shaded thickets, with Alnus rubra, Rhamnus purshiana, Thickets, base of steep west-facing slope above beach, partial Rubus ursinus, Rubus spectabilis, Rubus laciniatus, Rubus bifrons, shade, with putative parents, Prunus emarginata, Acer Pteridium. Arching shrubs 12-3 meters tall, nearly thornless, locally macrophyllum, Corylus, Rubus ursinus, R. spectabilis. Arching common. SHEET 1 OF 2. Phenology: Fruits. Origin: Introduced. shrubs 2 meters tall, with foliage intermediate. Phenology: Fruits. Origin: Introduced. Peter F. Zika 25730 24 Aug 2011 Arthur L. Jacobson 25729 5 Aug 2011 WTU-388212 WTU-388158 Rosaceae Rosaceae Rubus allegheniensis Porter Rubus bifrons Vest U.S.A., WASHINGTON, KING COUNTY: Near northeast shore of Christmas Lake; B. Wilson's station. U.S.A., WASHINGTON, KING COUNTY: Elev. 967 ft. Near northeast shore of Christmas Lake. 47° 25.9' N, 121° 45.7' W Elev. 967 ft. Dense partly shaded thickets, with Alnus rubra, Rhamnus purshiana, 47° 25.9' N, 121° 45.7' W Rubus ursinus, Rubus spectabilis, Rubus laciniatus, Rubus bifrons, Dense partly shaded thickets, with Alnus rubra, Rhamnus purshiana, Pteridium. Arching shrubs 12-3 meters tall, nearly thornless, locally Rubus ursinus, R. spectabilis, R. laciniatus, Pteridium. Arching common. SHEET 2 OF 2. Phenology: Fruits. Origin: Introduced. shrubs 1-2 meters tall, pink flowers, fruit black, subglobose, 12-20 x 13-19 millimeters, occasional. SHEET 2 OF 2. Phenology: Flowers & Peter F. Zika 25730 24 Aug 2011 Fruits. Origin: Introduced. WTU-388211 Peter F. Zika 25732 24 Aug 2011 WTU-388157 Rosaceae Rubus laciniatus Willd. Rosaceae U.S.A., WASHINGTON, KING COUNTY: Rubus bifrons Vest North end of Lincoln Park, 0.75 air miles north-northeast of Point Williams, Seattle. U.S.A., WASHINGTON, KING COUNTY: Elev. 13 ft. Near northeast shore of Christmas Lake. 47° 32.2' N, 122° 23.7' W Elev. 967 ft. Thickets, base of steep west-facing slope above beach, partial 47° 25.9' N, 121° 45.7' W shade, with Prunus emarginata, Acer macrophyllum, Corylus, Rubus Dense partly shaded thickets, with Alnus rubra, Rhamnus purshiana, ursinus, R. spectabilis. Arching shrubs 2 meters tall, fruit black, Rubus ursinus, R. spectabilis, R. laciniatus, Pteridium. Arching 13-22 x 14-18 millimeters, local. SHEET 2 OF 3. Phenology: Fruits. shrubs 1-2 meters tall, pink flowers, fruit black, subglobose, 12-20 x Origin: Introduced. 13-19 millimeters, occasional. SHEET 1 OF 2. Phenology: Flowers & Fruits. Origin: Introduced. P. F. Zika 25773 12 Sep 2011 with Arthur L. Jacobson Peter F. Zika 25732 24 Aug 2011 WTU-388448 WTU-388156 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus allegheniensis Porter U.S.A., WASHINGTON, KING COUNTY: CANADA, BRITISH COLUMBIA: North end of Lincoln Park, 0.75 air miles north-northeast of Point Southwest shore of Still Creek, 0.6 air kilometers north-northwest of Williams, Seattle. Burnaby Lake, Burnaby. Elev. 13 ft. Elev. 49 ft. 47° 32.2' N, 122° 23.7' W 49° 15.2' N, 122° 57.8' W Thickets, base of steep west-facing slope above beach, partial Damp thickets and margin of wet floodplain woods, with Alnus rubra, shade, with Prunus emarginata, Acer macrophyllum, Corylus, Rubus Populus tremuloides, Rubus laciniatus, R. armeniacus, R. ursinus, R. spectabilis. Arching shrubs 2 meters tall, fruit black, spectabilis, Impatiens glandulifera, Phalaris arundinacea. Arching 13-22 x 14-18 millimeters, local. SHEET 1 OF 3. Phenology: Fruits. shrubs 1.5-2.5 meters tall, fruit black, 12-17 x 12-15 millimeters, Origin: Introduced. common, naturalized. Phenology: Fruits. Origin: Introduced. P. F. Zika 25773 12 Sep 2011 P. F. Zika 25777 17 Sep 2011 with Arthur L. Jacobson with F. Lomer WTU-388447 WTU-388155 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus vestitus Weihe & Nees U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: North end of Lincoln Park, 0.75 air miles north-northeast of Point Center of Lincoln Park, 0.25 air kilometers northeast of Point Williams, Seattle. Williams, Seattle. Elev. 13 ft. Elev. 164 ft. 47° 32.2' N, 122° 23.7' W 47° 31.8' N, 122° 23.9' W Thickets, base of steep west-facing slope above beach, partial Shady thickets, with Thuja, Pseudotsuga, Alnus rubra, Oemleria, shade, with Prunus emarginata, Acer macrophyllum, Corylus, Rubus Polystichum munitum. Arching shrubs 1-2 meters tall, petals dark ursinus, R. spectabilis. Arching shrubs 2 meters tall, fruit black, pink. SHEET 2 OF 2. Phenology: Flowers. Origin: Introduced. 13-22 x 14-18 millimeters, local. SHEET 3 OF 3. Phenology: Fruits. Origin: Introduced. P. F. Zika 25728 22 Aug 2011 with Arthur L. Jacobson P. F. Zika 25773 12 Sep 2011 with Arthur L. Jacobson WTU-388437 WTU-388445 Rosaceae Rosaceae Rubus vestitus Weihe & Nees Rubus laciniatus Willd. U.S.A., WASHINGTON, KING COUNTY: Center of Lincoln Park, 0.25 air kilometers northeast of Point U.S.A., WASHINGTON, KING COUNTY: Williams, Seattle. Near northeast shore of Christmas Lake. Elev. 164 ft. Elev. 967 ft. 47° 31.8' N, 122° 23.9' W 47° 25.9' N, 121° 45.7' W Shady thickets, with Thuja, Pseudotsuga, Alnus rubra, Oemleria, Dense partly shaded thickets, with Alnus rubra, Rhamnus purshiana, Polystichum munitum. Arching shrubs 1-2 meters tall, petals dark Rubus ursinus, R. spectabilis, R. bifrons, Pteridium. Arching shrubs pink. SHEET 1 OF 2. Phenology: Flowers. Origin: Introduced. 1-2 meters tall, pink flowers, common. SHEET 1 OF 2. Phenology: Flowers. Origin: Introduced. P. F. Zika 25728 22 Aug 2011 with Arthur L. Jacobson Peter F. Zika 25733 24 Aug 2011 WTU-388440 WTU-388446 Rosaceae Rosaceae Rubus vestitus Weihe & Nees Rubus laciniatus Willd. U.S.A., OREGON, LANE COUNTY: U.S.A., WASHINGTON, KING COUNTY: East bank of Patterson Slough, just east of Autzen Stadium, west of Near northeast shore of Christmas Lake. Martin Luther King Boulevard, Delta Ponds, Eugene. Elev. 967 ft. Elev. 426 ft. 47° 25.9' N, 121° 45.7' W 44° 3.5' N, 123° 3.8' W Dense partly shaded thickets, with Alnus rubra, Rhamnus purshiana, Shaded thickets, near water, with Acer macrophyllum, Fraxinus, Rubus ursinus, R. spectabilis, R. bifrons, Pteridium. Arching shrubs Rubus bifrons, Oemleria. Pink flowers, 1 meter arching shrub, 1-2 meters tall, pink flowers, common. SHEET 2 OF 2. Phenology: common. Phenology: Flowers. Origin: Introduced. Flowers. Origin: Introduced. P. F. Zika 25610 13 Jul 2011 Peter F. Zika 25733 24 Aug 2011 with E. R. Alverson, B. Newhouse WTU-388443 WTU-388439

Rosaceae Rosaceae Rubus ulmifolius Schott Rubus bifrons Vest U.S.A., OREGON, LANE COUNTY: U.S.A., IDAHO, ADAMS COUNTY: Madison Meadow, greenbelt near junction of Payette National Forest; along Kleinschmidt Madison Street and 22nd Avenue, Eugene. Grade (Forest Road 050) circa 1/2 mile from Elev. 475 ft. bottom. 44° 2.1' N, 123° 6.2' W Elev. 2066 ft. Partly shaded thickets, with Rubus bifrons, 45.04854° N, 116.80674° W; WGS 84, Daucus. Bright pink flowers (darker pink than R. uncertainty: 3 m., Source: GPS bifrons here), common naturalized arching Dry, rocky slope and rock road cut, with unarmed shrub. Specimen used for illustration in bunchgrasses, Hackelia, Purshia. Petals very Revision of Flora of the Pacific Northwest, WTU pale pink; clambering shrubs, local in draw Herbarium, 2015. Phenology: Flowers. Origin: beneath Morus. Phenology: Flowers. Origin: Introduced. Introduced. P. F. Zika 25597 Lacie Schulte 47 with E. R. Alverson, B. Newhouse 25 Jun 2011 13 Jul 2011 WTU-406284, WTU-V-17042 WTU-388441, WTU-V-021998 Rosaceae Rosaceae Rubus bifrons Vest Rubus parviflorus Nutt. U.S.A., IDAHO, ADAMS COUNTY: U.S.A., WASHINGTON, KING COUNTY: Payette National Forest. East side of Hells Carkeek Park, Ballard, Seattle. Canyon Road. Elev. 400 ft. 45.10892° N, 116.76373° W; WGS 84, 47.709542°, -122.370423°; WGS 84, uncertainty: 1000 m., uncertainty: 30 m., Source: GPS Source: Digital Map, Georef'd by Joseph Stormer West-facing rocky outcropping alongside road; Woody shrub/tree; approximately 4 feet in height; palmately veined; Ericameria resinosa common. Uncommon along white flowers; many stamens. Collected for Biology 317 (Plant base of rocky outcropping. Phenology: Fruits. Identification and Classification). Phenology: Flowers. Origin: Native. Origin: Introduced. June Landenburger 1219 19 May 2012 David Giblin 3910 WTU-389784 26 Jun 2011 WTU-403516, WTU-V-011104 Rosaceae Rubus ursinus Cham. & Schltdl. Rosaceae U.S.A., WASHINGTON, KING COUNTY: Carkeek Park, Ballard, Seattle. Rubus parviflorus Nutt. Elev. 200 ft. 47.709893°, -122.371633°; WGS 84, uncertainty: 1000 m., U.S.A., IDAHO, ADAMS COUNTY: Source: Digital Map, Georef'd by Ibro Polovina Payette National Forest. Confluence of Crooked On hillside and amongst Rubus spectabilis. Viny with small thorns; River and Wildhorse River; 3.7 miles down white flowers. Collected for Biology 317 (Plant Identification and Wildhorse River from junction with Council Classification). Phenology: Flowers. Origin: Native. Cuprum Road. Elev. 3378 ft. June Landenburger 1213 6 May 2012 45.95865° N, 116.7246° W; WGS 84, uncertainty: 500 m., Source: Nat. Geog. WTU-389781 TOPO!, Georef'd by John Bassett Riparian habitat, roadside, and dry rocky slopes. White-flowered shrubs. Phenology: Flowers. Rosaceae Origin: Native. Rubus spectabilis Pursh Richard G. Olmstead 2011-47 U.S.A., WASHINGTON, KING COUNTY: 24 Jun 2011 Carkeek Park, Ballard, Seattle. Elev. 200 ft. WTU-405345, WTU-V-015699 47.709542°, -122.371796°; WGS 84, uncertainty: 1104 m., Source: GeoLocate, Georef'd by Joseph Stormer Gravelly slope descending into creek bed and surrounded by wooded hill crests. Petals magenta; many stamens; aggregate fruit. Rosaceae Collected for Biology 317 (Plant Identification and Classification). Phenology: Flowers. Origin: Native. Rubus idaeus L. ssp. strigosus (Michx.) Focke June Landenburger 1201 15 Apr 2012 U.S.A., IDAHO, ADAMS COUNTY: Payette National Forest; Kleinschmidt Grade WTU-390035 Road (Forest Service Road 050) circa halfway down to Snake River. Elev. 3050 ft. Rosaceae 45.05684° N, 116.78396° W; WGS 84, uncertainty: 3 m., Source: GPS Rubus laciniatus Willd. Steep, rocky west-facing slope with draws and outcrops; scattered shrubs, bunchgrasses. U.S.A., WASHINGTON, THURSTON COUNTY: Shrubs circa 1 meter tall along road cuts. Glacial Heritage Preserve. Phenology: Fruits. Origin: Native. Elev. 164 ft. 46.85865° N, 123.05176° W; UTM Zone 10, 496055E, 5189458N; Don Knoke 2088 NAD 83 Along roadways. Campanula rotundifolia, Galium sp., Holcus 25 Jun 2011 lanatus, Agrostis sp., Hypericum formosum, Carex inops, Rumex acetosella, Hypochaeris radicata, Arrhenatherum elatius. Flowers WTU-406087, WTU-V-17246 white, scattered. Phenology: Flowers & Fruits. Origin: Introduced. Pat Montague 116 1 Jul 2008 with Dan Montague WTU-389907 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus pensilvanicus Poir. U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Vendovi Island. Near cove on north side of North bank of Salmon River, mouth of Grouse island. In general vicinity of house. Creek, 0.2 miles east of Jim Moore Place, 2 air Elev. 16-65 ft. miles southeast of Rabbit Point. 48° 36' 46.44" N, 122° 36' 50.4" W; WGS 84, Elev. 2361 ft. uncertainty: 50 m., Source: Digital map 45° 29.1' N, 115° 20.57' W Open, mowed area around house and Seen naturalized 3 places in shady thickets over outbuildings. Common at edge of forest where it granite, with Amelanchier, Philadelphus, can get sun. Phenology: Flowers. Origin: Native. Toxicodendron, Berberis aquifolium, Symphoricarpos, Prunus virginiana, Rubus Peter Dunwiddie 2011-48 leucodermis. Arching shrub 1.5 meters tall, fruit black; perhaps introduced in vicinity many 1 Jun 2011 decades ago by settlers. Specimen used for illustration in Revision of Flora of the Pacific WTU-404326, WTU-V-012391 Northwest, WTU Herbarium, 2015. Phenology: Fruits. Origin: Introduced. Peter F. Zika 26010 29 Jul 2012 Rosaceae WTU-390317, WTU-V-021996 Rubus spectabilis Pursh U.S.A., WASHINGTON, WHATCOM COUNTY: Rosaceae Viewcrest Ridge, circa 2 miles south of Bellingham. Rubus pedatus Sm. 48.69255°, -122.481702°; WGS 84, uncertainty: 3200 m., Source: Digital Map, U.S.A., WASHINGTON, PACIFIC COUNTY: Georef'd by Joseph Stormer KO Peak in Willapa Hills. In gravely loam. Petals purplish-red, shrub circa Elev. 2700 ft. 1 meter tall. Phenology: Flowers. Origin: Native. T11N R7W Sec. 4; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Carl A. Linde Jr. s.n. Beneath overstory of Vaccinium alaskaense and Rubus spectabilis. Phenology: Flowers. Origin: Native. 19 Apr 1957 Cathy Maxwell 128 31 May 1981 WTU-390380, WTU-V-042599 with Ed Maxwell WTU-392738 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., IDAHO, IDAHO COUNTY: North bank of Salmon River, near mouth of Grouse Creek, 0.2 miles U.S.A., OREGON, GRANT COUNTY: east of Jim Moore Place, 2 air miles southeast of Rabbit Point. Malheur National Forest. Strawberry Lake. Elev. 2361 ft. Northern shores of the lake, where the trail runs 45° 29.1' N, 115° 20.57' W by the shore. In shady thickets over granite, with Amelanchier, Philadelphus, Elev. 6310 ft. Toxicodendron, Berberis aquifolium, Symphoricarpos, Prunus 44° 18.571' N, 118° 40.914' W virginiana. Stem glaucous, fruit dark purple to black; arching 1.5 tall Open, coniferous woodland. Infrequent by lake shrub, uncommon. Phenology: Fruits. Origin: Native. shore. Flowers white. Prickly shrub to 1.5 meters. Phenology: Flowers. Origin: Native. Peter F. Zika 26011 29 Jul 2012 Wendy McClure 10-31 WTU-390342 16 Jul 2010 WTU-401000, WTU-V-8180 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus pubescens Raf. U.S.A., OREGON, GRANT COUNTY: U.S.A., WASHINGTON, PEND OREILLE Malheur National Forest. Strawberry COUNTY: Campground. Flat open areas around Colville National Forest; along Lost Creek at campground and campsites. junction of Forest Service Road 4685 and spur Elev. 5766 ft. road 550; in wetland circa 100 meters northeast 44° 19.169' N, 118° 40.449' W of junction. Along banks of stream and in open coniferous Elev. 3432 ft. woodland with pine, fir, larch. Erect shrub to 1 48.68087° N, 117.52019° W; WGS 84, meter, infrequent in creek banks; flowers white. uncertainty: 50 m. Phenology: Flowers. Origin: Native. Wet meadow with fen in center, dominated by Sphagnum, Carex spp., Drosera, Alnus sinuata, Wendy McClure 10-52 and Picea engelmannii. Fruits deep red; stems low, from rhizomes; frequent. Phenology: Fruits. 16 Jul 2010 Origin: Native. WTU-401008, WTU-V-8188 Karen Peterson 2012-11 20 Jul 2012 Rosaceae WTU-408670, WTU-V-022449 Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, KING COUNTY: Green River bike route. Rubus parviflorus Nutt. 47.440374°, -122.232285°; WGS 84, uncertainty: 10129 m., Source: Digital Map, U.S.A., WASHINGTON, PEND OREILLE Georef'd by Joseph Stormer COUNTY: Sandy soil open field. Phenology: Flowers. Colville National Forest. Forest Road 245 circa Origin: Native. 5.8 miles above junction of Forest Road 250 and Sullivan Creek Road, on road to Sullivan Jerry Sheldon 3 Mountain. Elev. 5046 ft. 17 Apr 1973 48.86627° N, -117.22564° W; WGS 84, uncertainty: 500 m., Source: Nat. Geog. WTU-390493, WTU-V-18371 TOPO!, Georef'd by John Bassett Flowers white. Phenology: Flowers. Origin: Native. Rosaceae Melissa Barker 2012-13 Rubus parviflorus Nutt. 21 Jul 2012 U.S.A., WASHINGTON, WHATCOM COUNTY: WTU-407507, WTU-V-020581 Above Clayton Bay, circa 9 miles south of Bellingham. 48.64655°, -122.485135°; WGS 84, uncertainty: 1300 m., Source: Digital Map, Georef'd by Joseph Stormer Rosaceae In dry soil in open area alongside road. Shrub circa 2 decimeters tall. Phenology: Flowers. Origin: Native. Rubus idaeus L. ssp. strigosus (Michx.) Focke Carl A. Linde Jr. s.n. 7 May 1957 U.S.A., WASHINGTON, PEND OREILLE COUNTY: WTU-391173 Colville National Forest. Approximately 8 kilometers south on Dry Canyon Road from junction with Sullivan Lake Road. Elev. 900 m. 48.6965° N, 117.3498° W; WGS 84, uncertainty: 100 m., Source: Digital map, Georef'd by Maria Yousoufian East-facing roadside edge with large-bouldered talus slope above. Philadelphus lewisii common; boulders covered with lichens and moss. Uncommon; fruits red; shrubs to 1 meter. Phenology: Fruits. Origin: Native. Nisa Karimi 12-1 20 Jul 2012 WTU-406934, WTU-V-019665 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, PEND OREILLE U.S.A., WASHINGTON, PEND OREILLE COUNTY: COUNTY: Colville National Forest. Along Round Top Colville National Forest. Sullivan Creek Road, Mountain Trail #512 (Shedroof Divide trail). 10.2 miles from junction with Sullivan Lake Elev. 1830 m. Road. 48.8061° N, 117.1298° W; WGS 84, Elev. 887 m. uncertainty: 1000 m., Source: Digital map, 48.88888° N, 117.09759° W Georef'd by Maria Yousoufian Flowers white. Phenology: Flowers. Origin: Mixed coniferous open forest along trail. Some Native. rocky outcrops and seeps. Phenology: Flowers. Origin: Native. Kathleen Whitson 2012-24 Susan Kephart 763 20 Jul 2012 21 Jul 2012 WTU-406876, WTU-V-019592 WTU-407926, WTU-V-021153 Rosaceae Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, THURSTON COUNTY: Rubus vestitus Weihe & Nees Logging road above southern junction of US 101 and Old Highway 101, just south of Mason U.S.A., OREGON, LANE COUNTY: county line. 0.45 air kilometers southwest of summit of Spencers Butte, Eugene. Elev. 100 ft. Elev. 450 m. 47° 5' 19.7" N, 123° 3' 58.29" W 43.981° N, 123.1° W Silt loam. With Alnus rubra, Abies grandis, In shade of Pseudotsuga, near meadow. Pink flowers, arching Pseudotsuga menziesii, Oemleria cerasiformis, shrubs 1-1.5 meters tall, forming a dense stand. Phenology: Flowers Rubus armeniacus. Pseudotsuga and Fruits. Origin: Introduced. menziesii/Abies grandis forest. Locally common. Leaf material removed for RADseq analysis by P. F. Zika 26307 16 Jul 2013 Megan Ruffley at WTU, February 2016. with E. R. Alverson, B. N. Newhouse Phenology: Flowers. Origin: Native. WTU-393998 Regina Johnson 2013-013 14 Apr 2013 Rosaceae WTU-394399, WTU-V-019098 Rubus vestitus Weihe & Nees U.S.A., OREGON, LANE COUNTY: 0.45 air kilometers southwest of summit of Spencers Butte, Eugene. Rosaceae Elev. 450 m. 43.981° N, 123.1° W Rubus spectabilis Pursh In shade of Pseudotsuga, near meadow. Pink flowers, arching shrubs 1-1.5 meters tall, forming a dense stand. Phenology: U.S.A., WASHINGTON, MASON COUNTY: Flowers. Origin: Introduced. Olympic National Forest, South Fork Skokomish, Forest Road 23 about 0.5 mile P. F. Zika 26307 16 Jul 2013 uphill from Forest Road 2351. with E. R. Alverson, B. N. Newhouse Elev. 1000 ft. 47.357032° N, 123.320925° W; T22N R5W WTU-393900 S33 Sandy loam. With Alnus rubra, Populus trichocarpa, Oemleria cerasiformis, Acer Rosaceae circinatum, Rubus ursinus. Riparian alder grove in Pseudotsuga menziesii/Tsuga heterophylla Rubus pedatus Sm. forest. Abundant. Leaf material removed for RADseq analysis by Megan Ruffley at WTU, U.S.A., WASHINGTON, PEND OREILLE February 2016. Phenology: Flowers. Origin: COUNTY: Native. Colville National Forest. Sullivan Creek Road, 10.2 miles from junction with Sullivan Lake Regina Johnson 2013-025 28 Apr 2013 Road. Elev. 887 m. WTU-395366, WTU-V-019096 48.88888° N, 117.09759° W Flowers white. Phenology: Flowers. Origin: Native. Kathleen Whitson 2012-4 20 Jul 2012 WTU-407464, WTU-V-020927 Rosaceae Rosaceae Rubus bifrons Vest Rubus parviflorus Nutt. U.S.A., OREGON, GRANT COUNTY: U.S.A., MONTANA, GRANITE COUNTY: Highway 395; 2.4 air miles south of Canyon City. Lolo National Forest. Edith Peak Road. Elev. 3500 ft. Elev. 5460 ft. UTM Zone 10, 344327E, 4913142N; WGS 84 47.1069° N, 114.34924° W; WGS 84, Disturbed waste ground and eroding slopes. With Thinopyrum uncertainty: 500 m., Source: GPS, Invalid intermedium Populus trichocarpa, Clematis ligusticifolia, Rosa. coordinates (located outside county bounds). Flowers light pink. Plants forming dense mounds. Phenology: Mostly open forest dominated by Pinus contorta, Flowers. with Larix occidentalis and Pseudotsuga menziesii, scattered Abies amabilis. Understory R. E. Brainerd 2000 29 Sep 2011 dominanted by Xerophyllum tenax, Carex rossii, Vaccinium, Lupinus, Ceanothus, Arctostaphylos. WTU-394879 Common to dominant along old closed road. Phenology: Flowers. Origin: Native. Rosaceae Ryan Boyle 99 Rubus parviflorus Nutt. 13 Jul 2008 U.S.A., WASHINGTON, PEND OREILLE WTU-399824, WTU-V-006114 COUNTY: Colville National Forest. Slate Creek, 7 miles from State Route 31 along Forest Road 3155. Approximately 1 mile after trail, at road closure. Rosaceae Elev. 4197 ft. 48.96341° N, 117.20316° W Rubus parviflorus Nutt. Sloping hillside, gravel road. Mixed coniferous montane forest. Shrub to 1.5 meters; common. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Phenology: Flowers. Origin: Native. Wynoochee Falls. Elev. 1000 ft. Katie Parker 13 47° 28.278' N, 123° 31.569' W Clay loam. With Rubus spectabilis, Ribes bracteosum, Acer 20 Jul 2012 circinatum, Tsuga heterophylla, Dicentra formosa, Athyrium filix-femina. Riparian area in Tsuga heterophylla forest. Common. WTU-407745, WTU-V-021824 Phenology: Flowers. Origin: Native. Regina Johnson 2013-079 4 Jul 2013 Rosaceae WTU-397087 Rubus vestitus Weihe & Nees U.S.A., OREGON, LINN COUNTY: Rosaceae Parking area at north end of Green Peter Dam; 9.8 air miles east-northeast of Sweet Home. Rubus pedatus Sm. Elev. 1040 ft. UTM Zone 10, 535683E, 4922081N; WGS 84 U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Shady seepy bank at edge of Douglas-fir-bigleaf maple forest. With Wynoochee Pass Trail, in southern Olympics. Pseudotsuga menziesii, Acer macrophyllum, Athyrium filix-femina, Elev. 2500 ft. Rhamnus purshiana, Carex amplifolia. Sprawling shrub to 1 meter 47° 31.009' N, 123° 32.302' W tall, arching canes much longer; inflorescence axis glandular hairy; Steep rocky sandy loam. With Vaccinium parviflorum, Tsuga flowers light pink. Phenology: Flowers. Origin: Introduced. heterophylla, Polystichum munitum. Steep Tsuga heterophylla forest. Common. Phenology: Flowers. Origin: Native. R.E. Brainerd 2168 28 Jun 2013 Regina Johnson 2013-082 5 Jul 2013 WTU-396492 WTU-397084 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus idaeus L. var. sachalinensis (H. Lév.) Focke U.S.A., OREGON, LANE COUNTY: Deschutes National Forest. Pacific Crest Trail at U.S.A., ALASKA COUNTY: stream crossing east of Island Meadow. Fairbanks. Elev. 1651 m. 64.837778°, -147.716389°; WGS 84, 43.97501° N, 121.85798° W uncertainty: 10000 m., Source: GeoLocate, Moist streamside. White flowers. Phenology: Georef'd by Joseph Stormer Flowers. Origin: Native. Phenology: Fruits. Origin: Native. Richard G. Olmstead 2013-19 Clayton Haberman s.n. 12 Jul 2013 21 Jul 1990 WTU-412024, WTU-V-026517 WTU-399319, WTU-V-005134 Rosaceae Rosaceae Rubus bifrons Vest Rubus parviflorus Nutt. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: East shore Port Susan, north Tulare Beach. Near Arlington. 48.105718°, -122.34509°; WGS 84, 48.19889°, -122.12389°; WGS 84, uncertainty: 367 m., Source: Digital Map, uncertainty: 5000 m., Source: GeoLocate, Georef'd by Maria Yousoufian Georef'd by WTU Staff Phenology: Flowers. Origin: Introduced. Field. Phenology: Fruits. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 29 May 1987 Sep 2008 WTU-399321, WTU-V-005135 WTU-399325, WTU-V-005139

Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Happy Valley Tree Farm. East shore Port Susan, north Tulare Beach. 48.187751°, -122.262495°; WGS 84, 48.105718°, -122.34509°; WGS 84, uncertainty: 264 m., Source: Digital Map, uncertainty: 367 m., Source: Digital Map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Phenology: Flowers. Origin: Native. Base of sandy-clay bluff. "Large flower" specimen. Phenology: Flowers. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 5 May 1998 13 May 2005 WTU-399322, WTU-V-005136 WTU-399326, WTU-V-005140 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, SNOHOMISH Happy Valley Tree Farm. COUNTY: 48.187751°, -122.262495°; WGS 84, Happy Valley Tree Farm, Happy Valley. uncertainty: 264 m., Source: Digital Map, 48.187751°, -122.262495°; WGS 84, Georef'd by Maria Yousoufian uncertainty: 264 m., Source: Digital map, Phenology: Flowers. Origin: Native. Georef'd by Maria Yousoufian Phenology: Vegetative. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 21 May 1990 28 Jan 2003 WTU-399323, WTU-V-005137 WTU-399327, WTU-V-005168 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus parviflorus Nutt. U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, SNOHOMISH Arlington area. COUNTY: 48.198712°, -122.125142°; WGS 84, East Warm Beach Senior Community, uncertainty: 10000 m., Source: GeoLocate, Stanwood. Georef'd by Ibro Polovina 48.181907°, -122.348357°; WGS 84, Phenology: Vegetative. Origin: Native. uncertainty: 224 m., Source: Digital Map, Georef'd by Maria Yousoufian Clayton Haberman s.n. Trailside, mesic, mixed woods. Phenology: Fruits. Origin: Native. Feb 2009 Clayton Haberman s.n. WTU-399324, WTU-V-005138 2 Aug 2010 WTU-399328, WTU-V-005169 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus spectabilis Pursh U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: East Warm Beach Senior Community. Tulare Beach. 48.181907°, -122.348357°; WGS 84, 48.101763°, -122.343202°; WGS 84, uncertainty: 224 m., Source: Digital map, uncertainty: 650 m., Source: GeoLocate, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Trail side, mixed mesic woods. [Attached packet Garden weed seedling. Phenology: Vegetative. says Happy Valley Tree Farm, Happy Valley, Origin: Native. 6-20-2009.] Phenology: Fruits. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 9 May 1993 18 Aug 2009 WTU-399333, WTU-V-005174 WTU-399329, WTU-V-005170 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus parviflorus Nutt. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Happy Valley Tree Farm. East Warm Beach Senior Community, 48.187751°, -122.262495°; WGS 84, Stanwood. uncertainty: 264 m., Source: Digital Map, 48.181907°, -122.348357°; WGS 84, Georef'd by Maria Yousoufian uncertainty: 224 m., Source: Digital Map, Phenology: Flowers. Origin: Native. Georef'd by Maria Yousoufian Trailside mixed mesic woods. Phenology: Clayton Haberman s.n. Flowers. Origin: Native. 28 Mar 2000 Clayton Haberman s.n. WTU-399334, WTU-V-005175 9 Jun 2009 WTU-399330, WTU-V-005171 Rosaceae Rosaceae Rubus pedatus Sm. Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: United States 2. U.S.A., WASHINGTON, SNOHOMISH 47.710977°, -121.301686°; WGS 84, COUNTY: uncertainty: 17000 m., Source: Digital Map, Happy Valley Tree Farm. Georef'd by Ibro Polovina 48.187751°, -122.262495°; WGS 84, Roadside. Evergreen forest. Phenology: uncertainty: 264 m., Source: Digital Map, Vegetative. Origin: Native. Georef'd by Maria Yousoufian Phenology: Flowers. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 15 May 2004 20 May 1999 WTU-399335, WTU-V-005176 WTU-399331, WTU-V-005172 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus spectabilis Pursh U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: East Warm Beach Senior Community, Happy Valley Tree Farm, Happy Valley. Stanwood. 48.187751°, -122.262495°; WGS 84, 48.181907°, -122.348357°; WGS 84, uncertainty: 264 m., Source: Digital map, uncertainty: 224 m., Source: Digital Map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Phenology: Vegetative. Origin: Native. Trailside, mesic, mixed woods. Phenology: Fruits. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 28 Jan 2003 7 Aug 2010 WTU-399332, WTU-V-005173 WTU-339336, WTU-V-005177 Rosaceae Rosaceae Rubus chamaemorus L. Rubus chamaemorus L. U.S.A., ALASKA COUNTY: U.S.A., ALASKA, SITKA COUNTY: Kotzebue. Sitka. 66.877849°, -162.453845°; WGS 84, 56.966317°, -135.207777°; WGS 84, uncertainty: 10000 m., Source: Digital Map, uncertainty: 106622 m., Source: Digital Map, Georef'd by Joseph Stormer Georef'd by Joseph Stormer Phenology: Fruits. Origin: Native. Sphagnum bog. Phenology: Flowers. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 8 Jul 1990 8 Jun 1994 WTU-399367, WTU-V-005196 WTU-399371, WTU-V-005200

Rosaceae Rosaceae Rubus arcticus L. Rubus bifrons Vest CANADA, BRITISH COLUMBIA: Summit Lake, Mile 383 Alaska Highway. U.S.A., WASHINGTON, SNOHOMISH Phenology: Flowers. Origin: Native. COUNTY: Happy Valley Tree Farm, Happy Valley. Clayton Haberman s.n. 48.187751°, -122.262495°; WGS 84, uncertainty: 264 m., Source: Digital map, 28 Jun 1990 Georef'd by Maria Yousoufian Phenology: Vegetative. Origin: Introduced. WTU-399368, WTU-V-005197 Clayton Haberman s.n. Feb 2009 WTU-399374, WTU-V-005202 Rosaceae Rosaceae Rubus arcticus L. Rubus bifrons Vest U.S.A., ALASKA COUNTY: Lower Troublesome Creek, Mile 137 Parks U.S.A., WASHINGTON, SNOHOMISH Highway. COUNTY: Phenology: Flowers. Origin: Native. East shore Port Susan, north Tulare Beach. 48.105718°, -122.34509°; WGS 84, Clayton Haberman s.n. uncertainty: 367 m., Source: Digital Map, Georef'd by Maria Yousoufian 9 Jul 1990 Phenology: Flowers. Origin: Introduced. WTU-399369, WTU-V-005198 Clayton Haberman s.n. Jun 1989 WTU-399375, WTU-V-005203 Rosaceae Rosaceae Rubus chamaemorus L. Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: Safeway parking area, North Bend. U.S.A., WASHINGTON, SNOHOMISH 47.48733°, -121.791499°; WGS 84, COUNTY: uncertainty: 200 m., Source: Digital Map, East Warm Beach Senior Community, Georef'd by Ibro Polovina Stanwood. Landscape ground cover. Berry yellow. 48.181907°, -122.348357°; WGS 84, Phenology: Fruits. Origin: Native. uncertainty: 224 m., Source: Digital Map, Georef'd by Maria Yousoufian Clayton Haberman s.n. Mixed mesic woods, trail. Sprout from second year wood. Phenology: Vegetative. Origin: 9 Aug 2002 Native. WTU-399370, WTU-V-005199 Clayton Haberman s.n. 22 Jun 2009 WTU-399413, WTU-V-005217 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: East Warm Beach Senior Community, East Warm Beach Senior Center, Stanwood. Stanwood. 48.181907°, -122.348357°; WGS 84, 48.181907°, -122.348357°; WGS 84, uncertainty: 224 m., Source: Digital map, uncertainty: 224 m., Source: Digital Map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Opening, mesic mixed woods. Phenology: Trailside, mixed, mesic woods. First year ground Flowers. Origin: Native. shoot. Phenology: Vegetative. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 21 Apr 2010 4 May 2009 WTU-399418, WTU-V-005222 WTU-399414, WTU-V-005218 Rosaceae Rosaceae Rubus spectabilis Pursh Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: East of Warm Beach Senior Community, Tulare Beach. Stanwood. 48.101763°, -122.343202°; WGS 84, 48.181907°, -122.348357°; WGS 84, uncertainty: 650 m., Source: GeoLocate, uncertainty: 224 m., Source: Digital map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Roadside. Male. Phenology: Flowers. Origin: Trail side, mixed woods. Phenology: Flowers. Native. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 3 May 2006 6 Apr 2009 WTU-399419, WTU-V-005223 WTU-399415, WTU-V-005219 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus spectabilis Pursh U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Tulare Beach. Happy Valley Tree Farm, Happy Valley. 48.101763°, -122.343202°; WGS 84, 48.187751°, -122.262495°; WGS 84, uncertainty: 650 m., Source: GeoLocate, uncertainty: 264 m., Source: Digital map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Roadside. Female. Phenology: Flowers. Origin: Phenology: Vegetative. Origin: Native. Native. Clayton Haberman s.n. Clayton Haberman s.n. Feb 2009 3 May 2006 WTU-399416, WTU-V-005220 WTU-399420, WTU-V-005224

Rosaceae Rosaceae Rubus spectabilis Pursh Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Happy Valley Tree Farm, Happy Valley. Happy Valley Tree Farm. 48.187751°, -122.262495°; WGS 84, 48.187751°, -122.262495°; WGS 84, uncertainty: 264 m., Source: Digital map, uncertainty: 264 m., Source: Digital Map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Part shade. Seedling with stolon. Phenology: Phenology: Flowers. Origin: Native. Vegetative. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 21 May 1990 23 May 2010 WTU-399421, WTU-V-005225 WTU-399417, WTU-V-005221 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus bifrons Vest U.S.A., WASHINGTON, SNOHOMISH U.S.A., OREGON, GRANT COUNTY: COUNTY: Camp Creek, a major tributary of the Middle East Warm Beach Senior Community, Fork John Day River, a little upstream of a Stanwood. concrete bridge crosses the creek. 48.181907°, -122.348357°; WGS 84, Elev. 3760 ft. uncertainty: 224 m., Source: Digital Map, 44.6592° N, 118.82098° W; WGS 84, Georef'd by Maria Yousoufian Source: Specimen Label Trailside, opening, mesic mixed woods. Male. In light shade, with weedy grasses, near wide Sheet 2 of 2. Phenology: Flowers. Origin: place at edge of road. One plant with several Native. stems coming from one root crown. About 1 meter tall. Phenology: Vegetative. Origin: Clayton Haberman s.n. Introduced. 26 Apr 2010 Barbara L. Wilson 18343 with Richard E. Brainerd WTU-399423, WTU-V-005226 Rosaceae 8 Oct 2014 Rubus ursinus Cham. & Schltdl. WTU-401503, WTU-V-8642 U.S.A., WASHINGTON, SNOHOMISH COUNTY: East Warm Beach Senior Community, Rosaceae Stanwood. 48.181907°, -122.348357°; WGS 84, Rubus leucodermis Douglas ex Torr. & A. Gray uncertainty: 224 m., Source: Digital Map, Georef'd by Maria Yousoufian U.S.A., WASHINGTON, KITTITAS COUNTY: Trailside, opening, mesic mixed woods. Female. Along Forest Road 1115 0.1 miles from United Sheet 2 of 2. Phenology: Flowers. Origin: States Highway 97. Native. Elev. 750 m. 47.24805° N, 120.69647° W; T20N R17E S3; Clayton Haberman s.n. WGS 84 Fairly moist soil; 20° south facing slope; open 26 Apr 2010 forest. Other plants: Symphoricarpos albus, Abies grandis, Holodiscus discolor, Osmorhiza WTU-399424, WTU-V-005227 chilensis. Flowers white; occasional. Phenology: Rosaceae Flowers. Origin: Native. Rubus bifrons Vest Don Knoke 2379 U.S.A., OREGON, SHERMAN COUNTY: 11 Jun 2013 Deschutes River State Recreation Area, 13.3 miles east of The Dalles. WTU-409661, WTU-V-023649 Elev. 200 ft. UTM Zone 10, 663076E, 5054997N; WGS 84 Weedy riparian woodland; partial shade. With Alnus rhombifolia, Conium maculatum, Phalaris Rosaceae arundinacea, Urtica dioica. Flowers light pink; prickles straight to slightly curved. Phenology: Rubus parviflorus Nutt. Flowers. Origin: Introduced. U.S.A., WASHINGTON, KITTITAS COUNTY: R. E. Brainerd 1509 Along Forest Road 1115 0.2 miles from United States Highway 97. 22 Jun 2009 Elev. 750 m. 47.24805° N, 120.69647° W; T20N R17E S3; WTU-400096, WTU-V-006292 WGS 84 Moist soil near stream bank; 15° south-facing slope; coniferous forest. Other plants: Populus trichocarpa, Symphoricarpos albus, Rosaceae Pseudotsuga menziesii, Cornus stolonifera. Flowers white. Phenology: Flowers. Origin: Rubus parviflorus Nutt. Native. U.S.A., WASHINGTON, KLICKITAT COUNTY: Don Knoke 2424 Along Northwestern Lake Road. 45.779874°, -121.515627°; WGS 84, 4 Jul 2013 uncertainty: 627 m., Source: Digital Map, Georef'd by Joseph Stormer WTU-409195, WTU-V-023207 Shady roadside with grass, forbs. Douglas-fir forest. Phenology: Flowers. Origin: Native. David Gross 997 21 May 2014 WTU-400786, WTU-V-007302 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., MONTANA, BEAVERHEAD COUNTY: U.S.A., WASHINGTON, MASON COUNTY: Beaverhead-Deerlodge National Forest. Circa 2 Olympic National Forest. Cushman Lake Road. miles south of Steel Creek Campground on Elev. 241 m. Forest Road 2420 near South Fork Steel Creek. 47.49646° N, 123.30341° W; WGS 84, Elev. 2133 m. uncertainty: 200 m., Source: GPS 45.57967° N, 113.34314° W; Source: GPS Disturbed south-facing slope with loose, gravelly White flowers. Phenology: Flowers. Origin: soil. Hypericum perforatum and Hypochaeris Native. radicata common. Scattered throughout; heavily armed; branches glaucous. Phenology: Fruits. Jared Grummer 2014-29 Origin: Native. 12 Jul 2014 David Giblin 5542 WTU-414636, WTU-V-031679 26 May 2015 WTU-417791, WTU-V-038983 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., MONTANA, BEAVERHEAD COUNTY: Rubus ursinus Cham. & Schltdl. Beaverhead-Deerlodge National Forest. Circa 2 miles south of Steel Creek Campground on U.S.A., WASHINGTON, WHATCOM COUNTY: Forest Road 2420 near South Fork Steel Creek. Above Clayton Bay, circa 9 miles south of Elev. 2133 m. Bellingham. 45.57967° N, 113.34314° W; Source: GPS 48.653446°, -122.482956°; WGS 84, White flowers. Phenology: Flowers. Origin: uncertainty: 1000 m., Source: Digital Map, Native. Georef'd by Ibro Polovina In open area. Flower white on long creeping Jared Grummer 2014-34 stem circa 1.5 meters long. Phenology: Flowers. Origin: Native. 12 Jul 2014 Carl A. Linde Jr. s.n. WTU-414880, WTU-V-032290 7 May 1957 WTU-390464, WTU-V-010477 Rosaceae Rubus spectabilis Pursh Rosaceae U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Rubus ursinus Cham. & Schltdl. North Bay Natural Area Preserve. Elev. 0 ft. U.S.A., WASHINGTON, KING COUNTY: 47° 2' 56.37" N, 124° 4' 11.39" W Millikens yard. Silty clay loam. With Salix sp., Myrica gale, Open yard, loam soil. Phenology: Flowers. Rubus ursinus, Lysichiton americanum, Origin: Native. Gaultheria shallon, Polystichum munitum. Edge of bog. Common. Phenology: Flowers. Origin: Jerry Sheldon 38 Native. 18 May 1973 Regina Johnson 2013-019 WTU-391216, WTU-V-010486 20 Apr 2013 WTU-396310, WTU-V-010104 Rosaceae Rosaceae Rubus nivalis Douglas ex Hook. Rubus pedatus Sm. U.S.A., WASHINGTON, SKAGIT COUNTY: U.S.A., WASHINGTON, SKAMANIA COUNTY: A softly rounded ridge, 0.1 air miles northeast of Northwest side of Mount Adams. Bathtub Meadow. the eastern shore of Lizard Lake on Blanchard Elev. 1306 m. Mountain in the Washington Department of 46° 16.475' N, 121° 36.253' W; Source: Field notes Natural Resources, NRCA designated area. On more or less open ground under Alnus sinuata and Menziesia Elev. 1965 ft. ferruginea in an open Picea engelmannii - Abies amabilis forest 48.63294° N, 122.40374° W; WGS 84 above the meadow. Uncommon; in fruit. Phenology: Fruits. Origin: Second growth forest understory, with Berberis Native. nervosa, Alnus rubra, Tsuga heterophylla, Pseudotsuga menziesii, Vaccinium parvifolium. David Biek 04-182 17 Jul 2004 Leaf material removed for RADseq analysis by with Susan McDougall Megan Ruffley at WTU, February 2016. Phenology: Vegetative. Origin: Native. Herbarium: WTU Bud Hardwick 14-92 Rosaceae 8 Dec 2014 Rubus laciniatus Willd. WTU-404826, WTU-V-013163 U.S.A., WASHINGTON, YAKIMA COUNTY: South side of Mount Adams. Morrison Creek trailhead. Elev. 1390 m. 46° 10.107' N, 121° 29.398' W; Source: Field notes Rosaceae Open area of disturbed soil at the trailhead parking area; surrounded by Abies grandis and Pinus contorta but sunny. Phenology: Rubus spectabilis Pursh Vegetative. Origin: Introduced. U.S.A., WASHINGTON, SKAMANIA COUNTY: David Biek 04-331 19 Aug 2004 North side of Mount Adams. Midway Meadow. with Susan McDougall Elev. 1280 m. 46.353149°, -121.533784°; WGS 84, uncertainty: 250 m., Source: Herbarium: WTU Digital map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Rosaceae David Biek 04-61 18 Jun 2004 with Susan McDougall Rubus laciniatus Willd. Herbarium: WTU U.S.A., WASHINGTON, MASON COUNTY: Bog peatland approximately 1 kilometer south of Dewatto Bay Road. Rosaceae Elev. 120 m. 47.4246° N, 123.03613° W; WGS 84, Rubus leucodermis Douglas ex Torr. & A. Gray uncertainty: 200 m., Source: GPS Mix of Kalmia microphylla/Rhododendron U.S.A., WASHINGTON, YAKIMA COUNTY: groenlandicum/Sphagnum bog and more open Southeast side of Mount Adams. Road up Snipes Mountain. peatland with muddy to deep water channels; Elev. 1280 m. Juncus acuminatus and Rhynchospora alba 46° 6.119' N, 121° 28.076' W; Source: Field notes common. Uncommon in middle of bog; Roadside and on slopes above and below road, west exposure, 20° apparently bird-dispersed. Phenology: Fruits. slope, loose stony soil. With Collomia grandiflora, Penstemon Origin: Introduced. attenuatus, grasses, Achillea millefolium. Phenology: Flowers. Origin: Native. Jeff Walker 815 with Peter Zika, David Giblin David Biek 04-89 3 Jul 2004 with Susan McDougall 24 Aug 2015 Herbarium: WTU WTU-417830, WTU-V-038760

Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus pensilvanicus Poir. U.S.A., WASHINGTON, SKAMANIA COUNTY: Northwest side of Mount Adams. Bathtub Meadow. U.S.A., WASHINGTON, WHATCOM COUNTY: Elev. 1306 m. North of East Badger Road (State Route 546). 46° 16.475' N, 121° 36.253' W; Source: Field notes 48° 57.93' N, 122° 21.52' W On more or less open ground under Alnus sinuata and Menziesia Tree and shrub area between agricultural fields. ferruginea in an open Picea engelmannii - Abies amabilis forest Immature fruits. Phenology: Fruits. Origin: above the meadow. Common. Phenology: Flowers. Origin: Native. Introduced. David Biek 04-181 17 Jul 2004 Laurel Baldwin s.n. with Susan McDougall 27 May 2015 Herbarium: WTU WTU-415253, WTU-V-032574 Rosaceae Rosaceae Rubus bifrons Vest Rubus nivalis Douglas ex Hook. U.S.A., WASHINGTON, FRANKLIN COUNTY: U.S.A., WASHINGTON, JEFFERSON Near bank of swim area at Lyons Ferry State COUNTY: Park at confluence of the Palouse and Snake Along Duckabush River Trail on west slope of Rivers. The Little Hump, ca 1.6 air km west of trailhead. 46.59529° N, 118.21853° W; Source: Elev. 248 m. Specimen Label 47.683° N, 123.063005° W; WGS 84, Growing with Elaeagnus angustifolia, Amorpha uncertainty: 20 m., Source: GPS fruticosa, Echinocystis lobata. In fruit. Shaded Pseudotsuga menziesii forest, with Phenology: Fruits. Origin: Introduced. Gaultheria shallon, Linnaea borealis, Polystichum munitum, and Hylocomnium Jeff Walker 825 splendens. Petals white, nearly linear, ca. 1-1.5 mm wide; leaves deep glossy-green; plants 8 Sep 2015 trailing, locally common on cut trail bank. Photographed. Phenology: Flowers. WTU-404655, WTU-V-013001 Ben Legler 14054 7 May 2016 Rosaceae WTU-409992, WTU-V-023945 Rubus idaeus L. U.S.A., IDAHO, VALLEY COUNTY: Rosaceae On north face of the first peak north of where Forest Road 375 crosses Monumental Summit. Rubus spectabilis Pursh Elev. 2700 m. 44.908553° N, 115.267077° W; WGS 84, U.S.A., WASHINGTON, CLALLAM COUNTY: uncertainty: 8 m., Source: GPS Near Ozette Lake. Steep, north-facing slope with broken rock just 48.14585° N, 124.5927° W; Source: GPS below subalpine summit; with Arnica, Juncus, Forested wetland in managed timberland with Penstemon montanus, Saxifraga bronchialis. Lysichiton americanus understory, Thuja Small shrubs among large rocks and in rock plicata-Tsuga heterophylla-Picea sitchensis crevices. Phenology: Flowers. Origin: Native. overstory, and a lot of decayed large woody debris as substrate. Small pools of shallow S. Legler 2015-23 water present. Abundance: Common. Growth with Ben Legler, Russ Holmes, Don Estberg Habit: vegetative shrub. Site ID: NWCA16-4861. Visit #1. Collection #Q2 (QA Voucher). 10 Jul 2015 Phenology: Vegetative. Origin: Native. WTU-415618, WTU-V-033479 Patricia Johnson NWCA16-4861-V1-Q2 28 Jun 2016 Rosaceae WTU-411089, WTU-V-025525 Rubus parviflorus Nutt. U.S.A., IDAHO, VALLEY COUNTY: Rosaceae Along Forest Service Road 674 on west bank of South Fork Salmon River just south of junction Rubus spectabilis Pursh with East Fork Salmon River. Elev. 1135 m. U.S.A., WASHINGTON, MASON COUNTY: 45.003155° N, 115.718227° W; WGS 84, Near Matlock. uncertainty: 19 m., Source: GPS 47.12901° N, 123.44805° W; Source: GPS West-facing rock road cut among burned conifer Forested edge of beaver-impounded wetland. forest, with Chamerion angustifolium, Sedum Alnus rubra, Frangula purshiana overstory. stenopetalum, Heuchera cylindrica, shrubs. Athyrium filix-femina, Polystichum munitum, Shrubs circa 1 meter tall; common at base of Carex obnupta and C. exsiccata understory. rock. Phenology: Flowers and Fruits. Origin: Abundance: Dominant. Growth Habit: Fruiting Native. shrub. Site ID: NWCA16-4891. Visit #1. Collection #Q4 (QA Voucher). Phenology: S. Legler 2015-28 Fruits. Origin: Native. with Ben Legler, John Bassett, Veanna Willard, Mike Carlson Patricia Johnson NWCA16-4891-V1-Q4 11 Jul 2015 25 May 2016 WTU-415495, WTU-V-033759 WTU-410811, WTU-V-024861 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, MASON COUNTY: Residence of Richard G. Olmstead, 19226 40th Olympic National Forest off of FS-210. Place Northeast, Lake Forest Park. 47.39296° N, 123.2862° W; Source: GPS Elev. 100 m. Forested slope wetland with Acer circinatum, 47.76949° N, 122.2849° W Frangula purshiana, Rubus spectabilis Garden. Phenology: Vegetative. Origin: Native. overstory; Athyrium filix-femina understory. Cultivated. Growing on Thuja plicata stump. Abundance: Sparse. Growth Habit: Vining/climbing up Richard G. Olmstead 2013-49 stump, non-flowering. Site ID: NWCA16-4887. Visit #1. Collection #Q4 (QA Voucher). 18 Aug 2013 Phenology: Vegetative. Origin: Native. WTU-410295, WTU-V-024273 Patricia Johnson NWCA16-4887-V1-Q4 12 May 2016 Rosaceae WTU-411368, WTU-V-025762 Rubus parviflorus Nutt. U.S.A., WASHINGTON COUNTY: Rosaceae C-4 Road. [possibly Stevens County] 48.400756°, -117.874671°; WGS 84, Rubus bifrons Vest uncertainty: 80000 m., Source: Digital map, Georef'd by Joseph Stormer U.S.A., WASHINGTON, KLICKITAT COUNTY: Forest edges and thickets. Phenology: Flowers. Between Klickitat River and Klickitat Rail Trail near Pitt. unknown s.n. 45.79469° N, 121.19924° W; Source: GPS Riparian bench. Populus balsamifera and 24 May 2004 Crataegus douglasii dominant overstory with Equisetum hyemale and Phalaris arundinacea in WTU-405603, WTU-V-016935 understory. Abundance: Dominant. Growth Habit: Fruiting perennial vine. Site ID: NWCA16-4890. Visit #1. Collection #Q1 (QA Voucher). Phenology: Fruits. Origin: Introduced. Rosaceae Patricia Johnson NWCA16-4890-V1-Q1 Rubus ursinus Cham. & Schltdl. 23 Aug 2016 U.S.A., WASHINGTON, SKAGIT COUNTY: Near Anacortes on refinery property. WTU-411381, WTU-V-025749 48.46472° N, 122.54925° W; Source: GPS Forested wetland. With Populus balsamifera, P. tremuloides, Alnus rubra, Symphoricarpos albus, Rubus spectabilis, Ranunculus repens. Rosaceae Abundance: Common. Growth Habit: Creeping along the substrate, rhizomatous, perennial, Rubus phoenicolasius Maxim. woody vine with white 5-petaled flowers. Site ID: NWCA16-4858. Visit #1. Collection #Q3 (QA U.S.A., MASSACHUSETTS, HAMPSHIRE Voucher). Phenology: Flowers. Origin: Native. COUNTY: Pelham Road, Amherst. Patricia Johnson NWCA16-4858-V1-Q3 Escaped from cultivation. Reference specimen for illustration in Revision of Flora of the Pacific 3 May 2016 Northwest WTU Herbarium, 2016. Origin: Introduced. WTU-411362, WTU-V-025768 R. E. Torrey s.n. with R. Livingston 28 Jul 1951 WTU-138654, WTU-V-024777 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., WASHINGTON, STEVENS COUNTY: U.S.A., IDAHO, VALLEY COUNTY: Monumental Mountain. On lower part of slope circa 300 meters 48.490455°, -118.064988°; WGS 84, southwest of south end of Rainbow Lake. uncertainty: 1807 m., Source: GeoLocate, Elev. 2376 m. Georef'd by Maria Yousoufian 44.827944° N, 115.58035° W; WGS 84, Open forests, roadsides, seepage areas. [Mixed uncertainty: 4 m., Source: GPS collection. Also on sheet: Oplopanax horridus, Steep, east-facing granite rock faces and blocky WTU-V-016898. Filed under Oplopanax talus vegetated with Arnica longifolia, Valeriana horridus.] Phenology: Vegetative. Origin: Native. sitchensis, Castilleja miniata, Veratrum viride, and Sibbaldia procumbens. Low shrubs in dry Sue Winterowd s.n. rocky areas. Phenology: Flowers. Origin: Native. with Cindy Jennings Don Estberg 2015-61 26 Aug 2008 with John Bassett, Sara Legler, Ben Legler WTU-405642B, WTU-V-026277 12 Jul 2015 WTU-415623, WTU-V-033483 Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rosaceae U.S.A., IDAHO, VALLEY COUNTY: Rubus parviflorus Nutt. Along Forest Service Road 674 on west bank of South Fork Salmon River just south of junction U.S.A., WASHINGTON, PIERCE COUNTY: with East Fork Salmon River. Joint Base Lewis-McChord, TA18, Road to Elev. 1135 m. Nisqually River. 45.003155° N, 115.718227° W; WGS 84, 46.97675° N, 122.59113° W; WGS 84, uncertainty: 19 m., Source: GPS Source: GPS West-facing rock road cut among burned conifer In opening along roadside in Pseudotsuga forest, with Chamerion angustifolium, Sedum menziesii-Acer macrophyllum forest. stenopetalum, Heuchera cylindrica, shrubs. Occasional. Flowers white. Phenology: Flowers. Arching shrubs; stems glaucous; occasional. Origin: Native. Phenology: Flowers and Fruits. Origin: Native. Peter Dunwiddie 16-151 S. Legler 2015-29 with Sarah Krock, Rod Gilbert with Ben Legler, John Bassett, Veanna Willard, Mike Carlson 2 Jun 2016 11 Jul 2015 WTU-417426, WTU-V-037880 WTU-415630, WTU-V-033492 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus parviflorus Nutt. U.S.A., IDAHO, SHOSHONE COUNTY: Coeur d'Alene National Forest. Shoulder of U.S.A., IDAHO, VALLEY COUNTY: Forest Service Road 945. Payette National Forest. Forest Road 361 Elev. 1075 m. approximately 3 kilometers northeast from 47.782° N, 115.932743° W; WGS 84, junction with Forest Road 48. uncertainty: 4 m., Source: GPS Elev. 1312 m. Rocky road cut among mixed montane conifer 45.04877° N, 115.744° W; WGS 84, forest, with Hypericum perforatum, Holodiscus uncertainty: 30 m., Source: GPS discolor, Ceanothus velutinus. Frequent shrubs Southwest facing slope with Pinus ponderosa to over 1 meter tall. Phenology: Flowers and and Carex rossii. Locally common on edge of Fruits. Origin: Native. road adjacent to draw; berries red. Phenology: Flowers and Fruits. Origin: Native. Julia Bent 2016-02 Audrey Ragsac 2015-38 8 Jul 2016 with Veanna Willard, Mike Carlson, David Giblin WTU-419194, WTU-V-041093 12 Jul 2015 WTU-415625, WTU-V-033481 Rosaceae Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., IDAHO, SHOSHONE COUNTY: U.S.A., IDAHO, SHOSHONE COUNTY: Coeur d'Alene National Forest. Area around Character Peak Area #3. [Elevation bridge over Little North Fork Coeur d'Alene approximate, from digital map.] River. Elev. 4400 ft. Elev. 890 m. 47.63349° N, 116.04842° W; WGS 84, 47.88454° N, 116.13179° W; WGS 84, Source: GPS, Digital map uncertainty: 30 m., Source: Digital map Rocky east-northeast slope. Boulder field. With Disturbed roadside edge and bouldery area Pteridium aquilinum, Acer glabrum var. along banks of river. Uncommon along edge of douglasii, Philadelphus, Lycopodium, river; fruits red. Phenology: Fruits. Origin: Calamagrostis. Phenology: Flowers and Fruits. Native. Origin: Native. David Giblin 5892 Terri Knoke 2016-58 with Richard Robohm 9 Jul 2016 10 Jul 2016 WTU-419430, WTU-V-041008 WTU-420514, WTU-V-047671 Rosaceae Rubus parviflorus Nutt. Rosaceae U.S.A., IDAHO, SHOSHONE COUNTY: Rubus parviflorus Nutt. Coeur d'Alene National Forest. Shadow Falls, along and above trail at Yellow Dog Creek. U.S.A., IDAHO, SHOSHONE COUNTY: [Elevation approximate, from digital map.] Coeur d'Alene National Forest. Forest Road Elev. 3200 ft. 602. Approximately 2 kilometers south of 47.7596° N, 116.1049° W; Source: GPS, trailhead for Mount Sentinel. Digital map Elev. 5243 ft. West-facing slope along and above trail. 47.7964° N, 116.0121° W; WGS 84, Western hemlock dominant, ericaceous uncertainty: 50 m., Source: GPS understory. (Vaccinium, etc.) Phenology: Locally common on edge of road; annual; Flowers. Origin: Native. corollas pink. Locally common and abundant along edge of road. Phenology: Flowers. Origin: Ashley Powell 2016-44 Native. 9 Jul 2016 Shannon Kimball 2016-7 WTU-420964, WTU-V-50094 8 Jul 2016 WTU-420566, WTU-V-048132 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus nivalis Douglas ex Hook. U.S.A., IDAHO, SHOSHONE COUNTY: Coeur d'Alene National Forest. Hairpin bend in U.S.A., WASHINGTON, JEFFERSON Devil's Elbow Road to area approximately 2 COUNTY: kilometers south along road. Olympic National Forest, Duckabush River Trail, Elev. 1245 m. west side of Big Hump, before 5 Mile Camp. 47.7345° N, 116.0332° W; WGS 84, Elev. 1200 ft. uncertainty: 1000 m., Source: Digital map 47° 41.549' N, 123° 7.068' W; Source: Forest edge, disturbed roadside; surrounding Specimen Label forest mixed coniferous. Uncommon; prickly, Sandy loam. With Thuja plicata, Tsuga arching shrub to 1.5 meters tall. Phenology: heterophylla, Pteridium aquilinum, Viola Fruits. Origin: Native. sempervirens, Tiarella trifoliata, Rubus pedatus, Linnaea borealis, Polystichum munitum. Fruit Clayton Wiley 2016-48 red. Common. Phenology: Fruits. Origin: Native. 10 Jul 2016 Regina Johnson 2013-125 WTU-420792, WTU-V-049137 24 Aug 2013 WTU-409149, WTU-V-023066 Rosaceae Rosaceae Rubus idaeus L. ssp. strigosus (Michx.) Focke Rubus bifrons Vest U.S.A., IDAHO, SHOSHONE COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Coeur d'Alene National Forest. Junction of West Cascades. Forest Service Roads 6310 and 3099. Elev. 250 ft. Elev. 2900 ft. T19N R2E S5 47° 52.762' N, 116° 12.495' W; WGS 84, Serious pest. Origin: Introduced. uncertainty: 200 m., Source: GPS Riparian corridor and adjacent west-facing rocky Mengert 4 West outcropping with Heuchera glabra; riparian corridor with Salix sp. and Phalaris arundinacea. 1 May 1977 Uncommon at base of rocky outcropping. Phenology: Fruits. Origin: Native. WTU-405737, WTU-V-016783 Rene Dufour 2016-4 9 Jul 2016 Rosaceae WTU-420670, WTU-V-048675 Rubus spectabilis Pursh U.S.A., WASHINGTON, KING COUNTY: Rosaceae 2 miles north on Tinkham Road. Elev. 2000 ft. Rubus parviflorus Nutt. 47.40383°, -121.533895°; WGS 84, uncertainty: 4250 m., Source: Digital Map, U.S.A., IDAHO, SHOSHONE COUNTY: Georef'd by Joseph Stormer Coeur d'Alene National Forest. Border of Idaho Under the powerline right-of-way. Moist woods, and Montana (Sanders County). Forest Road lowland to mid mountain. Origin: Native. 430, 0.4 miles north of Jordan Summit. Elev. 1552 m. Takekawa 24 West 47.94377° N, 116.00731° W; Source: GPS with Mengert Flowers white. Phenology: Flowers. Origin: Native. 6 May 1977 Richard G. Olmstead 2016-64 WTU-405739, WTU-V-016785 with Sheila Olmstead, Evan Kelly, Veanna Willard, Mike Carlson, Wendy McClure, Regina Johnson, Jim Duemmel, Richard Robohm, Terri Knoke Rosaceae 8 Jul 2016 Rubus idaeus L. ssp. strigosus (Michx.) Focke WTU-420796, WTU-V-049208 U.S.A., IDAHO, VALLEY COUNTY: Payette National Forest. Profile Creek Road. Approximately 3 kilometers from intersection with Stibnite Road. Rosaceae Elev. 1718 m. 44.97774° N, 115.41935° W; WGS 84, Rubus ursinus Cham. & Schltdl. uncertainty: 10 m., Source: GPS East-facing rocky slope with loose soil. Locally U.S.A., WASHINGTON, PIERCE COUNTY: common on dry slope; stems glaucous. No locality given. Phenology: Flowers. Origin: Native. Elev. 250 ft. T19N R2E S5; WGS 84, uncertainty: 62000 Don Estberg 2015-37 m., Source: Digital Map, Georef'd by Ibro with Russ Holmes, David Giblin Polovina The only native blackberry, highly palatable as it 11 Jul 2015 starts to leaf out. Phenology: Flowers. Origin: Native. WTU-416250, WTU-V-034965 Mengert 3 West 1 May 1977 WTU-405721, WTU-V-016767 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus spectabilis Pursh U.S.A., WASHINGTON, GRAYS HARBOR U.S.A., WASHINGTON, WHATCOM COUNTY: COUNTY: Marine Drive near airport, 1.8 air kilometers east Wynoochee Falls. of mouth of Nooksack River, Bellingham. Elev. 1000 ft. Elev. 20 m. 47° 28.278' N, 123° 31.569' W; Source: 48.78° N, 122.54° W; Source: Specimen Specimen Label Label Clay loam. With Rubus spectabilis, Ribes Partly shaded understory thickets, Acer bracteosum, Acer circinatum, Tsuga macrophyllum, Holodiscus, Oemleria, Rubus heterophylla, Dicentra formosa, Athyrium armeniacus. Flowers purple-pink, shrubs 2.5 filix-femina. Riparian area in Tsuga heterophylla meters tall. Phenology: Flowers. Origin: Native. forest. Common. Phenology: Fruits. Origin: Native. Peter F. Zika 27632 Regina Johnson 2013-079 9 Apr 2016 4 Jul 2013 WTU-410449, WTU-V-024486 WTU-409652, WTU-V-023660 Rosaceae Rosaceae Rubus bifrons Vest Rubus laciniatus Willd. U.S.A., OREGON, KLAMATH COUNTY: U.S.A., WASHINGTON, STEVENS COUNTY: Highway 97, circa 9 air miles north of Klamath Off Valley Westside Road. Falls, 2.4 air miles south of Hagelstein Park. 48.175173°, -117.724685°; WGS 84, Elev. 4150 ft. uncertainty: 500 m., Source: GeoLocate, 42.34909° N, 121.82759° W; Source: Georef'd by Sara Shiina Specimen Label Waste areas and crop land. Phenology: Bank of seasonally wet ditch on gravelly Flowers. Origin: Introduced. highway and railroad fill. Associated taxa: Ericameria nauseosa, Penstemon deusus, CKJ s.n. Bromus tectorum, Sambucus mexicana. with SLW Phenology: Flowers. 25 Sep 2006 R.E. Brainerd 2384 WTU-405674, WTU-V-016872 19 Jun 2015 WTU-412921, WTU-V-028429 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus nivalis Douglas ex Hook. U.S.A., WASHINGTON, STEVENS COUNTY: Off Valley Westside Road. U.S.A., OREGON, DOUGLAS COUNTY: 48.175173°, -117.724685°; WGS 84, Above Ragsdale Butte Road, 2.7 air kilometers uncertainty: 500 m., Source: GeoLocate, northwest of Pilot Rock. Georef'd by Sara Shiina Elev. 1235 m. Waste areas and crop land. Origin: Introduced. 42.782° N, 122.839° W; Source: Specimen Label CKJ s.n. Steep partly shaded mesic north-facing slope, with SLW with Tsuga heterophylla, Pseudotsuga, Acer macrophyllum, Trientalis, Adenocaulon, Luzula 25 Sep 2006 comosa, Hieracium albiflorum. Prostrate, shiny leaves, uncommon. Origin: Native. WTU-405673, WTU-V-016873 Peter F. Zika 27918 17 Jun 2016 WTU-410565, WTU-V-024676 Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus arcticus L. ? U.S.A., WASHINGTON, KING COUNTY: U.S.A., ALASKA COUNTY: Near southeast Middle Fork Road, North Bend, Kotzebue. Middle Fork Snoqualmie River drainage. 66.877849°, -162.453845°; WGS 84, Elev. 240 m. uncertainty: 10000 m., Source: Digital Map, 47.476° N, 121.702° W; Source: Specimen Georef'd by Joseph Stormer Label Phenology: Flowers. Origin: Native. Shady forest edge along old woods road, with Polystichum munitum, Trientalis, Rubus ursinus, Clayton Haberman s.n. Pteridium, Gaultheria shallon, Thuja, Tsuga, Pseudotsuga. Prostrate, immature fruit red; 8 Jul 1990 common. Phenology: Fruits. Origin: Native. WTU-407287, WTU-V-020998 Peter F. Zika 27863 6 Jun 2016 Rosaceae WTU-410561, WTU-V-024680 Rubus arcticus L. Rosaceae U.S.A., WASHINGTON, OKANOGAN COUNTY: [locality and coordinates withheld; status in Rubus parviflorus Nutt. Washington: G5T5/S1 (Threatened; WANHP)] Elev. 5377 ft. U.S.A., WASHINGTON, KING COUNTY: Slope 0%, aspect 90%. Peat, high elevation Near southeast Middle Fork Road, North Bend, wetland complex, this area with slow moving Middle Fork Snoqualmie River drainage. water, hummocks of Carex and some Elev. 240 m. Sphagnum hummocks. Associated plants: Picea 47.476° N, 121.702° W; Source: Specimen engelmannii, Pinus contorta, Carex scopulorum, Label Salix spp. Flower not opened yet but diagnostic Shady forest edge along old woods road, with pink color showing. Phenology: Flowers. Origin: Polystichum munitum, Trientalis, Rubus ursinus, Native. Pteridium, Gaultheria shallon, Thuja, Tsuga, Pseudotsuga. 1 meter tall shrub, white flowers Erica Heinlen 2017-9 46 millimeters diameter; common. Phenology: Flowers. Origin: Native. 19 Jun 2017 Peter F. Zika 27865 WTU-417219, WTU-V-036851 6 Jun 2016 WTU-410558, WTU-V-024681 Rosaceae Rubus pensilvanicus Poir. Rosaceae U.S.A., WASHINGTON, SNOHOMISH Rubus idaeus L. ssp. idaeus COUNTY: Gated forest road intersections of Kellogg Lake U.S.A., WASHINGTON, PEND OREILLE Road and Sierra Pacific Industries 24000 road COUNTY: and 25000 road. Plants growing 30 meters to Fawn Ridge above Trimble Creek. the east from the midpoint of that turnout, north 48.3667°, -117.3667°; WGS 84, uncertainty: of the paved roadway, about 3 meters north of 1500 m., Source: GeoLocate, Georef'd by the paved surface. Sara Shiina 47° 53.881' N, 121° 44.963' W; WGS 84, Phenology: Flowers. Origin: Native. Source: GPS [Geocoordinates provided by collector added Cindy Talbott 453 after label was printed}. Phenology: Fruits. Origin: Introduced. 25 Jun 1979 Noah Booker s.n. WTU-410689, WTU-V-024816 28 Jul 2017 WTU-416658, WTU-V-036264 Rosaceae Rosaceae Rubus laciniatus Willd. Rubus parviflorus Nutt. U.S.A., WASHINGTON, KLICKITAT COUNTY: U.S.A., WASHINGTON, STEVENS COUNTY: At Dallesport Treaty Indian Fishing Site along Western Colville National Forest and Vicinity: Columbia River. Black Canyon ridgetop, ca [about] 6 air mi ESE 45.619264°, -121.16441°; WGS 84, of Northport. uncertainty: 5000 m., Source: Digital Map, Elev. 4000-4680 ft. Georef'd by Dan Post T39N R41E S8 E½, also S17 NW¼; WGS 84, In rocky fill slope of railroad. Phenology: uncertainty: 3657 m., Source: GeoLocate, Flowers. Georef'd by Dan Post Cool, moist subalpine forest; Abies lasiocarpa, David Gross 1225 hemlock, cedar; also roadside. Phenology: Flowers. 25 May 2017 Jean Wood 5146 WTU-415918, WTU-V-034204 25 Jun 1996 Rosaceae WTU-416046, WTU-V-034586 Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, KING COUNTY: Rosaceae East half of South Woods Park; south Shoreline. Elev. 114 m. Rubus parviflorus Nutt. 47.738154° N, 122.303805° W; WGS 84, uncertainty: 12 m., Source: GPS, Georef'd by Ben Legler U.S.A., WASHINGTON, STEVENS COUNTY: Small opening in mature Pseudotsuga menziesii woods, with Rubus Colville National Forest, Twin Lakes bifrons, R. ursinus, R. leucodermis, Photinia davidiana, Polystichum T35N R41E S4; WGS 84, uncertainty: 500 m., munitum. Stems glaucous, arching, 1.5-2 meters tall, in clumps; Source: Digital Map, Georef'd by Dan Post leaves white-hairy beneath; primocane leaves palmately compound; Phenology: Flowers. floricane leaves ternately compound; inflorescence 3-5-flowered; plants locally common in opening. Phenology: Fruits. Ellman [s.n.] Ben Legler 14596 17 Sep 2017 Jun 1979 Herbarium: WTU WTU-416053, WTU-V-034589

Rosaceae Rubus idaeus L. ssp. idaeus Rosaceae U.S.A., WASHINGTON, KING COUNTY: Rubus pedatus Sm. South Woods Park: in center of east half of park; south Shoreline. Elev. 115 m. U.S.A., WASHINGTON, MASON COUNTY: 47.738101° N, 122.304011° W; WGS 84, uncertainty: 11 m., Olympic National Forest, 4 Stream Road. Source: GPS, Georef'd by Ben Legler Elev. 2200 ft. Small opening in mature Pseudotsuga menziesii forest created from 47.51838° N, 123.36403° W; Source: old disturbance, with Rubus spp., Cotoneaster spp., Photinia Specimen Label davidiana, small Thuja plicata, and a few old, persisting, ornamental Rocky sandy loam With Pseudotsuga menziesii, trees. Canes arching, to ca 1.5 meters tall, completely unarmed, Tsuga heterophylla, Chamerion angustifolium, glaucous; leaves white-tomentose beneath; primocanes with most Digitalis purpurea, Aquilegia formosa. Forest leaves pinnately 5-foliate; floricanes senesced, with leaves ternate Service road in coniferous forest. Common. and fruits in small clusters; single patch of ca 15 stems growing Phenology: Flowers. Origin: Native. among R. leucodermis, either adventive or persisting here for several decades, now appearing wild. Phenology: Fertile. Regina Johnson 2016-023 Ben Legler 14613 31 Oct 2017 3 Jun 2016 Herbarium: WTU WTU-414488, WTU-V-031538 Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus idaeus L. ssp. strigosus (Michx.) Focke U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, STEVENS COUNTY: Stevens Pass. Stevens Pass ski area. Western Colville National Forest and Vicinity: Elev. 4300 ft. Little Round Top Mountain: Middle Fork Mill 47° 44.527' N, 121° 5.164' W; T26N R13E Creek, circa 9 road miles northeast of Colville sec. 13; Source: Specimen Label on Aladdin Route. Meadows with seeps and springs surrounded by Elev. 2300-3300 ft. subalpine fir and douglas fir forest. Assoc. with T36N R40E S15 NW1/4; also S16 NE1/4; Spiraea douglasia, Alnus, Vaccinium, Valerian WGS 84, uncertainty: 1000 m., Source: Digital sitchensis. Phenology: Flowers. Map, Georef'd by Sara Shiina Cedar at creekside; mixed Douglas fir forest, P. Camp 3720 and aspen groves on lower slopes; Douglas fir, ponderosa pine and larch with pinegrass and 11 Jul 2017 ninebark on upper slopes; rock outcrops of shale/slate. Phenology: Flowers and Fruits. WTU-415754, WTU-V-033414 Origin: Native. Jean Wood 901 5 Jun 1995 Rosaceae WTU-410688, WTU-V-024999 Rubus pubescens Raf. Rosaceae U.S.A., WASHINGTON, STEVENS COUNTY: Western Colville National Forest and Vicinity: Rubus idaeus L. ssp. strigosus (Michx.) Focke Little Twin Lakes, circa 11 air miles east of Colville. U.S.A., WASHINGTON, FERRY COUNTY: Elev. 3700 ft. Western Colville National Forest and Vicinity: T35N R41E S4 NW1/4; also T36N R41E S33 Lambert Creek, circa 9 air miles northeast of SW1/4; WGS 84, uncertainty: 1000 m., Republic. Source: Digital Map, Georef'd by Sara Shiina Elev. 3800-4700 ft. Mixed forest surrounding lake; Pinus contorta T37N R34E S8 NE1/4 and S1/2; WGS 84, and Vaccinium scoparium. Phenology: uncertainty: 4221 m., Source: Digital Map, Vegetative. Origin: Native. Georef'd by Isaac Afolayan Creekside. Phenology: Flowers and Fruits. Jean Wood 6740 Origin: Native. 1 Aug 1996 Jean Wood 6379 WTU-410701, WTU-V-024991 25 Jul 1996 WTU-410687, WTU-V-025000 Rosaceae Rubus pedatus Sm. Rosaceae U.S.A., WASHINGTON, PEND OREILLE Rubus pubescens Raf. COUNTY: Western Colville National Forest and Vicinity: U.S.A., WASHINGTON, STEVENS COUNTY: Granite Meadows, circa 3.5 air miles east of Colville National Forest, Kettle Falls Ranger Little Pend Oreille Lakes, off Tacoma Divide District about 25 miles north of Kettle Falls. Road. From Kettle Falls take Highway #395 north Elev. 4350 ft. about 23 miles to the Sand Creek Road #4013. T36N R42E S14 SW1/4; WGS 84, Head east on it about 2.5 miles to Forest uncertainty: 600 m., Source: Digital Map, Service Road #100, then northeast about 1.25 Georef'd by Joseph Stormer miles to Forest Service Road #115 junction. Go Old beaver ponds with some open water; northwest on Forest Road #120 about 0.5 miles, extensive peat bog with multiple Carex species. then west 0.2 miles to a calcareous stream Phenology: Fruits. Origin: Native. north of the road. Elev. 2360 ft. Jean Wood 2738 T40N R36E S25 NE1/4; WGS 84, uncertainty: 31567 m., Source: GeoLocate, Georef'd by 26 Jul 1995 Crystal Shin Plants were growing with Sanicula marilandica. WTU-410697, WTU-V-024992 Phenology: Flowers. Origin: Native. Kirk Larson 336 28 May 1998 WTU-410698, WTU-V-024990 Rosaceae Rosaceae Rubus pedatus Sm. Rubus idaeus L. U.S.A., WASHINGTON, PEND OREILLE U.S.A., WASHINGTON, FERRY COUNTY: COUNTY: Colville National Forest, Ward Lake about 15 Colville [National Forest], Salmo River. miles north of Republic. From Republic take Elev. 4100 ft. County Road #270 north 4.5 miles, then go T40N R45E [S]11 NESW; WGS 84, north on County Road #203 for 3 miles. Turn uncertainty: 1500 m., Source: Digital Map, west on County Road #201 about 1 mile, then Georef'd by Sara Shiina take County Road #517 north [for] 3.5 miles. Go Associated plants: WH/Pamy. Phenology: north on Forest Service Road #2149 for 0.5 Flowers. Origin: Native. miles, then east on Forest Service Road #2150 for 1 mile. RCP 511 Elev. 2600 ft. T38N R32E S14 SW1/4 SW1/4; WGS 84, 24 Sep 1974 uncertainty: 500 m., Source: Digital Map, Georef'd by Sara Shiina WTU-410696, WTU-V-024994 Associated plants include: Juncus ensifolius, Carex aquatilis, Carex diandra and Carex lasiocarpa. Phenology: Flowers. Origin: Native. Rosaceae Kathy Ahlenslager 594 10 Jul 1999 Rubus parviflorus Nutt. with Jean Wood U.S.A., WASHINGTON, PEND OREILLE WTU-410686, WTU-V-024997 COUNTY: Newport Ranger District, Colville National Rosaceae Forest. Spring Creek Road. T32N R42E S27 SW1/4; WGS 84, Rubus pedatus Sm. uncertainty: 400 m., Source: Digital Map, Georef'd by Joseph Stormer : Roadbank next to Grand fir, Douglas-fir, larch and lodgepole with ninebark. Phenology: Phenology: Flowers. Origin: Native. Flowers. Origin: Native. Thomas Burke 163 Cindy Jo Talbott 444 21 Aug 1974 20 Jun 1979 WTU-410694, WTU-V-024998 WTU-410693, WTU-V-024995

Rosaceae Rubus pedatus Sm. Rosaceae U.S.A., WASHINGTON, FERRY COUNTY: Colville National Forest, Sherman Creek. Rubus vestitus Weihe & Nees Elev. 3547 ft. T36N R35E S15; WGS 84, uncertainty: 1790 U.S.A., WASHINGTON, GRAYS HARBOR m., Source: Digital Map, Georef'd by Isaac COUNTY: Afolayan Margin of Bowerman airfield, Hoquiam, 4.7 air Associated plants: PIEN, Mertensia sp., LIBO-2, kilometers west-northwest of Rennie Island. EQAR, GYDR. Many plants in swampy/wooded Elev. 4 m. area near stream. Phenology: Vegetative. 46.9729° N, 123.938° W; Source: Specimen Origin: Native. Label Uncommon in thickets, full sun, with Salix J. L. Murray s.n. hookeriana, Rubus spectabilis, R. armeniacus. Petals pink, arching shrub 2.5 meters tall. 19 Aug 1980 Phenology: Fruits. Origin: Introduced. WTU-410695, WTU-V-024996 Peter F. Zika 28987 12 Aug 2017 WTU-416902, WTU-V-036561 Rosaceae Rosaceae Rubus vestitus Weihe & Nees Rubus cf. anglocandicans A. Newton U.S.A., OREGON, LANE COUNTY: U.S.A., OREGON, LANE COUNTY: North Shore Boat Ramp on Fall Creek North Shore Boat Ramp on Fall Creek Reservoir. 4.5 kilometers northeast of Lowell. Reservoir. 4.5 kilometers northeast of Lowell. Elev. 265 m. Elev. 265 m. 43° 57' 25.88" N, 122° 45' 18.72" W; Source: 43° 57' 24.46" N, 122° 45' 16.17" W; Source: Specimen Label Specimen Label Near parking lot for boat ramp, open, south Near parking lot for boat ramp, open, south aspect, adjacent to forest of Pseudotsuga, aspect, adjacent to forest of Pseudotsuga, Calocedrus, Populus, Fraxinus, Quercus. Pink Calocedrus, Populus, Fraxinus, Quercus. flowers. Sample collection site of L.V. Clark Sample collection site of L.V. Clark (see (see Biological Invasions vol. 15 p. 1331) where Biological Invasions vol. 15 p. 1331) where this this species was identified with DNA markers. species was identified with DNA markers. Origin: Introduced. Phenology: Fruits. Origin: Introduced. Edward R. Alverson 2016-004 Edward R. Alverson 2016-006 18 Jun 2016 18 Jun 2016 WTU-417241, WTU-V-036880 WTU-417244, WTU-V-036881

Rosaceae Rosaceae Rubus laciniatus Willd. Rubus cf. anglocandicans A. Newton U.S.A., OREGON, LANE COUNTY: U.S.A., OREGON, LANE COUNTY: North Shore Boat Ramp on Fall Creek North Shore Boat Ramp on Fall Creek Reservoir. 4.5 kilometers northeast of Lowell. Reservoir. 4.5 kilometers northeast of Lowell. Elev. 265 m. Elev. 265 m. 43° 57' 24.61" N, 122° 45' 16.25" W; Source: 43° 57' 24.22" N, 122° 45' 11.32" W; Source: Specimen Label Specimen Label Near parking lot for boat ramp, open, south Near parking lot for boat ramp, open, south aspect, adjacent to forest of Pseudotsuga, aspect, adjacent to forest of Pseudotsuga, Calocedrus, Populus, Fraxinus, Quercus. Pink Calocedrus, Populus, Fraxinus, Quercus. White flowers. Sample collection site of L.V. Clark flowers. Sample collection site of L.V. Clark (see Biological Invasions vol. 15 p. 1331) where (see Biological Invasions vol. 15 p. 1331) where this species was identified with DNA markers. this species was identified with DNA markers. Phenology: Flowers. Origin: Introduced. Phenology: Flowers. Origin: Introduced. Edward R. Alverson 2016-003 Edward R. Alverson 2016-002 18 Jun 2016 18 Jun 2016 WTU-417242, WTU-V-036882 WTU-417240, WTU-V-036875

Rosaceae Rosaceae Rubus cf. anglocandicans A. Newton Rubus cf. anglocandicans A. Newton U.S.A., OREGON, LANE COUNTY: U.S.A., WASHINGTON, COWLITZ COUNTY: North Shore Boat Ramp on Fall Creek Between South Kelso Drive and Interstate 5 Reservoir. 4.5 kilometers northeast of Lowell. near Exit 39, 0.24 kilometers south of Allen Elev. 265 m. Street, Kelso. 43° 57' 24.46" N, 122° 45' 16.17" W; Source: Elev. 5 m. Specimen Label 46° 8' 35.21" N, 122° 53' 47.16" W; Source: Near parking lot for boat ramp, open, south Specimen Label aspect, adjacent to forest of Pseudotsuga, Forming dense colony along chain link fence on Calocedrus, Populus, Fraxinus, Quercus. roadside. Sample collection site of L.V. Clark Sample collection site of L.V. Clark (see (see Biological Invasions vol. 15 p. 1331) where Biological Invasions vol. 15 p. 1331) where this this species was identified with DNA markers species was identified with DNA markers. Phenology: Fruits. Origin: Introduced. Origin: Introduced. Edward R. Alverson 2016-014 Edward R. Alverson 2016-006 28 Jul 2016 18 Jun 2016 WTU-417234, WTU-V-036865 WTU-417243, WTU-V-036879 Rosaceae Rosaceae Rubus cf. anglocandicans A. Newton Rubus cf. anglocandicans A. Newton U.S.A., WASHINGTON, COWLITZ COUNTY: U.S.A., WASHINGTON, KING COUNTY: Between South Kelso Drive and Interstate 5 Sea-Tac rest area along Interstate 5 north near Exit 39, 0.24 kilometer south of Allen bound near milepost 141, circa 2 kilometers Street, Kelso. northeast of Milton. Elev. 5 m. Elev. 75 m. 46° 8' 35.21" N, 122° 53' 47.16" W; Source: 47° 16' 12.95" N, 122° 18' 51.58" W; Source: Specimen Label Specimen Label Forming dense colony along chain link fence on Along east edge of rest area adjacent to Milton roadside. Sample collection site of L.V. Clark Road, fairly open but adjacent to Pseudotsuga (see Biological Invasions vol. 15 p. 1331) where menziesii. Sheet 1 of 2. Phenology: Flowers. this species was identified with DNA markers. Origin: Introduced. Sheet 3 of 3. Phenology: Fruits. Origin: Introduced. Edward R. Alverson 2016-015 Edward R. Alverson 2016-014 28 Jul 2016 28 Jul 2016 WTU-417237, WTU-V-036871 WTU-417233, WTU-V-036866 Rosaceae Rosaceae Rubus bifrons Vest Rubus cf. anglocandicans A. Newton U.S.A., OREGON, LANE COUNTY: U.S.A., WASHINGTON, COWLITZ COUNTY: North Shore Boat Ramp on Fall Creek Between South Kelso Drive and Interstate 5 Reservoir, 4.5 kilometers northeast of Lowell. near Exit 39, 0.24 kilometer south of Allen Elev. 265 m. Street, Kelso. 43° 57' 25.61" N, 122° 45' 20.53" W; Source: Elev. 5 m. Specimen Label 46° 8' 35.21" N, 122° 53' 47.16" W; Source: Near parking lot for boat ramp, open, south Specimen Label aspect, adjacent to forest of Pseudotsuga, Forming dense colony along chain link fence on Calocedrus, Populus, Fraxinus, Quercus. roadside. Sample collection site of L.V. Clark Sample collection site of L.V. Clark (see (see Biological Invasions vol. 15 p. 1331) where Biological Invasions vol. 15 p. 1331) where this this species was identified with DNA markers. species was identified with DNA markers. Sheet Sheet 1 of 3. Phenology: Vegetative. Origin: 1 of 2. Phenology: Flowers. Origin: Introduced. Introduced. Edward R. Alverson 2016-005 Edward R. Alverson 2016-014 18 Jun 2016 28 Jul 2016 WTU-417236, WTU-V-036870 WTU-417239, WTU-V-036872

Rosaceae Rosaceae Rubus cf. anglocandicans A. Newton Rubus bifrons Vest U.S.A., WASHINGTON, KING COUNTY: U.S.A., OREGON, LANE COUNTY: Sea-Tac rest area along Interstate 5 north North Shore Boat Ramp on Fall Creek bound near milepost 141, circa 2 kilometers Reservoir, 4.5 kilometers northeast of Lowell. northeast of Milton. Elev. 265 m. Elev. 75 m. 43° 57' 25.61" N, 122° 45' 20.53" W; Source: 47° 16' 12.95" N, 122° 18' 51.58" W; Source: Specimen Label Specimen Label Near parking lot for boat ramp, open, south Along east edge of rest area adjacent to Milton aspect, adjacent to forest of Pseudotsuga, Road, fairly open but adjacent to Pseudotsuga Calocedrus, Populus, Fraxinus, Quercus. menziesii. Sheet 2 of 2. Phenology: Vegetative. Sample collection site of L.V. Clark (see Origin: Introduced. Biological Invasions vol. 15 p. 1331) where this species was identified with DNA markers. Sheet Edward R. Alverson 2016-015 2 of 2. Phenology: Vegetative. Origin: Introduced. 28 Jul 2016 Edward R. Alverson 2016-005 WTU-417238, WTU-V-036874 18 Jun 2016 WTU-417235, WTU-V-036869 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus bifrons Vest U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Shoulder of NE 165th Street ca 150 meters east of 15th Avenue NE, Olympic National Forest. Beaver Lake. on north margin of Hamlin Park; Shoreline. Elev. 182 m. Elev. 113 m. 48.1137° N, 124.24568° W; WGS 84, 47.748407° N, 122.311033° W; WGS 84, uncertainty: 14 m., uncertainty: 20 m., Source: Field notes Source: GPS, Georef'd by Ben Legler Shoreline of Lake dominated by Spiraea Forested road edge under Pseudotsuga menziesii, with Rubus douglasii and Salix sp. and adjacent waters to bifrons and Prunus avium. Primocane leaves mostly palmate with 5 1.5 meters deep. Locally common in disturbed leaflets but a few pinnate-palmate with 7 leaflets; leaflets soil at edge of parking area. Phenology: Fruits. white-tomentose beneath; filaments slightly flattened towards their Origin: Native. base; stem and prickles recurved; stems glaucous, without gland-tipped hairs; small patch of plants on road edge, to 1.5 meters David Giblin 6279 tall. Photographed. Phenology: Flowers. 28 Aug 2018 Ben Legler 14662 20 Jun 2018 WTU-419841, WTU-V-043590 Herbarium: WTU

Rosaceae Rosaceae Rubus lasiococcus A. Gray Rubus leucodermis Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, GRAYS HARBOR U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: COUNTY: Olympic National Forest. Forest Service Road Olympic National Forest, Junction of Forest 2372. Approximately 3 road kilometers north of Service Roads 2270 and 868. North and south Satsop Lakes. From GPS point to 2 kilometers sides of bridge along east bank. up road. Elev. 269 m. Elev. 775 m. 47.44773° 123.55046°; Source: Specimen 47.42763° N, 123.52593° W; WGS 84, Label, Inverted longitude. uncertainty: 1000 m., Source: Specimen Sandy riparian zone dominated by Salix Label scouleri; shrubby understory and cobbly open South-facing slopes, exposed, with scree, rocky areas. Uncommon; shrub to 1 meter tall; outcrops, and road cuts; adjacent Psuedotsuga flowers white. Phenology: Flowers. Origin: menziesii forest. Locally common on edge of Native. forest; sterile. Phenology: Vegetative. Origin: Native. David Giblin 6091 David Giblin 6103 22 May 2018 23 May 2018 WTU-419555, WTU-V-042654 WTU-417995, WTU-V-039878 Rosaceae Rosaceae Rubus bifrons Vest Rubus spectabilis Pursh U.S.A., WASHINGTON, ASOTIN COUNTY: Along edge of gravel parking lot adjacent to U.S.A., WASHINGTON, GRAYS HARBOR Asotin Creek in Asotin City Park. COUNTY: Elev. 780 ft. Olympic National Forest. Forest Service Road 46.3399° N, 117.05621° W; Source: 2372. Approximately 3 road kilometers north of Specimen Label Satsop Lakes. From GPS point to 2 kilometers Growing with Alnus and Bromus. Phenology: up road. Fruits. Origin: Introduced. Elev. 775 m. 47.42763° N, 123.52593° W; WGS 84, Jeff Walker 1091 uncertainty: 1000 m., Source: GPS South-facing slopes, exposed, with scree, rocky 1 Sep 2018 outcrops, and road cuts; adjacent Psuedotsuga menziesii forest. Scattered along forest edge; WTU-418479, WTU-V-039148 corollas purple-red; to 2 meters tall. Phenology: Flowers. Origin: Native. David Giblin 6115 23 May 2018 WTU-417985, WTU-V-039868 Rosaceae Rosaceae Rubus phoenicolasius Maxim. Rubus ursinus Cham. & Schltdl. U.S.A., MASSACHUSETTS, BARNSTABLE U.S.A., WASHINGTON, PIERCE COUNTY: COUNTY: JBLM, TA 10. Edge of old sand quarry, N of Brick Kiln Road, Elev. 115 m. 0.25 air km W of Mares Pond, Falmouth, Cape 47.05692° N, 122.47941° W; WGS 84, Source: GPS Cod. Occasional along roadside in openings, Pseudotsuga menziesii Elev. 10 m. forest. Origin: Native. 41.5874° N, 70.6025° W; Source: Specimen Label Peter Dunwiddie PWD-16-227 13 Jun 2016 Large colony naturalized in partly shaded with Rod Gilbert, Sarah Krock, Peter Zika thickets, with Celastrus orbiculatus, Scrophularia lanceolata. Purple glands on Herbarium: WTU sepals and stems, 1.5-2.5 m tall, fruit orange ripening to shiny red. Phenology: Fruits. Origin: Introduced. Rosaceae Peter F. Zika 28927B Rubus ursinus Cham. & Schltdl. with Pam Polloni, Don Schall U.S.A., WASHINGTON, PIERCE COUNTY: 25 Jul 2017 JBLM, TA-13, Lynch Wetland. Elev. 104 m. WTU-417091, WTU-V-036725 47.01406° N, 122.51003° W; WGS 84, Source: GPS Near vernal pool surrounded by Fraxinus and Quercus garryana. Rosaceae Common trailing on ground in openings along road edge under Pseduotsuga menziesii. Immature berries red. Origin: Native. Rubus laciniatus Willd. Peter Dunwiddie PWD-16-241 13 Jun 2016 U.S.A., WASHINGTON, LEWIS COUNTY: with Rod Gilbert, Sarah Krock, Peter Zika Cowlitz Valley. 46.538271°, -121.852778°; WGS 84, Herbarium: WTU uncertainty: 20000 m., Source: Digital Map, Georef'd by Dan Post Blackberry Rosaceae R. Ernesti E2136 Rubus laciniatus Willd. 12 Jul 1938 U.S.A., WASHINGTON, PIERCE COUNTY: JBLM, TA-10, next to Johnson Marsh. WTU-385777, WTU-V-041399 Elev. 104 m. 47.01406° N, 122.51003° W; WGS 84, Source: GPS Near vernal pool surrounded by Fraxinus and Quercus garryana. Low shrub in openings along road edge under Pseduotsuga Rosaceae menziesii. Immature berries red. Origin: Introduced. Rubus parviflorus Nutt. Peter Dunwiddie PWD-16-242 13 Jun 2016 with Rod Gilbert, Sarah Krock, Peter Zika U.S.A., WASHINGTON, PIERCE COUNTY: JBLM, TA18, Road to Nisqually River. Herbarium: WTU Elev. 65 m. 46.97675° N, 122.59113° W; WGS 84, Source: GPS Occasional in opening along roadside in Pseudotsuga Rosaceae menziesii-Acer macrophyllum forest. Flwrs: White. Origin: Native. Rubus leucodermis Douglas ex Torr. & A. Gray Peter W. Dunwiddie PWD 16-151 2 Jun 2016 with Sarah Krock, Rod Gilbert U.S.A., WASHINGTON, PIERCE COUNTY: JBLM, TA-5. Herbarium: WTU Elev. 105 m. 47.05059° N, 122.57736° W; WGS 84, Source: GPS Along roadside, with Pteridium aquilinum. Stems glaucus, fruit black, Rosaceae dome-shaped. Origin: Introduced. Rubus bifrons Vest Peter Dunwiddie PWD-16-277 24 Jun 2016 with Rod Gilbert, Sarah Krock, Peter Zika U.S.A., WASHINGTON, PIERCE COUNTY: JBLM, TA 10. Herbarium: WTU Elev. 115 m. 47.05692° N, 122.47941° W; WGS 84, Source: GPS to 3 m tall, common along roadside in openings, Pseudotsuga menziesii forest. Origin: Introduced. Peter Dunwiddie PWD-16-226 13 Jun 2016 with Rod Gilbert, Sarah Krock, Peter Zika Herbarium: WTU Rosaceae Rosaceae Rubus spectabilis Pursh Rubus parviflorus Nutt. U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: JBLM, TA-5, roadside wetland. Patos Island along north shore. Elev. 105 m. 48.786094° N, 122.946467° W; WGS 84, Source: Digital Map, 47.05059° N, 122.57736° W; WGS 84, Source: GPS Georef'd by Linnea Johnson Several plants at edge of wetland. Fruits orange. Origin: Native. Local, in most areas. Petals large, white. Phenology: Fruits. Origin: Native. Peter Dunwiddie PWD-16-278 24 Jun 2016 with Rod Gilbert, Sarah Krock, Peter Zika Peter Dunwiddie PWD-18-192 30 May 2018 with Peter Zika, Shelley Evans Herbarium: WTU Herbarium: WTU Rosaceae Rosaceae Rubus laciniatus Willd. Rubus parvifolius L. U.S.A., WASHINGTON, THURSTON COUNTY: Scatter Creek North Unit U.S.A., ILLINOIS, CHAMPAIGN COUNTY: Elev. 200 m. Urbana; Busey Woods; at southwest end of 46.84173° N, 123.00143° W; WGS 84, Source: GPS boardwalk, under powerline right of way. A few plants along trail in Pseudotsuga understory, with Elev. 215 m. Symphoricapos albus, Rubus armeniacus, Stachys cooleyae Petals 40.1274° N, 88.211474° W; WGS 84, white to pink, variously lobed Phenology: Flowers. Origin: uncertainty: 100 m., Source: Specimen Label Introduced. Edge of Trail along powerline right of way bordering wet deciduous woods. Patch 2-4 Peter Dunwiddie PWD-17-493 30 Jun 2017 meets wide; sprawling arching stems, flowers with Etsuko Reistroffer purple. Phenology: Flowers. Origin: Native. Herbarium: WTU Gordon C. Tucker 16357 Rosaceae 24 May 2015 Rubus bifrons Vest WTU-420411, WTU-V-045399 U.S.A., WASHINGTON, THURSTON COUNTY: Scatter Creek North Unit Elev. 200 m. Rosaceae 46.84173° N, 123.00143° W; WGS 84, Source: GPS A few plants along trail in Pseudotsuga understory, with Rubus parviflorus Nutt. Symphoricapos albus, Rubus armeniacus, Stachys cooleyae Petals white Phenology: Flowers. Origin: Introduced. U.S.A., OREGON, UNION COUNTY: Wallowa-Whitman National Forest, Forest Peter Dunwiddie PWD-17-494 30 Jun 2017 Service Road 7312. Approximately 1 km from with Etsuko Reistroffer junction with F. S. Road 4380. Elev. 1567 m. Herbarium: WTU 45.01701° N, 118.15694° W; WGS 84, uncertainty: 50 m., Source: GPS, Invalid coordinates (located outside county bounds). Rosaceae Riparian corridor and adjacent upland forest with Pinus contorta and Pseudotsuga menziesii. Rubus spectabilis Pursh Scattered at forest edge. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, THURSTON COUNTY: Scatter Creek North Unit Wendy McClure 2018-17 Elev. 200 m. 46.84173° N, 123.00143° W; WGS 84, Source: GPS 13 Jul 2018 A few plants along trail in Pseudotsuga understory, with Symphoricapos albus, Rubus armeniacus, Rubus laciniatus, Stachys WTU-421637, WTU-V-053304 cooleyae Berries orange Phenology: Fruits. Origin: Native. Peter Dunwiddie PWD-17-495 30 Jun 2017 with Etsuko Reistroffer Herbarium: WTU Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus ulmifolius Schott U.S.A., WASHINGTON, YAKIMA COUNTY: U.S.A., WASHINGTON, KITSAP COUNTY: Okanogan-Wenatchee National Forest. West of Jamestown S'Klallam Reservation. Clear Lake on Forest Service Road 540. 48.024773°, -122.996217°; uncertainty: 2000 Elev. 1006 m. m., Georef'd by Marina DeMarco, Invalid 46.65179° N, 121.32599° W; WGS 84, coordinates (located outside county bounds). uncertainty: 50 m., Source: GPS Phenology: Fruits. Origin: Introduced. Massive talus slope largely without vegetation except at base; north-facing. Holodiscus David Brownell s.n. discolor and Woodsia oregana common. Petals white; branches covered with prickles; locally 10 Oct 2019 common at base of talus. Phenology: Flowers and Fruits. Origin: Native. WTU-423245, WTU-V-060450 David Giblin 5235 17 Jun 2014 Rosaceae WTU-424723, WTU-V-067839 Rubus laciniatus Willd. U.S.A., WASHINGTON, KLICKITAT COUNTY: Railroad Island Park. Rosaceae 45.72214° N, 120.70325° W; WGS 84, Source: GPS Along dry roadside, with Bromus diandrus, Sisymbrium altissimum. Rubus ursinus Cham. & Schltdl. Petals white-pale pink. Phenology: Fruits. Origin: Introduced. U.S.A., WASHINGTON, CLARK COUNTY: Peter Dunwiddie PWD-19-173 23 Jul 2019 Lacamas Prairie Natural Area Preserve. with Peter Zika Elev. 195 ft. 45° 38' 20.1" N, 122° 27' 31" W; WGS 84, uncertainty: 3 m., Herbarium: WTU Source: GPS Oak/Ash association with Symphoricarpos albus, Trillium ovatum, Oemleria cerasiformis, Tellima grandiflora, Amelanchier alnifolia, Rosaceae Pseudotsuga menziesii. Phenology: Flowers. Origin: Native. Rubus arcticus L. Roger T. George 2018-34 3 May 2017 U.S.A., WASHINGTON, OKANOGAN COUNTY: Herbarium: WTU [locality and coordinates withheld; status in Washington: G5T5/S1 (Threatened; WANHP)] Elev. 5439 ft. Rosaceae In hummocky edge of a wetland complex that is mostly marshy in this location with Salix ssp., Rubus bifrons Vest Carex utriculata, Carex ssp., Picea engelmannii, Pinus contorta. The wetland also has fen U.S.A., WASHINGTON, ASOTIN COUNTY: components to it. Slope 0%, Aspect 90 degrees. Approximately 4 kilometers from end of Snake No flower, petals present. Need to relocate River Road on south side of Grande Ronde earlier in year to verify color, and also didn't dig River. deep enough to collect rhizomes. Origin: Native. Elev. 266 m. 46.06184° N, 116.99362° W; WGS 84, Erica Heinlen 2016-2 uncertainty: 30 m., Source: GPS Moist, north-facing rocky outcropping with 28 Jul 2016 Lomatium dissectum and Selaginella wallacei common. Locally common and excluding other WTU-423160, WTU-V-059993 vegetation. Phenology: Flowers. Origin: Introduced. David Giblin 4809 Rosaceae 22 May 2013 Rubus arcticus L. WTU-410041 U.S.A., WASHINGTON, OKANOGAN COUNTY: [locality and coordinates withheld; status in Washington: G5T5/S1 (Threatened; WANHP)] Elev. 5440 ft. In a peatland at high elevation with Poa sp., Carex utriculata, Carex scopulorum, Salix sp. Slope 0%, Aspect 270 degrees. Collected leaves that show rhizomes. Took pictures of dark pink flowers. 50 or so plants in a 400 sq. ft area. There may be more hiding under thick grass. Origin: Native. Erica Heinlen 2017-3 20 Jul 2017 WTU-423158, WTU-V-059990 Rosaceae Rosaceae Rubus nivalis Douglas ex Hook. Rubus praecox Bertol. U.S.A., WASHINGTON, KING COUNTY: U.S.A., OREGON, WASCO COUNTY: Greenwater Lakes Trail, 0.4 air km E of S end of 2.4 air km WSW of W end of Miller Island. N-most of Greenwater Lakes, Cascade Range. Elev. 60 m. Elev. 880 m. 45.6492° N, 120.9564° W; Source: Specimen 47.0885° N, 121.4488° W; Source: Specimen Label label, Georef'd by Marina DeMarco On dry sandy ground, full sun, at base of basalt Trailing evergreen, shaded forest floor, with cliffs, with Bromus tectorum, Achillea Maianthemum stellatum, Polystichum munitum, millefolium, Heterotheca villosa, Chondrilla Athyrium filix-femina, Trillium ovatum. Leaves juncea, Ericameria nauseosa. Arching adventive shiny dark green, old growth, occasional in area. shrub 2.5 m tall, petals white to pale pink. 10-11.5 x 7-8 mm, flowers 20 mm diameter, Peter F. Zika 29526 thorns the same color as stem; uncommon. U.S. Army Corps Columbia River Plant Survey. 27 May 2019 Peter F. Zika 29670 WTU-422872, WTU-V-059417 26 Jun 2019 WTU-421869, WTU-V-055218 Rosaceae Rubus bifrons Vest s.str. Rosaceae U.S.A., WASHINGTON, SNOHOMISH COUNTY: Rubus sp. Sunday Lake Road, 120 m N of Sunday Lake. Elev. 70 m. U.S.A., OREGON, BENTON COUNTY: 48.232° N, 122.2572° W; Source: Specimen Near NW Peavy Arboretum Road, 1 air km SSE Label of Calloway Reservoir. Thickets at edge of forest, partial shade, with Elev. 125 m. Alnus rubra, Acer macrophyllum, Populus 44.6554° N, 123.2315° W; Source: Specimen trichocarpa, Oemleria cerasiformis. Arching Label colonial shrubs, petals pale pink 12-20 x 13-14 Shaded streamside thicket, with Pseudotsuga mm, shaded stem surface green, with red-based menziesii, Rubus armeniacus. Cultivated shrub, prickles; common adventive. petals pink. Cultivated. Peter F. Zika 29647 Peter F. Zika 29752 with Richard E. Brainerd 24 Jun 2019 16 Sep 2019 WTU-422918, WTU-V-059536 WTU-423001, WTU-V-059589 Rosaceae Rubus bifrons Vest s.str. Rosaceae U.S.A., WASHINGTON, SNOHOMISH Rubus parviflorus Nutt. COUNTY: Sunday Lake Road, 120 m N of Sunday Lake. U.S.A., WASHINGTON, GRAYS HARBOR Elev. 70 m. COUNTY: 48.232° N, 122.2572° W; Source: Specimen Pacific Coast Range: 15 Overlake Lane, Label McCleary, 0.5 miles S of WA Hwy 8, ca 18 miles Thickets at edge of forest, partial shade with W of Olympia. Alnus rubra, Acer macrophyllum, Populus Elev. 104 m. trichocarpa, Oemleria cerasiformis. Arching 47.046943° N, 123.281231° W; T18N R5W colonial shrubs, petals pale pink 12-20 x 13-14 S14 SW4 NW4 NE4 SW4; Source: Specimen mm, shaded stem surface green, with red-based Label prickles; common adventive. Edge of Pseudotsuga menziesii-Alnus rubra forest with thicket of Rubus armentiacus, R. Peter F. Zika 29647 nutkanus, R. spectabilis, R. laciniatus, and R. ursinus. Shrub 5 ft tall 24 Jun 2019 Walter Fertig 32523 WTU-422919, WTU-V-059537 22 May 2018 WTU-423325, WTU-V-060059 Rosaceae Rosaceae Rubus parviflorus Nutt. Rubus praecox Bertol. U.S.A., WASHINGTON, COLUMBIA COUNTY: U.S.A., OREGON, UMATILLA COUNTY: Along Turkey Ck Trail (FS trail 3136), between Near N end of access road into McNary Wildlife Panjab and Tepee TH, Wenaha-Tucannon Nature Area, 1.2 air km W of the S end of John Wilderness, Umatilla NF. 17 mi. SE of Dayton, Day Dam. WA. Elev. 80 m. Elev. 4200 ft. 45.93096° N, 119.3125° W; Source: 46.1457° N, 117.7134° W; Source: Specimen Specimen Label Label At edges of grassy openings and on pond Mature fir forest. Thimbleberry. shores, full sun or partial shade over basalt, with Elaeagnus angustifolia. Arching adventive Heidi Jump 20180624-HJ02 1/2 shrubs 1.5 m tall, petals white or pale pink, 12-14 x 9-10 mm, shaded stems same color as 24 Jun 2018 prickles (green); common. WTU-423230, WTU-V-060415 Peter F. Zika 29661 26 Jun 2019 Rosaceae WTU-422925, WTU-V-059525 Rubus praecox Bertol. U.S.A., WASHINGTON, KING COUNTY: Rosaceae M Street NW, 3.2 air km E of Lake Dolloff, Auburn. Rubus bifrons Vest Elev. 15 m. 47.3284° N, 122.2442° W; Source: Specimen U.S.A., WASHINGTON, KLICKITAT COUNTY: Label SE side of gravel Maryhill Highway, 1.9 air km Adventive, sunny damp thickets near marsh, WSW of Preachers Eddy Island. with Phalaris arundinacea, Crataegus Elev. 55 m. monogyna. 2 m tall arching shrubs, shaded 45.7052° N, 120.7412° W primocanes with red-based green thorns on Riparian zone, full sun over basalt, with green stem, in sun stem and thorn dark purple; Artemisia dracunculus, Asclepias fascicularis. petals 12-15 x 9-11 mm, styles 2 mm, stamens Petals pink, 20 mm, prickles red-based on green 7.5 mm. [Sheet 2 of 2] stem; 17-72 flowers/inflorescence; adventive arching shrub. [U.S. Army Corps Columbia Peter F. Zika 29685 River Plant Survey] 1 Jul 2019 Peter F. Zika 29557 WTU-422941, WTU-V-059507 3 Jun 2019 WTU-422218, WTU-V-057774 Rosaceae Rubus praecox Bertol. Rosaceae U.S.A., WASHINGTON, KING COUNTY: Rubus parviflorus Nutt. M street NW, 3.2 air km E of Lake Dolloff, Auburn. U.S.A., WASHINGTON, CHELAN COUNTY: Elev. 15 m. Peshastin Creek, W of Etienne Creek. 47.3284° N, 122.2442° W; Source: Specimen 47.364391°, -120.652704°; WGS 84, Label uncertainty: 1500 m., Source: Digital Map, Adventive, sunny damp thickets near marsh, Georef'd by Crystal Shin with Phalaris arundinacea, Crataegus monogyna. 2 m tall arching shrubs, shaaded Harold Smith 745 primocanes with red-based green thorns on green stem, in sun stem and thorn dark purple; 31 May 1936 petals 12-15 x 9-11 mm, styles 2 mm, stamens 7.5 mm; common. [Sheet 1 of 2] WTU-38869, WTU-V-045444 Peter F. Zika 29685 1 Jul 2019 WTU-422950, WTU-V-059508 Rosaceae Rosaceae Rubus pubescens Raf. Rubus laciniatus Willd. U.S.A., COLORADO, DOUGLAS COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Mesic canyon, Billy Smith Gulch, tributary to Olympic National Forest. West Fork Humptulips River. Forest Sugar Creek, 7 mi. NE of Deckers Service Road 2204 approximately 2 km west of Campbell Grove 39.306279°, -105.196356°; WGS 84, Campground. uncertainty: 513 m., Source: Digital Map, Elev. 300 m. Georef'd by Isaac Afolayan, Tissue removed 47.47363° N, 123.70374° W; WGS 84, uncertainty: 100 m., by Gwen Ellis at University of Washington for Source: Digital map DNA analysis. February 4, 2020. Wet ditch along road with Oenanthe sarmentosa, Rubus nutkanus, Phenology: Fertile. Origin: Native. and Boykinia intermedia common. Locally common on roadside. Phenology: Vegetative. Origin: Introduced. W. A. Weber 13109 David Giblin 7109 27 Aug 2019 22 Jun 1967 Herbarium: WTU WTU-242185, WTU-V-060994 Rosaceae Rosaceae Rubus leucodermis Douglas ex Torr. & A. Gray Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, LINCOLN COUNTY: Miles-Creston Road approximately 2 km south of Hawks Cove U.S.A., WASHINGTON, CHELAN COUNTY: Campground. Okanogan-Wenatchee National Forest. White River Road. Elev. 523 m. Approximately 2 km east of Grasshopper Meadow Campground. 47.80424° N, 118.32224° W; WGS 84, uncertainty: 50 m., Elev. 596 m. Source: GPS 47.92873° N, 120.90678° W East-facing basalt face with seasonal seep down face and adjacent South-facing boulder and talus slope and adjacent rocky dry slope on other side of road. Scattered on dry shrub; prickly outcroppings in broken canopy, Pseudotsuga menziesii and Pinus shrubs with arching branches. Phenology: Flowers. Origin: Native. ponderosa pine forest. Pteridium aquilinum and Moehringia macrophyllum common. Rock brownish-orange, possibly ultramafic. David Giblin 6850 24 May 2019 Trailing vine; locally common at edge of talus; berries red. Phenology: Fruits. Origin: Native. Herbarium: WTU David Giblin 6880 16 Jul 2019 Rosaceae Herbarium: WTU Rubus idaeus L. ssp. strigosus (Michx.) Focke Rosaceae U.S.A., WASHINGTON, OKANOGAN COUNTY: Sinlahekin Wildlife Refuge. Forde Lake, northwest corner. Rubus leucodermis Douglas ex Torr. & A. Gray Elev. 470 m. 48.73706° N, 119.66973° W; WGS 84, uncertainty: 50 m. U.S.A., WASHINGTON, GRAYS HARBOR Large pond with shoreline dominated by Typha latifolia and shallow COUNTY: waters by Potamogeton richardsonii; adjacent Pinus Pacific Coast Range, woods west of Overlake ponderosa-dominated dry slope. Scattered in upland adjacent to Lane, 0.6 miles south of Hwy 8 and McCleary, pond; stems arching; lower leaves white. Phenology: Fruits. Origin: 18 miles W of Olympia. Native. Elev. 122 m. 47.048326° N, 123.284789° W; T18N R5W David Giblin 7359 19 Sep 2019 S14 NW4 SW4. Edge of Pseudotsuga menziesii-Tsuga Herbarium: WTU heterophylla-Frangula purshiana forest and gravel logging road. Soil loamy clay to rocky. Scrambling shrub over Gaultheria shallon. Rosaceae Leaflets mostly 3, stems glaucous on young growth, flowers with stamens and pistils. Rubus lasiococcus A. Gray Walter Fertig 33140 U.S.A., WASHINGTON, SKAMANIA COUNTY: Gifford Pinchot National Forest. Bluff Mountain Trail just below Silver 22 Jul 2018 Star Summit and for 500 meters east along trail. Elev. 1232 m. WTU-423605, WTU-V-063280 45.74815° N, 122.23737° W; WGS 84, uncertainty: 500 m., Source: GPS Subalpine meadows with exposed rock and talus in places; Xerophyllum tenax, Eriophyllum lanatum and Vaccinium sp. common. Scattered in dry soil; stoloniferous; petals white. Phenology: Flowers. Origin: Native. David Giblin 7536 25 Jun 2020 Herbarium: WTU Rosaceae Rosaceae Rubus ursinus Cham. & Schltdl. Rubus ursinus Cham. & Schltdl. U.S.A., WASHINGTON, SKAMANIA COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Gifford Pinchot National Forest. Bluff Mountain Trail just below Silver Anderson Butte, between West Fork Satsop and Wynoochee Rivers. Star Summit and for 500 meters east along trail. South side, on large rock outcrop at end of road bed. Elev. 1232 m. Elev. 2765 ft. 45.74815° N, 122.23737° W; WGS 84, uncertainty: 500 m., 47.38042° N, 123.55962° W; WGS 84, Source: GPS Source: GPS Mid elevation rock outcrop in coniferous forest. Very coarse sandy Subalpine meadows with exposed rock and talus in places; soil. With Pseudotsuga menziesii, Thuja plicata, Tsuga heterophylla, Xerophyllum tenax, Eriophyllum lanatum and Vaccinium sp. Holodiscus discolor, Leucanthemum vulgare, Hypericum perforatum, common. Locally common on dry south-facing slope; petals white. Anaphalis margaritacea, Acer circinatum, Achillea millefolium. Phenology: Flowers. Origin: Native. Uncommon on balds, abundant in forest. Phenology: Flowers. Origin: Native. David Giblin 7537 25 Jun 2020 Regina Johnson 2021-045 19 Jun 2021 Herbarium: WTU Herbarium: WTU Rosaceae Rosaceae Rubus spectabilis Pursh Rubus spectabilis Pursh U.S.A., WASHINGTON, SKAMANIA COUNTY: Gifford Pinchot National Forest. Mckinley Ride parking area to U.S.A., WASHINGTON, ISLAND COUNTY: approximately 2 kms west along Bluff Mountain Trail. Hobbit Trail, Camano Island shrub wetland (former estuarine Elev. 1076 m. wetland). 45.78005° N, 122.16697° W; WGS 84, uncertainty: 2000 m., 48.213213° N, 122.448391° W Source: GPS Saturated/moist, part sun, scrub-shrub swamp - former estuary. Mix of Abies lasiocarpa forest edge and wind swept talus field along Common, shrub, rhizomes, erect. Phenology: Fruits. Origin: Native. the ridgeline dominated by Xerophyllum tenax, Castilleja miniata, and Penstemon fruticosus. Scattered on forest edge; petals Patricia Johnson NWC21-WA-10017-Q14 26 May 2021 purple-red. Phenology: Flowers. Origin: Native. Herbarium: WTU David Giblin 7501 24 Jun 2020 Herbarium: WTU Rosaceae Rubus bifrons Vest Rosaceae U.S.A., WASHINGTON, ISLAND COUNTY: Rubus parviflorus Nutt. Hobbit Trail, Camano Island shrub wetland (former estuarine wetland). U.S.A., WASHINGTON, SKAMANIA COUNTY: 48.213213° N, 122.448391° W Gifford Pinchot National Forest. Mckinley Ride parking area to Saturated/moist, full sun, scrub-shrub swamp - former estuary. approximately 2 kms west along Bluff Mountain Trail. Common, vine, rhizomes, arubrching, climbing Phenology: Fertile. Elev. 1076 m. Origin: Introduced. 45.78005° N, 122.16697° W; WGS 84, uncertainty: 2000 m., Source: GPS Patricia Johnson NWC21-WA-10017-Q23 26 May 2021 Mix of Abies lasiocarpa forest edge and wind swept talus field along the ridgeline dominated by Xerophyllum tenax, Castilleja miniata, Herbarium: WTU and Penstemon fruticosus. Locally common on forest edge; petals white; shrubs to 1.5 meters tall. Phenology: Flowers. Origin: Native. Rosaceae David Giblin 7500 24 Jun 2020 Rubus ursinus Cham. & Schltdl. Herbarium: WTU U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: A sedge meadow dominated valley surrounded by managed Rosaceae timberland that is likely in the North River watershed of Willapa Bay. 47.431833° N, 123.97305° W Rubus lasiococcus A. Gray Moist; full sun; headwater valley marsh - clibming on Carex obnupta. Common, vine, rhizomes, trailing, climbing; flowers white, 5 petals. U.S.A., WASHINGTON, YAKIMA COUNTY: Phenology: Flowers and Fruits. Origin: Native. Okanogan-Wenatchee National Forest. Sand Ridge Trail. Approximately 5 air km east of White Pass. Shellock Lake. Patricia Johnson NWC21-WA-10045-Q24 2 Jun 2021 Elev. 1482 m. 46.67936° N, 121.34132° W; WGS 84, uncertainty: 200 m., Herbarium: WTU Source: GPS Small montane lake at base of north-facing basalt scree slope with Vaccinium sp., Xerophyllum tenax, Phyllodoce empetriformis, and Rhododendron columbianum common, Mosquitoes plaque like. Scattered in moist upland; stems to 50 cm tall. Phenology: Flowers. Origin: Native. David Giblin 7586 14 Jul 2020 Herbarium: WTU Rosaceae Rubus spectabilis Pursh U.S.A., WASHINGTON, CLALLAM COUNTY: Adjacent to an unnamed tributary to the Soleduck River, west of the community of Shuwah. 48.031666° N, 124.397378° W Saturated to very moist, part shade, forested wetland near creek. Common, shrub, erect; Flowers pink, 5 petals. Phenology: Fruits. Origin: Native. Patricia Johnson NWC21-WA-10039-Q2 29 Jun 2021 Herbarium: WTU