Arch. Bioi. Sci., Belgrade, 55 (1-2), 23-32, 2003.
r .ON THE DIVERSITY OF SOME SPIDERS (ARANEA : ARACHNIDA) FROM SERBIA " 1<::' L)< :' ;f.:' B. P. M. CURCIL:, 'C. C. DELTSHEV, 'G. A. BLAGOEV,S. B. CURCiC and V. T. TOMIC
Institute of Zoology, Faculty of Biology, University of Belgrade, 11000 Belgra,qe? Serbia and Montenegro; Centre for Biospeleology of Southeast Europe, 11000 Belgrade, Serbi~ ~d M6ntenegro; , and lInstitute of Zoology, Bulgarian Academy of SCiences, I 000 So~~~ Bulgaria.
Abstract - A total of 76 species from 16 families: Dysderidae (6), Theridiidae (3), Nesticidae (1), Linyphiidae (35), Tetragnathidae (1), Araneidae(l), Lycosidae (5), Agelenidae (3), Cybaeidae (1), Dictynidae (1), Amaurobiidae (5), Liocranidae (2), Clubionidae (3), Gnaphosidae (3), Thomisidae (3) and Salticidae (3) were established from 42 localities in Serbia. Ten species and three genera are new for the Serbian spider fauna. The most diverse is the family Linyphiidae, otherwise represented by 35 species.
UDC
INTRODUCTION 1970). Ten years later, N i k 0 lie and P 0 I e n e c (1981) combined the data concerning Yugoslav spiders The existing information on Serbian spiders-is due and reported 525 species from 35 families from Serbia. to the efforts of many .araneologists from different More recent publications list the cave fauna (D eel e m countries. The first information concerning the spiders of an - Rei n h 0 I d 1985; De e I e m a n - Rei n h 0 I d Serbia appeared at the end ·of 19th century in the paper and De e I e m a n 1988; T h a I e rand Ho fer 1987; of Spa s 0 j e vie (1891). Three years later, C h Y z e r Del t s he v et at. 1996; D. e Its h e v and Cur c i c and K ul c z Y Ii ski (1894, 1897) announced more 1997; Del t she v et al. 1997; Del t she v et al. precise data. The information concerned single localities 1998; Cur c i c et at. 1999a) and leaf-litter spiders in Northern Serbia. Much more information was reported of Western and Eastern Serbia (D e Its h e v et at. by B res j an c e va (1907), who recorded 148 species. 1998; Cu.rcic et at. 1999b;Curcic et al. 1999c; The first huge work came from S to j ice vie (1929). Cur c i c et at. 2000; Del t she v and Cur c i c He reported 432 species from 34 families, which 2002). constituted a rev\':le o.f all literature available by that time. Finally, Del t she v et al. (2003) concluded that A considerable contribution was made by K rat 0 - the spider fauna of Serbia is represented by 618 species, ch v f I (1934) who published the list of cave spiders in belonging to 221 genera and 36 families. The number of Yugoslavia. Fag e (1931) published a revue of cave species is comparatively high compared with a number spiders in EJlrope, where 71 species from the Balkan of spiders recorded from other countries from the Balkan Peninsula (9 of which from Serbian caves) were re Peninsula: the highest number of species is recorded ported. Some years later, D r ens k y (1936) compiled for the territories of Bulgaria (910) (D e Its h e v and all faunistic data about the spiders of the Balkan B I ago e v 2001), Greece (810) (B 0 s m a n s in Peninsula mentioned in the literature and reported press), Croatia (614) (D e Its h e v 1999) and Ma 447 species from 34 families for Serbia. In this period cedonia (595). This reachness, however, depends not Ko I 0 s v a r y (1938, 1940) published a list of spiders only on the size of the regions, but also on the degree of of Yugoslavia that included 40 species from the terri the explorations by araneologists. Best represented are tory of Serbia. In the same period K rat 0 c h v f I and the families Linyphiidae (156 species), Lycosidae (62 Mill e r (1938) continued their work of cave spiders species), Gnaphosidae (59 species), Salticidae (49 spe from the Balkan countries, with an emphasis on research cies), Araneidae (48 species), Theridiidae (42 species) on the distribution of cave Centromerus species. S i I - and Thomisidae (37 species). h a v y (1944) described new species from the family The goal of the present study is to give a critical Thomisidae. Much later, Cur c i c (1969) examined the systematic revue as well as a zoogeographical analysis cave spiders in the Iron Gate region and reported 17 of the spiders collected from 42 additional localities in species. Some data concerning Serbian spiders can be Serbia, based on intensive field research and new data found in the papers of van He I sdi n g e n (1969, accumulated (Table 1).
23 24 B. P. M. CURCIC et al.
Table 1. - List of established spider species in Serbia. Abbreviations: f == female, ff = females, m = male, mm = males, juv. = juvenile(s). Species new to the fauna of Serbia in bold letters.
Geousandspeeies Collectol'$*
DYSDERIDAE
Oysdera adriatica KulCzynski. 11397 Rimski Most, v. Kumanica, nr. lvanjica, 4ft. 1 m. 24 SPMC, SBC, ;I)v" 14.05.2001. SMM.SVD Oysdera ninrH (Canestrioi. 1868) IzvorS$kioac, Mt Allala. or. Belgrade. 1 f. 12juv•• 16. SSC 04.2001. Oysderocrates storkani (Kratoclwll, V. Osonica. MlCeroemo, nr.lvaojica, 2 mm, 5juv., 16- SPMC. S8C. 1935) 17. OB. 2001; nr. l.,ake Ti6ar (:; DajiQko Jezero Lake). SMM.SVD Mt .. Golija. nr.lvanjics.• 1 rn. August 2001; Zrnajinac. v. Milice. Mt Radoeelo, nr. Ivanjica. 1 m, .13. OB. 2001. Harpactea rubicum:l(l{C .. L Koch. 183S) Toranj, Ml Avala. nr. Selg~de, 1 m, 23juv., 1(t 04. S8C 2001. Plarpactea saeva (Herman, 1879) In front of the Mala Pe6ina Cave, v.Rli,nf. Kotraza. 1 BPMC. SVD m•. 9 j~" .• 02. 05.2()01 . .Kaemiscarnicus Deeleman.-Reinhold, L6vaeka Ce$ma,. v. ljepojeviCi, Mt. Javor (1350 rn), 2 BPMC.SSC. 1993 mm. 11-12,~. 2001; v. Mo{;ioci,nr. KatiCi,Mt. Mooanj SMM.SVD (11501'1'1), 2ft. 2 mm, 09. 08. 2001 .
THERIDIIDAE
EuryopiS flavomaculata (C. L Koch, 10 front of the MalaPe6ina CelVE!, v.Rti,nr. KotraiCl. 1f. SPMC. SVD 1836) 2 mm.2juv•• 02. 05.2001. Robertus livid1.ls (Slackwall, ·1836) Golij$~ Reka, MtGolija, nr.Ivanjjca.1 f.13Jl.lv,. 12. SPMG. SEM. 05. 2001;SiserVode, v. Koritnik, Mt. Gotija (1300 m). 1 S8C.BMM, f.7 mm,3 j\.IV.,07,08. 2001; Lovacka(~e$fO NESTICIDAE Nesticus cellulant.ls (Clerck. 1757) Peth~Pecina.Cave. v. Petnica.nr.·Valjcwo.a ft. 1. rh. SPMG.SeM. 6 jull" 21. 06. 2001; Hadtl-Pro(illnov~ PeCinaC~ve. v. $SC.BMM. Ra~~i. or. IlIanjica, 1 f. 11. OS, 2001; Virovska pecinaSVO.GC Cave,V. Virovo,nr.. ... 5 ft. 61uII., 30, ..04.2001; Disina Peeina Calle, v. .Mt. Miroo. 1 f, 2 jU\I •• 18. 08.2001. UNYPHIIDAE ./3olyphantesaltic&p$ (Sundavall, Lovae~ Cesn"lq, v •• t~pojeviQi, Mt. Javor (1$SOm),nr. SPMG.SSC. .1833) ...... lvahjica, 1f, 11-12.08.2001. . / BMM,SVD BO/fphantes /colosvaryi(CapPOfiacQ, V.OsoniCa,Mt Cememo, nr. Ivanjica, 2 mm, 5juv., 1f3.. 8Pf.!1C; sac. 1936) 17; oa. 2001, BMM.SVO Centromeru.scavemarum (L. Koch. Mala Pe6ina Celve. v. Rti. nr. Kotrata.n.2jUII., 12. OS. BPMC. SEM. 1872) 2001; 8 ft.S juv" 30 ..04. 2001;liadti·Prodanova SSC.SMM. Pecina Cave. v, Raieici. nr. IvanjiC$, 3 ft, 4 mm. 1 juv.• SVO,GC 11. De .. 2001;3 ff. 1JUY, •• 14.05 2001; Vi.rovska Pe6ina Cave. v. Virovo,nl'.Arilje, 111"\,$0.04. 200.1; 8!ser Vode. v.Korithik. Mt. Golija (130~ rh). hr. lvanjica, 141f, 07. ~. 2001;,4. cave oPPRsite to the Kr&t$ta PeCiOCl SPIDES FROM SERBIA 25 Table 1 - Continued. Gen .... s and species Locality Collectors'" Cave. v. Trudovo. Mt Javor. nr. Nova Varo!>,1 t,1 juv., 22. OB. 2001; v.MqciQCi. nr.l Table 1 - Continued. Genus, and species Locality Cotlecto(s* MiHee, Mt.RadQ6elo"nr. Ivanjica, 4ff, 13. OS. 2001 Lepthyphantes leprosus ( Table 1 - Continued. G~nusandspeci~s Locality Collectors* Kapilijama" Cave, V. Truc;lovo, Mt. Javor. nr. Nova Varo~ , 4 ft, 1 mrn. 4 jllV., 19.,08..2091,; a cave i!lt!'le Arsovica Kr~. v. Maskova, Of: Ku~iCi . Mt. Javor.20 ft. 20 mm. t6juv, 20. 08. 2001 ; peci(laBanja Cave, v. Miroe. Mt. Miroc. e ft, 2 mm, 17, OS.?001 ; Velika Grad i ~nica Cave, V. Miroc, Mt. Miroc, 2ft. 16.08. 2001. Saaristoa firma (O.P.-Cambridge. Golijska Re~a . Mt. Golija, or. IvanjiCa" 1 m, 01 ~03 . " 12. J:jPMC. sec, 1900) 052001 . BMM,SVD ScoiarguspilosusSimon, 191.3 GolijskaReke, Mt. Golija, ot. Ivanjica; 1 t. 1 m, August BPMc. sec, 2001; BiserVoc;le, v. Koritnik, Mt. Golija (1300 m). or. 8MM,SVD Ivanjica, 1 m. 07. 08.2001; v. Osonica, Mt. Cememo. Ilf.lv8nJica, 1 m; 16-17. os. 2001 :Zmajinac. v. Milice, Mt. Radole{o, nr. Ivanjica, 1 f,13. OS. 200f. St~monyphantes/in~atus (Lirmaeus. Golijska Reka, Mt. Golija ~ nr. Ivanjica, 1, '(,12.0$,2001; BPMC.SBC. 1758) v.O~onica, Mt. Cemerno. nr: Ivanjica, 1 f, 16-17. 08. BMM. SVD 2001 . Tenuiphantes flavipes (Bleckwan, 1 a54) Mt Avala,nr. eelgrade, 1 t, 09. 06_ 2001; in front of the BPMC.SBC. Mala Petina Cave. veRti, nr. Kotra!a, 1 f.4Juv., 02; 05. SVD 2001.. Tenulphantes flQriana '(van Ml, AvalC\, or. Belgrade. 1 f; 1 m, 3juv. • 09. ·06.2001. SBC Helsdfngen, 1977) Tenuiphantes tenebricola (Wider, 1834) Golijska Reka, Mt. GOlij;a. nr. Iv.anjica, 1 f,2 mm. AugU$t BPMC. SEM. 2001; Lake Ti6ar. Mt. Gonja, nr.. Ivanjica, 3 ft, 2 mm, . SBC, BMM. August 2001 ; BiserVode. v. Koritnik. Mt.Golija (1300 SVD m). nr. Ivanjica. 1 f,07: 0S. 2001; LovaekaCesma, v. Ljepojevici,Mt. Javor (1350 m),nr. lvanjica.4 ft, 1' m, 11-12.08.2001; y.. Motioti. nr.Ka~c i ,Mt Mu~anj (1150 m). 2ft. 09. 08. 2001; v. Osonica, Mt. Ce~rno . ni'. Ivanjica. 7 ft, 5mm, 1&;17. OS. 2001; Zmajiilac, v. Milice; Mt.. Radoeelo, hr. Ivanjica, 22 ·ft. 12 mm, 13:08.2001. 1richoncus hackmani Millidge, 1955 Rimski Most. v. Kumanica; nr. Ivaniica, 1 f. 1 11').24 juv .• BPMC, SEM, 14. 05. 2001. sec, BMM, svb Walckenaeria clJcullata(C. L Koctt, ZmaJinac, v. MiliCe. Mt. Radocelo. nt. Ivanjica. 1 f, 13. BPMC. SEM. 1836) " , " OS. 2001. $BC.BMM, SVD Walckenaeria furcilata (Menge. 1868) Mt.Avala. nr. Belgrade; 1 f. 09. 06. 2001. SBC Walckenaeria mitrata (Menge, 1868) GolijskaReka, Mt. Golija, nr. Ivanjica., 1 t, 1 m. 01-03. BPMC, SEM, 05. 2001; Lake Ticar, Mt. .Golija. nr. •lvanjica, 1f; August sec. BMM 2001 ; BiserVode, v::Koritn[k. Mt.GOlija(t300m); nr. Ivanjica, 11. 07. OB. 2001. .' TETRAGNATHIDAE r ~ Meta mfjnardl (latreil~,1804~ , Mala Petiha ~ Cave, v. Rti, nr; Kqtratc;l, 3 ff.<30,, 04J200W SeM;' SBO.'·;'·, A cave <>pposite to the Krstata Pecina.Cave,v. . BMM, GC ;TrYdOvp, Mt' Javot. hr. Nova Varo~, 1 m. 5jlJv., 22.08. 20Q;1 ;Krstata Pacina Cave. v. Debelja, Mt. Javor, nr. Nova Varo$, 1 f, 5 juv., 21 . 08. 2001 ; VelikaGradisnica , > " ' c Ca"e " v . Mir~, Mt. Miroc, 10 juv., 16,. ~ . J~OO1 . - ; " " -. • ARANEIDAE ~ : . -- ~- ', .'. ' -~.- Cerciciiaprominens (Westrlng, 1851 ) Rimski Most, v ..Kumanica d lrrlvanjica, 1f; 14. 015 . BPMC.SEM, . 2001. · , 1 .. SBC,BMM;' SVD 28 B. P. M. CURCIC et al. Table 1 - Continued. Genus and species locality Collectors· LYCOSIDAE Pardosa amentata (Clerck. 1757) V. MoCioci. nr. Katici,Mt. Mue AGELENIDAE Histoponaluxurians (Kulczynski, 1897) Izyor Sakinac, Mt.Avala. or. . B~fgradei2 ft;5 juv,.16. BPMC, SEM, 04.2001; 1 f, 5 juv., 16. QEl, 2001; Mt. Avala, or. SBC,BMM, Belgrade, 2 ft, 5juv., 09: 06, 2001; Golijska Reka, ty1t. 'SVD Golrja, nr. Ivanjica,3 ft', 2mm, Ot.03. 05.2001; 2 ft, 4 . jOV" .12.. 05.2001; 6ff.2juv., August20Ql;Lpvacka Cesma, v; .LJepojevici, Mt Javor (1350 m), nr. rvanjica; 6 ft, 1 rn, 11 ~ 12. 08.2001; V. Moeioci, tu. Katici, Mt M'-h'::.anj(11!$0 m), 3ft, 1 m,09; 08. Z001;v. O~()nica, Mt . Q~memo.l'lr.lvanji~i 5 ft,23 JOV., 16·17; 0.8 .. 2001 ; ZmaJinac, 1/. Milice; Mt Radotelo, I'Ir.lvanjica,. 12ft, 2 mm, 13. 08. 2001. Histopona torpida fC, LKoch,1837) Biser VOde, v. Kpritnil<;, Mt. Golija(1300 nil, nr,lvanjica, BPMC, SNlM, 2ft, 07.08.2001. SVD Tegenaria doinestica (Clerck, 1757) PetnickaPecinaGave. v : P~tnica. nr. Valjevo, 1 (21. SBC 06. 2001, CYBAEIDAE Cybaeusbalkanus Deltshev. 1997 Krstata PeGina Gave, 11 . Debelja, Mt Javor, nr; Nova saC, BMM Varos,1f, 1 m,.21. 08,. 200t. DYCTINIDAE Oictyna arundinacfia (Linnaeus, 1758) Gc;>IiJskaReka, Mt Gplija, nr.lvanjica,l m, 01 ,,03.05. BPMC, SBC, 2001. · BMM, SVD AMAUROBIIDAE AmaurobiuspallidusL Koch, 1868 Izvor Sakinac,Mt.Avala; . nr.Belg~qe. 1 f,1 m;' 16. 64: SBC 2001; 1 f, 2 j(Jv., 16. 06.200t Mt. AYala, nr. SelQrade; t f. 2 juv:, 1 2. 06.2001; roadfork to v. Zi,lce. v. Beli Pdtok and\;. Pin()$ava,MLAvala,-hr: Belg~de. 1t, 2 jw.1 6. "04.2001. ' .. . Coe/otes inermis (L KOCh; 1855) GolijskaReka, Mt. GOlija, nr.lvanjica.1 m, August ~PMC,SBC, 2001; LoVacka Cesma, ,,: LjepOjevi61, Mt Jayor{1350 BMM,SVD m). 3 mm, 1.1·12.08. 2001. CoelotesjlJ{initschi (Drensky, 1915) KrstataPeci9aCave, v ..Debelja,Mt. Jav6r,N~ Nova SBC. BMM Varo$. 1 f,21. 08. 2001 , •. . \ Coe/otes·titaniacusBr.ignoU; 19Ti GolijSkaReka. Mt. Golija, nr.lvanjica,lm, 01-03, P5. BfJrI£, SEM, 2001. . ~,8M\1,~ SPIDES FROM SERBIA 29 Table 1 - Continued. '. ... GenusanQs~ies - Locality Collectors. . Utocoraslongispinus(Kulczynski. H}~7)lzvor Sakinac; M~Av~~ nnSelgrade, 2ft, 12juv., 1fi. sac 04 . .2001; en routeJzV9rS",kioac -:roranj, Mt. Avala, nr. Belgrade, 1 t, 2 juv., '16. 04;2001; en route v. Zuce - v, Beli Potok.Jv1LAVala; M; Belgr'liIde, 1f,3juv., 1t). 04, 20()1. UOCRANIDAE 1, : " ;!l UoCraniJm rupicola(Wcllcl . CLUBIONI.oAE C/ubibna comta(C, l.Koch,t~39) V. Osoniqa, Mt. Cel'l'lerno, .nr. lvanjica, 2 ff,1.6.;17. 08. BPMC, SBC, 2001, BMM, SVD CIl)/)iona saxatills L, ~och, 18.66 V. Osonica, MtCemerno, nr. Ivanjica. 1 f, 16'--17.08. 8PMC, S8C, 2001. BMM,SVD Clubion,a terrestris Westrlng, 1851 Ml Avala, or. Belgrade,1 t,1 juv., 09. 06.2001; en SBC rou,le IZV9rSakinac - Toranj, Ml. Ayallil, nr. Belgn:tde, 1 t, 16 ~ 04. 2001, . GNAPHOSIDAE Calli/epiS noctuma (Linnaeus, 175~) Rimski Most, v. Kumanica,. nr.lvanjica, 1 f, 14.05. BPMC, SEM, 2001 . SBC,BMM, SVD Micaria (u/gens (Walckenaer, 1802) Rirnski Most, v.Kumaniea, nr. Ivanjica,1 m, 14. 05. 8PMC, SEM, 2001. S8C, BMM, svo le/otes apiicorum(l. Koch, 1876) V, Osonica, Mt.Cemerno, or. Ivanjica.1 f. 16~ 17. 08. BPMC, SVD 2001 . Ozyptiia b/ackwal/iSimon. 1875 Mt. Avala, nr. Belgrade, 1 rn, 6juv" 09.06.. 2001; in sac fl'ontoftheMala PeCinaC~ve, V. Rti,nr. KotrlilZa, 1 f, 1 1l"I' 7 j(JV., 02. 05 .. 2001; enro\-jt~ v ~ Zuce - v. Beli . Potok, Mt. Avala. nr.Belgrade,1 m,1 juv., 16. 04. 2"001. . Xysticus /tm/O C. l. Koch, 1 835 Mt. Avala,nr. Belgrade, 1 m, 12.1'06. 2001; roadforkto SBC . v. Zure, v. Beli Potok and v. Pinosava. Mt. Avala, nr ~ Belgtade, 1 rn, 16. 04. 20p1 . _ . Xysticus sabu/osUs(M~nge, 1875) V. Osonica, Mt.Cemerno, nr. Ivanjica,1 m,16-1T 08. BPMC, SBo..).,. 2001. BMM,SVD SALTICIDAE Bal/us chalybeius (Walckenaer, 1802) Toranj, Mt. Avala, nr. aelgrade,1 rn, 16. 04. 2001. sec He!iophanus cupreus (Walckenaer, ~jmski Most, v. KI../manica,nr, IVclnJica, 2 ff. 14. 05. BPMC,SBC, 1802) 2001. aMM,SVD Neolf~ticulatus (Blackwall, 1853) Izvor Sakinac, Mt. Avallil, nr. Belgrade, 2 ff, 2 mm, 1 BPMC,·SBC, juv .• 16. 04. 2001; 1 t, 1 juv., 16. 04. 2001;Mt. Avala, aMM,sVD ne Belgrade,9ff, 2ml'l'l, 2juv., 09. 06. 2001: Rimski M9st, v. Kumanica. nr. Ivanjica, 5 ff, 1 juv., 14. 05. 30 B. P. M. CURCIC et al. Table 1 - Continued. Genus and species Locality Collec:tors* 2001.; in front of Virovska PecinaCavEt. v. Virovo. nr. Ariije. 1 m. 30. 04. 2()01; ToranJ. Mt. Avala, nr. Belgrade. 1 m,a juv .• 16.04. 2001; v. MociOci, nr. KatiCi, Mt. Mucanj (1150 m),2ff, .Q9. 08.2001. *Collectors: BPMC = Bo!idar P. M. CurCic.SBC =Sreeko B. Curcic. BMM =Bojan M. Mitic. SVD = Sonja V. Ojukic,. SEM = Slobodan E. Makarov,GC = Gorein Cvijanovic. CONCLUSION some cave-dwelling spiders (Araneae, Arachnida) from Serbia, Yugoslavia. Archives of Biological Sciences 51 (1), 7P-8P, The spider fauna of Serbia is well represented on Belgrade. the territory of Serbia but the species are distributed CurCii:, B. P. M., Deltshev, C., Makarov, S. E~, Karamata, O. S., unequally in different areas and districts. The number of Tomii:, V. T. (1999b). First report on some leaf-litter spiders established species (628) and genera (224) is com (Araneae, Arachnida) from Serbia. Archives of Biological Sciences 51 (1), 15P-16P, Belgrade. paratively high to the number of spiders recorded from CurCii:, B. P. M., Deltshev, C, Makarov, S. E., Tomii:, V. T., other countries from the Balkan Peninsula. The faunistic Curcii:, S. B., Curcii:, N. B. (1999c). On some some leaf diversity. of Serbian spiders shows that the Balkan litter spiders (Araneae, Arachnida) from West Serbia, Peninsula is a territory of considerable species reachness. Yugoslavia. Archives of Biological Sciences 51 (1), 21P- In the zoogeographical respect, the widely distributed 22P, Belgrade. spiders are dominant, which shows that the process of Deeleman-Reinhold, C, Deeleman, P. (1988). Revision des Dys colonization is considerable. The endemic species re derinae (Araneae, Dysderidae), les especes Mediterra present the local character of this fauna and point to the neennes occidentales ex~eptees. Tijdschrift,:voor Entomo logie 131,141-269, Amsterdam. main role of the genesis of the Balkan Peninsula in its Deeleman-Reinhold, C L, (1985). Contribution a la connaissance origin and evolution. 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M. liYPqHli, IX. X. ,l{EJIqEB, If. A. EJIAfOEB, C. E. liypqHli H B. T. TOMHli HHCWUWYW 3a 300J/OZUjy, EUOJ/O/uKU