Allan Hancock Foundation Monograph No. 11
DEEP-WATER POLYCHAETES FROM A TRANSECT OFF CENTRAL OREGON
by
Kristian Fauchald and Danil R. Hancock
Published by the Allan Hancock Foundation and the Institute for Marine and Coastal Studies, University of Southern California, Los Angeles, California 9Ü0Ü7 CONTENTS
Introduction 3 Kaniilv Oibiniidac 5 Familv Paraonidac 6 Family Cossuridae 9 Family Spionidae 11 Familv Cirratiilidac 14 F"amilv (lapiu'llidac 15 F'amilv Maldanidac 17 Famih Ophcliidac IX Familv Scalibrcgmidae 20 Familv PlivilodcK idac 21 Familv Folynoidac 23 Familv Sigalionidae 23 F'amilv Hesionidat' 24 Familv Pilargiidac 25 Family Syllidae 26 F'amilv Xercdiae 26 Faniily Glyceridae 26 Family Goniadidac 27 Family Nephtyidae 27 F'amily Tomopteridae 28 F'amilv Amphinomidae 28 Family Onuphidac 29 Familv Finiicidao 32 Familv Lumbrineridae 32 Familv Arabcllidac 35 Family Dorvilleidac 35 Familv Sternaspidae 35 Familv Ovveniidae 35 Family Flabclligci idae 36 Family Fauveliopsidac 37 Family Sabellariidae 38 F^amilv Pectinariidac 39 Familv Ampharetidae 39 Family Fcrcbellidac 42 Family Trichobranthidae 43 Family Sabellidae Malmgren 43 Station List 53 Illustrations 54 Literature Cited 70 Deep-water Polychaetes from a Transect off Central Oregon
by Kristian Fauchald and Danil R. Hancock
The polvrhaclc fauna of Oregon lias been studied very lilUe. The only large survey to date was by Hartman and Reish (1950); a few additional records can be found scattered in expedition reports. A list of polvchaetes from shelf areas identified bv Donald ). Reish was published by Carev (1972). The pre.sent. material was coliecied to characterize bathval and abyssal enviroinnenis off Oregon. The polychaete fraction of this material was treated first by Hancock (1969); later, pails of the material were reviewed by the senior author, and descriptions and illustrations were added. The samples were taken in a transect from Yaquina Bay, Oregon, extendingabout 320 km offshore. The dci)th langed from 100 lo 2900 m. The permanent stations, each visited a variable number of times, are referred toas .N.AI) stations. NAD stations with numbers below nine are in waters shallower than 200 m; those with numbers above 20 are on the Cascade Abyssal Plain, fhe iniermediate stations are on a steep slope. Figine 1 shows the position of the transect; the insert is a depth profile. Details on the stations can be found ill the station lisl. ,'\ total of one hundred and forty-lwo species of polychaetes are reported upon lieie. One hundred thirty-five have been identified to species and named; another .seven cannot be identified to species but belong lo geneia not otherwise represented. Thirty-seven families are represented. One, the iOMOP- TERID.AK, conlainsexclusivelv pelagic members, and the present specimens must either have been caught in the water colimin or have been brought in during the screening process. Hartman and Reish (19.50) reported 136 species from intcrtidal areas and shallow water off Oregon. Thirteen of these species have also been found in the present study. Carey (1972) reported 102 species from shellclepths off Oregon; his study was based in part on the same material treated here.' Synonyms have been eliminated as imu h as possible, as have differences of opinion on the identity of certain species. In view of the verv limited investigaiions in Oregon, the number of species known from Oregon compares favorably with the number known in Washington and California (see Banse and Hobson, 1974; Hartman, 1968 and 1969). The bathymétrie distribution of the species in the present material denumstrates the presence of a distinct deep-slope fauna, as well as a shelf fauna, of f Oregon. A few species are limited to the shelf region only (Xaineris unrinata; Polydora braihycephala; Travisia, nedr giga.s; and Artacamella havrorki). All others are limited to waters deeper than 200 m. This paper treats only two NAD stations from the upper slope (200 • 800 m), each of which was sampled only once. In contrast, stations farther down the slope are more numerous and were sampled more irecjuently. (See Fig. 1 and the Station List). The material is probably ade(iuale for deep-slope depths, given ihe limitations of the sampling gear, screenings, and sample processing specified b> I lancock ( 1969). However, the report is not adequate for the upper sfope and shelf. Therefore, no bathytnetric subdivisions can be indicated except for a distinct separation between the shelf fauna and the deep-slope fauna, sinced less than 10% of the species have been found in both areas. The best-represented families are: PARAONIDAE, SPIONIDAK, and LUMBRINERIDAE (11 species each); ONL'PHIDAE (10 species); A.VIPHARETIDAE (9species); CAPlTELI.lDAEand MALDANIDAE (7 species each); ORBINIIDAE, OPHELIIDAE, and PHYLLODOCIDAE (6 species each), PILARiillDAE and TEREBELLIDAE (5 species each). The number of cirratulid species reported is very low. More distinct taxa appear to be present, but current taxonomic practice does not allow species ideniification in most cirratulid genera.
'.All species cuneiilly kiiouii Iroiii Oregon are lislccl in Table 1. Allan Hancock Foundation Monograph No. 11
The familial composition of the deep-water fauna closely resembles that reported from other areas (Hartman, 1965; Hartman and Fauchald, 1971; Fauchald, 1972a). At the species level, it is very similar to the launa reported from western .Mexico (Fauchald, 1972a) in similar depths and lo the bathyal fauna of some of the open trenches and basins off southern California. Characteristic of the deep-water polychaete fauna is the dominance of paraonids, spionids, lum- bnneiids, and onuphids. The ampharetids are usually more important than the lerehcllids; however, members of the terebellid subfamily POLYCIRRINAE are present in large nuiubers in deep water, but are usually poorly reported because of difficult taxonomic problems. This paper is no exception in this regard. I he sigalionids are usually more important than the polynoids; this is also the case in the present material, but both families are poorly represented, and no conclusive statements can be made. Three taxa are characteristic in many ways of deeper water but always represented by only one or a few species, include the sternaspids, fauveliopsids, and trichobranchids. These three families are represented by large numbers of specimens off Oregon and, despite the low species numbers, characterize the deep-water samples if abundance is considered. New genera are described in the families SCALlBRFCiMIDAE, FL.ABFI.IJGERIDAE, and AM- PH ARETIDAF. All three genera are strikingly different from previously described taxa. New species are described in the PARAONIDAE. COSSLRIDAE, SP10MÜAE, OPHELIIDAE PHVl.l ODOCIDAF HESIONIDAE, AMPHINOMIDAE, ONUPHIDAE and FALVEI.IOPSIÜAE. Keys to species within each family are given, but no family key has been constructed, since such a key was recently published (Fauchald, 1977). The keys are constructed solely to distinguish among species reported in the present study. They will not segregate species reported in the present study from other species, described or undescribed. In some instances, characters used in the keys may be generally valid at the generic level only, but are sufficient in the present context to distinguish species included. Most of this material and a series of additional samples were discussed by Hancock (1969). Since this document was unpublished, the authors ask that it not be quoted and that species mentioned in that document not be cited except after an examination of the material, since some incomplete identifications and a few erroneous identifications have been discovered.
ACKNOWLEDGEMENTS
The authors are grateful to Dr. Andrew J. Carey, Jr., Department of Oceanography, Oregon State University, for allowing us to study this collection, and to Dr. Bernard C. Abbott, Director of the Allan Hancock Foundation, lor space, access to the collections and the library, and lor other material a.ssistance. Mrs. JoAnne Woodcock expertly typed the manuscript and organized the literature citations and station list. Deep-Water Polychaetes 5
ORDER ORBINIIDA
FAMILY ORBINIIDAE HARTMAN, 1942
KEY TO SPECIES la. Prostoniium anteriorly truncate or rounded Naineris unrinata b. Prostomium anteriorly sharply pointed 2 2a. Thoracic setae all simple capillaries ^ b. Thoracic setae of at least two different kinds 4 3a. Thorax with 1 1 setigers; first branchiae abdominal in position Haploscoloplos kerguelensis h. Thorax with 16 setigers; first branchiae on one of the last thoracic seúgers Haplos•loplos elongatus 4a. First three thoracic setigers with brush-tipped setae Califia mexicana b. Brush-tipped setae ab.sent .' • "." 5a. Last thoracic setigers with a few thick, harpoon-shaped neurosetae in addition to the capillaries Phylo nudus b. Last thoracic setigers without harpoon-shaped neurosetae Scoloplos (Leodamas), near mazatlanensis.
Califia mexicana Fauchald, 1972
Califia mexicana Fauchald, 1972a, pp. 164-166, pi. 33, figs. a-e. fípforrf^; AD-6, NAD-17 (2): AD-74, NAD-I4 (1). Remarks: Members of the genus Califia were reviewed by Fauchald (1972a). C. mexicana has branchiae limited to about 10 setigers. The first setigers have spherical postsetal lobes and lack simple dentate setae. Occurrence: Bathyal depths off western Mexico.
Haploscoloplos elongatus (Johnson, 1901)
Scoloplos elongata Johnson, 1901, pp. 412-413, pi. 10, figs. 105-110. Haploscoloplos elongatus: Hartman, 1969, pp. 19-20, 5 figs.; Hartman and Reish, 1950, p. 26; Fauchald, 1972a, p. 166; Carey, 1972, p. 438. Record: AD-41, NÂD-21 (1). Remarks: The present specimen agrees with H. elongatus as defined by Hartman ( 1969). The first pair of branchiae is on setiger 14. Occurrence: C^ommon in shallow water from Alaska to southern California.
Haploscoloplos kerguelensis (Mclntosh, 1885)
Scoloplos kerguelensis Mclntosh 1885, pp. 355-356, pi. 43, figs. 6-8, pi. 22A, fig. 19. Haploscoloplos kerguelensis: Fauchald, 1972a, pp. 166-167, pi. 34, figs. a-b. Records: AD-7, NAD-16 (2); AD-149, NAD-15 (1). Remarks: The piesent specimens have about 11 thoracic setigers and the first branchiae are on one of the abdominal setigers. They thus agree with H. kerguelensis. Occurrence: Appears to be widespread in deep water (Fauchald, 1972a).
Naineris uncinata Hartman, 1957
Naineris uncinata Hartman, 1957, pp. 301-302, pi. 38, figs. 1-8; Hartman, 1969, pp. 31-32, 6 figs. Record: AD-97, NAD-4 (4). Remarks: The present specimens have double neuropodial postsetal lobes in posterior thoracical setigers. All other species recorded from the Americas have exclusively simple postsetal lobes on the thorax. Occurrence: Alaska to southern California. Intertidal in the northern part of the range and submerged in lower latitudes.
Phylo nudus (Moore, 1911)
Aricia nuda Moore, 1911, pp. 311-315, pi. 21, figs. 172-176. Phylo nudus: Hartman, 1969, pp. 39-40, 4 figs.; Carey, 1972, p. 439. 6 Allan Hancock Foundation Monograph No. 11
Records: AD-56, NAÜ-25 (1): AD-148, NAD-I2 (1). Remarks: Members of the genus Phylo have thick acicular spines in posterior thoracic setigers. P. nudus differs from congeners from the Americas in the lack of a ventral fringe usually present on these same setigers. Occurrence: Bathyal depths off southern California.
Scoloplos (Leodamas), near mazatlanensis Fauchald, 1972
Scoloplns (Leodamas) mazatlancnsis Fauchald, \972d, pp. 169-171 pi 3.5 figs a-c Record: AD-7, NAD-16 (2). Remarks: The present specimen differs from Scoloplos (Leodamas) mnzatlanensis as originally described in havmg 14 rather than 15, thoracic setigers. It does not oihcr\\'i.se differ from thai species as described.
FAMILY PARAONIDAE CERUTTI 1909*
KEY TOSPF.C:iFS
la. Median antenna absent 2 b. Median antenna present 4 2a. Modified setae present in postbranchial neuropodia Tauberia graccilis b. Modified setae absent 3 3a. Branchiae absent Paraonella ahranchiata b. Five pairs of branchiae present Paraonella cedroensis 4a. Modified setae absent 5 b. Modified setae present g 5a. Antenna barely reaches the tip of the prostomium 6 b. Antenna reaches at least the .second seliger 7 6a. Maximally, 20 pairs of branchiae present Aedicira antennata b. At least 40 pairs of branchiae present Aedicira pacifica 7a. Postbranchial postsetal lobes long and thread-like Aedicira longicirrata h. Postbranchial postsetal lobes short and stubby Aedicira oregonensis 8a. Modified setae notopodial Cirrophorus sp. b. Modified setae neuropodial 9 9a. Notopodial and neuropodial capillary setae of similar thickness in postbranchial setigers Acesta lopezi h. Notopodial capillary setae distinctly thicker than the neuropodial ones in postbranchial .setigers 10 10a. Antenna branched Allia ramosa h. Antenna unbranched 11 1 la. Modified setae distally aristate Allia hartmani b. Modified setae without aristae Allia crassicapitis
Acesta lopezi Berkeley and Berkeley, 1956
Aricidea lopezi Berkeley and Berkeley, 1956, p. 542, figs. 1-3; Hartman, 1969, pp. 59-60, 3 figs. Aricidea (Acesta) lopezi lopezi: Strelzov, 1973, pp. 102-104, fig. 44. Aricidea ? lopezi: Fauchald, 1972a, p. 179. Records: AD-11, NAD-15 (1); AD-17, NAD-17 (3); PAD-44, NAD-22 (1); AD-65, NAD-21 (1). Remarks: The concept of the species is accepted as defined by Strelzov (1972); the described subspecies, rubra (Hartman 1963) and rosea Reish (1968), are here considered separate species; thus the subspecific de.signation used by Strelzov has been eliminated. Occurrence: May be widespread, but has been confused with similar species. Western Mexico and southern California in bathyal depths and in shelf depths in western Canada.
*The many .subgenem described in this family try Stelinv (1973) are here considered as distinct genera (see also Fauchald, 1977). Deep-Water Polychaetes
Aedicira antennata (Annenkova, 1934)
Aricidea antennata Annenkova, 1934, p. 658, figs. 2, 36. Aedicira antennata: Fauchald, 1972a, pp. 175-176. Aricidea (Allia) quadrilnhata: Strelzov, 1973, pp. 88-91, figs. 13 and 37 (partim). Aricidea uschakov'i: Carey, 1972, p. 439. Records: AD-6, \AD-17 (1); AD-9, NAD-21 (2); AD-UO, NAD-21 (4). Remarks: Strelzov (1973) combined a series oí'species that have comparativelv long, slender antennae and lack modified .setae. In ail of these forms, the neurosetae in poslbranchial setigcrs arc somewhat more sharply tapered than those in more anterior setigcrs and those in the notopodia. The relative lengths of the antennae vary, as do the development of the postsetal lobes and the number of pairs of branchiae. Without more precise data, the propo.sed combinations are unacceptable. The present specimens agree with the specimens designated Aedicira antennata from western Mexico. Occurrence: Cold-water areas of the northern Pacific as far south as western Mexico in slope depths.
Aedicira longicirrata Fauchald, 1972
Aedicira longicirrata Fauchald, 1972a, pp. 176-177, pi. 36, fig. a. Records: AD-6, NAD-17 (1); ?AD-7, NAD-16 (2). Remarks: Aedicira longicirrata has very long, cirriform postselal lobes in postbranchial setigcrs; the long, sofi postbranchial neurosetae are gently tapering. The specimens from .\fí-7 have the long postsetal lobes, but all postbranchial setae have been broken. Occurrence: Bathyal depths off western Mexico.
Aedicira oregonensis, new species
Records: AD-6, NAD-17 (I ) (pi. 1, fig. 1); AD-16, NAr)-13 (1); AD-17, NAD-17 (1); AD-18, NAD-23 (!)• AD-89, NAD-22A (I): AD-139, NAD-24 (26, HOLOTYPK, Poly 1162, PARA IPKS Poly 1163); AD-1.50 NAD-26 (10). Descriptions: The holotype is an incomplete specimen with 80 setigers; it is 9 mm long and 0.5 mm wide, excluding setae. Anterior .segments are narow; postbranchial segments are approximately as long as they are wide. The body is evenly tapered posterior to the branchial region. The specimen is yellow and has no color patterns. The prostomium (Fig. a) is broadly rounded; the antenna is clavata and sits on a slightly elevated base. Parapodia consist of low welts in prebranchial and branchial regions; ihev are indistinct, aside from the emerging selai bunflles, in postbranchial setigers. Postsetal lobes are present from setiger 2 to about seliger 15 or 16; the lobes increase in size up to setiger 8 or 9 and thereafter decrease. They are narrow and digitate where best developed. Pairs of branchiae are pre.sent on setigers 6 to 15; each branchia is thick; some have a tapering tip, but most are bluntly rounded distally. All setae are moderately long, slender capillaries in both rami. Aedicira oregonensis is characterized by the low number of pairs of branchiae. The only other member of the genus (see Fauchald, 1972a for a review) with less than 20 pairs of branchiae is A. helgicae (Fauvel, 1936, pp. 29-31, fig. 3), which may be a Paraonis species rather than oï Aedicira because the median antenna appears to be absent (Strelzov, 1973, pp. 68-69). Occurrence: Bathyal areas off central Oregon, hence the specific name. Detailed data for the type locality can be found in the Station List.
Aedicira pacifica (Hartman, 1944)
Aricidea pacifica Hartman, 1944b, pp. 316-317, pi. 27, figs. 8-9. Aricidea pacifica: Hartman, 1969, pp. 53-54, 3 figures. Aricidea (Aedicira) pacifica: Strelzov, 1973, pp. 66-68, pi. 17, fig. 3; text fig. 25. Records: AD-7, NAD-16 (2); AD-17, NAD-17 (7); AD-65; NAD-21 (1); AD-86, NAD-21 (2); AD-110 NAD-21 (1); AD-139, NAD 24 (5); AD-149, NAD-15 (4); AD-154, NAD-26 (10)
\ f^ Allan Hancock Foundation Monograph No. 11
Remarks: Ardirim parifica has up to 60 pairs of hranchiae. The antenna is long and slender. The postsetal lobes are long and slender, at least through the branchial region. The poslbranchial neiirosetae are abruptly tapered, often with a distinct bend. Ocnirrenre: Intertidal areas off southern California and shelf depths in the northwest Pacific Ocean.
Allia crassicapitis (Fauchald, 1972), new combinadon
Ariridea crassirapitis Fauchald, 1972a, pp. 177-179, pi. 37, figs. g-h. Records: AD-6, NAD-17 (2); An-9, NAD-21 (2); AD-19, NAD-22 (2); AD-33, NAD-21 (2); AD-41, NAD-21 (2); AD-65, NAD-21 (1): AD-86, NAI)-21 (2); AD-110, NAD-21 (2); AD-139, NAD-24 (4); AD-15(), NAD-26 (1). Remarks: The present species belongs to the genus/l//;« in that the neuropodial capillary setae are thicker than the notopodial ones in postbranchial setigers. The modified setae lack aristae. The present specimens agree well with the species as originally defined. Occurrence: Western Mexico in bathyal and abyssal depths.
Allia hartmani (Strelzov, 1968)
Aedicira hartmani Strelzov. 1968, pp. 80-81, fig. 3a-d. Allta hartmani: Strelzov, 1973, pp. 80-81, pi. 17, fig. 5. text fig. 33. Record: AD-13, NAD-18 (1). Remarks: The neurosetae are slightly less distinctly aristate than as described (Strelzov, 1973). Occurrence: Shelf depths in the Barents Sea.
Allia ramosa (Annenkova, 1934)
Ariciden ramosa Aiuienkova, 1934, pp. 657-658, fig. 3a. Ancidea (Allia) ramosa: Strelzov, 1973, pp. 82-83, fig. 34. Aedicira ramosa: Hartman, 1969, pp. 55-56, 1 fig. Records: AD-13, NAD-18 (1); AD-141, NAD-llB (1); AD-149, NAD-15 (1). Occurrence: Sea of Japan to Southern California in bathyal depths.
Cirrophoru^, species indeterminable
ßmW.- AD-150, NAD-26 (1). Remarks: The present specimen is too poorly preserved to allow closer identification.
Paraonella abranchiata, new species (Plate 1, Figs, b-c)
Records: AD-7, NAD-16 (3); AD-10, NAD-26 (2. HOLOTYPE, Poly 1164, PARAfYPE, Poly 1165); AD-1.^4, NAD-26 (3). Description: The holotype is a complete specimen, with approximately 70 setigers; it is 5 mm long and 0.25 mm wide, excluding setae. The body is cylindrical, yellowish, and lacks color patterns. The prostomium (fig. b) is triangular in outline, with a raised dorsomedian diamond-shaped area; it is slightlv concave ventrally. All anterior setigers are similar; each is biramous, and a distinct digitate notopodial postsetal lobe is present. These lobes increase slowly in length posteriorly and, near the far posterior end, are nearly as long as the body is wide. Branchiae are absent, hence the specific name. The pygidiiun (fig. c) is a swollen ring with five distinct anal cirri. One pair is on either side of the anal opening, distal and slightly dorsal in relation to the rest of the animal. A single, midventral cirus is present, and basallv on the ventral side is a pair of rather short, slender cirri. All cirri are digitate or slighdy clávate. All setae are long, slender capillaries. Paraonella abranchiata agrees with Paraonella Strelzov (1973, p. 146) in lacking an antenna and modified setae in either ramus. It differs from all members of the genus in also lacking branchiae. The pygidial appendages resembled those of some species of Cirrophorus (see Strelzov, 1973, figs. 49 and 51). Occurrence: Bathyal depths off central Oregon; data for the type locality can be found in the Station List. Deep-Water Polychaetes ^
Paraonclla cedroensis (Fauchald, 1972), new conibiiialion
Paraonides cedwciisis Fauchald. 1972a. pp. 181-182. Record: AD-17. S\Y)-\1 (1). Rnnarks: Tlii.s spfcios belongs lo the genusParaonclla (Slrelzov. 1973, p. 14(i). It was ditteientiated from similar species bv Fauchald (1972a). It most closely resembles the genotype of Paraonclla, P. nórdica (Strelzov. 1968, pp. 7.5-76. fig. 1) in that both have lew pairs of branchiae liniited to anterior setigers. The two can be separated bv the long ¡)ostbraiuhial posisetal lobes in P. ci'drocn.sis; these lobes are the same length on both pre- and postbranchial setigers in P. nórdica. Furthermore, P. nórdica has six pairs of branchiae, starting on setiger 5; /••. ccdroensis has live pairs, starting on seliger 4. Occurrence: Western Mexico in bathyal depths.
Tauheria gracilis (Tauber, 1879)
Aonides gracilis Vduber 1879. p. 11.5. Paraonis granlú oculata: Carev. 1972, p. 439. Paraonis gracilis: Fauchald. 1972a. p. 183. Tauberia gracilis: Strelzov. 1973, pp. 127-133. figs. 54-57. Records: AD-6. NAD-17 (4); AD-9. \AI)-21 (5); AD-13. NAD-18 (1); AD-16, NAD-13 (!)• AD-18 NAI)-23 (1); Al)-41, NAD-24 (1); AD-43, \AD-22 (1): AD-141. \AD-1 18 (1); An-148 NAD-T' (1)' AD-149, NAD-15 (157); AD-1,54, NAD-26 (2). Remarks: Tauberia gracilis is the most abundant paraonid in the present material. I he initial appearance and the number of pairs of branchiae are somew hat variable. Smaller specimens have branchiae starting on setiger 5 and usualK have about six pairs. Larger specimens have the first branchiae on either setiger 6 or 7 and maximally have 15 pairs; the number is usuallv around 12 pairs. This variability is well within the boundaries defined previouslv for this species. Species of this genus were well defined by Strelzov (1973). Occurrence: Apparently nearly worldwide in colder waters.
FAMILY COSSURIDAE DAY, 1963
KEY TO SPECIES
la. Tentacle inserted on setiger 4 Cossura módica b. I entacle inserted on setiger 3 2 2a. Body darkly pigniented; enlarged setae in both notopodial and neuropodial fascicles Cossura hrunnea b. Body pale; enlarged setae in neuropodial fascicles only Cossura rostrata
Cossura hrunnea Fauchald, 1972
Cossura hrunnea Fauchald, 1972a, pp. 208-210. pi. 41. figs. a-e. Records: AD-7, NAD-16 (1); AD-9, NAD-21 (1): AD-65. \AD-21 (1); AD-1 19, NAD-22 (1)- AD-141 NAD-118 (4); AD-149, NAD-15 (9). Remarks: Cossura hrunnea can be differentiated from the very similar C;. candida Hartman (1955, pp. 44-45, pi. 1, figs. 1-5) by a dark brown pigmentation on most of its segirients. C. candida is pale colored. Enlarged setae are present in both fascicles in C. hrunnea, and are limited to the neuropodial fascicles in C. candida. Occurrence: Bathyal depths off western Mexico. 10 Allan Hancock Foundation Monograph No. 11
Cossura módica, new species (Plate 1, Fig. d)
Records: AD-17, NAD-17 (4); AD-32, NAD-19 (1); AD-141, NAD-118 (35, HOLOTYFE AHFPoly 1158, PARATYPES AHF Poly 0000). Description: The type is an incomplete specimen with 35 setigers. It is 6 mm long and 0.5 mm wide, excluding setae. The body is cylindrical, anteriorly tapering, and white. The prostomium (Fig. d) is conical and approximately as long as it is wide. The two peristomial segments are equal in length; each is approximately as long as the prostomium. The first setiger has a single ramus; all other parapodia are biramous. Each notopodium has two distinct fascicles of setae emerging at different angles to the long axis of the parapodium; all are slender, evenly tapering capillaries. The longer setae are in a bundle of two or three; the shorter, in a bundle of about 20 in anterior setigers; the number of setae in each fascicle decreases posteriorly. Each neuropodium also has two fascicles of setae. The smaller bundle has two or three evenly tapering setae closely resembling those in the notopodium. In the first 15 to 20 setigers, the larger fascicle has thick, abruptly tapering, marginally limbate setae. These thick setae are replaced posteriorly by slender capillar- ies. All capillary setae from both rami are distally pilose. The tentacle is attached on setiger 4; when complete, it reaches to setiger 20. The genus Cossura was reviewed by Fauchald ( 1972a). The present species belongs to the group that has two complete peristomial segments. The only other species in this group that has the tentacle inserted posterior to setiger 3 isC. alba Hartman (1967, p. 119), which has smooth hmbate noto- and neurosetae in all setigers. The present species has both limbate and capillary setae; in addition, all of the capillary setae are distally pilose. Occurrence: Off the central Oregon coast in bathyal depths. Detailed data can be found m the Station List. Etymology: The name refers to the unremarkable appearance of this worm {modicus Ladn = ordinary, commonplace). Cossura rostrata Fauchald, 1972
Cossura rostrata Fauchald, 1972a, pp. 211-212, pi. 41, figs, f-h, pi. 42, fig. a. Records: AD-7, NAD-17 (6); AD-9, NAD-21 (5); AD-32, NAD-19 (2); AD-33, NAD-21 (3); AD-41, NAD-21 (3); AD-44, NAD-22 (1); AD-65, NAD-21 (10); AD-86, NAD-21 (7); AD-87, NAD-21 (4); AD-110, NAD-21 (3); AD-119, NAD-22 (9); AD-139, NAD-24 (6); AD-141, NAD-118 (40); AD-149, NAD-15 (24); PAD-150, NAD-26 (2 juveniles); AD-154, NAD-26 (2). Remarks: C. rostrata has two peristomial segments; the tentacle is inserted on the third setiger, and the first setiger has a neuropodial fascicle of setae only. The neuropodial fascicles have thick, abruptly tapering setae in anterior setigers; such setae are missing from the notopodial fascicles. Occurrence: Western Mexico in bathyal depths. This variability is well within the boundaries defined previously for this species. Species of this genus were well defined by Strelzov (1973). Occurrence: Apparendy nearly worldwide in colder waters.
Family Chaetopteridae Malmgren, 1867 Phyllochaetopterus limicolus Hartman, 1960.
PhyUochaetopterus limicolus Hartman 1960, pp. 120-122, pi. 10, figs. 3-5; Hartman, 1969, pp. 215-216, 3
^Records: AD-41, NAD-21 (1); AD-89, NAD-22A(1 and tubes); AD-119, NAD-22 (1); AD-141, NAD-1 IB (1); PAD-149, NAD-15 (1, postlarva). Remarks: Phyllochaetopterus limicolus can most easily be separated from Spiochaetopterus costarum (Claparède, 1870; see Hartman, 1969, pp. 219-220, 6 figs.), which is also common in deep-water coastal samples, by the number of segments in the midbody region. P. limicolus has only two setigers in the intermediate region; S. costarum has at least 30. The chisel-shaped seta of sefiger 4 is distally smooth in P. limicolus and crenulate in S. costarum. These characters will not separate P. limicolus from other species of Phyllochaetopterus. Occurrence: Deep shelf and bathyal depths off southern California. Deep-Water Polychaetes ,,
FAMILY SPIONIDAE, GRUBE, 1850
KEY TO SPECIES
la. Fifth setiger with modified setae Polydora hrachycephala b. Fifth setiger without modified setae 2 2a. Branchiae present 3 b. Branchiae absent 6 3a. Branchiae present on a large number of segments Scolelepis foliosa b. Branchiae limited to about 10 anterior setigers 4 4a. All branchiae cirriform 5 b. First and last pair of branchiae bipinnate I'rionospio anuncata 5a. Four pairs of branchiae; neuropodia! hooks first present from setiger 22 Minuspio minor b. At least six pairs of branchiae; neuropodial hooks first present from setiger 12 or 13 Minuspio cirrifera 6a. First setiger with capillary setae only Spiophanella pallida b. First setiger with large curved hooks in addition to the capillary setae 7 7a. Frontal horns long and slender, eyes absent Spiophanes anoculata b. Frontal horns short or absent; eyes present 8 8a. Median and posterior notopodial lobes folióse Spiophanes berkeleyorum b. Median and posterior notopodial lobes cylindrical or threadlike 9 9a. Hooded hooks present from about setiger 25; each hook quadridentate . . . .Spiophanes fimbriata b. Hooded hooks first present from sedger 15; each hook tridentate Spiophanes kroyeri
Minuspio cirrifera (Wirén, 1883)
Prionospio (?) cirrifera Wirén, 1883, pp. 409-410. Minuspio cirrifera: Foster, 1971, pp. 108-112, figs. 262-275. Records: AD-9, NAD-21 (11); AD-43, NAD-22 (1). Remarks: Foster's (1971) concept of Minuspio cirrifera appears to be somewhat broader than is acceptable. The present specimens have six pairs of cirriform branchiae; hooded hooks are present in the neuropodia from setiger 12 or 13, and each is distally multidentate. The branchiae are digitate rather than filiform. Occurrence: May be widely dispersed, but appears to have been confused wilh several similar species.
Minuspio minor, new species (Plate 2, Figs, c-d)
Record: AD-13, NAD-18 (2, HOLOTYPE, Poly 1152, PARATYPE Poly 1153). Description: The holotype is an incomplete specimen with 29 setigers. It is 5 mm long and 0.3 mm wide, excluding setae. The body is cylindrical and slightly inflated anteriorly, with the anterior end appearing to be truncately tapering toward the ventral side. The prostoniium (Fig. c) is an elongated flattened keel, with the transverse anterior end as broad as the width of the specimen. The peristomium has a pair of flattened, distally rounded wings which are held erect over the posterior part of the prostomium. The first parapodium is incomplete, with the notopodium missing. The neuropodial postsetal lobe is flattened and directed dorsally; it covers the base of the peristomial wings. All other parapodia are biramous. Anterior notopodia have triangular, elongated postsetal lobes; more posteriorly, these lobes become lower and distally rounded or truncate. They are distinct and folióse in all setigers. Neuropodial postsetal lobes are similar in all setigers; each is folióse and distally truncate. Four pairs of cylindrical branchiae are present on setigers 2 to 5. Each is slender and distally tapering. The anteriormost pair is slightly shorter than the others; all other pairs are similar in length. Anterior setigers have only slender capillary setae; pilose genital spines are present ventrally in each neuropodial fascicle from setiger 8 or 9. Neuropodial hooks (Fig. d) are first present from setiger 22; notopodial hooks are absent from both specimens. Each hook is double-hooded and distally quadriden- tate, with a thick proximal fang and three slender teeth distally. 12 Allan Hancock Foundation Monograph No. 11
Minuspio minor resembles M. chilensis (Hartmann-Schröder, 1962) in that both have reduced first setigers without notopodia. All other species of the genus have complete first setigers. M. chilensis has eyes, while M. minor does not. M. chilensis has six pairs of branchiae, and AÍ. minor has four. Lateral wings are indistinct on the peristomium in M. chilensis and distinct in M. minor. Hooks are first present from setiger 13 inM. chilensis, later in larger specimens; similar-sized specimens of M. minor have hooks from setiger 22. Both species have quadridentate hooks. Occurrence: Bathyal depths off central Oregon. Detailed locality data can be found in the Station List.
Minuspio, species A
Records: AD-9, NAD-21 (16); AD-13, NAD-18 (2); AD-17, NAD-17 (3); AD-33, NAD-21 (1); AD-65, NAD-21 (5). Remarks: All of these specimens belong to the same species. All lack branchiae, and the branchial scars are not sufficiently distinct to allow complete identification. They are referred loMinuspio since they resemble M. cirrifera in most external characteristics. However, the hooded hooks are bidentate rather than multidentate as in M. cirrifera; the first neuropodial hooks are present from about setiger 15 or 16 in most specimens.
Polydora brachycephala Hartman, 1936
Polydora brachycephala Hartman, 1936, pp. 48-49, figs. 3-5; Hartman, 1969, pp. 129-130, 3 figs. Record: AD-68, NAD-6 (1). Remarks: This single specimen agrees with Polydora brachycephala in that it has nearly straight, distally bidentate hooded hooks. The prostomium is deeply bifid and is hooded by large wings developed from the peristomium. The shape of the modified setae of setiger 5 agrees with the illustration given by Hartman (1969). Occurrence: Intertidal and shallow subtidal areas in central California.
Prionospio anuncata Fauchald, 1972
Prionospio (Prionospio) anuncata Fauchald, 1972a, pp. 193-194. pi. 39, figs. a-e. Records: ?AD-6, NAD-17 (3); AD-16, NAD-13 (2); AD-17, NAD-17 (2); AD-86, NAD-21 (1); AD-141, NAD-llB (9); AD-149, NAD-21 (7). Remarks: Prionospio anuncata was separated from similar species which also have four pairs of branchiae, the first and last bipinnate and the second and third smooth, on the late first occurrence of the hooded hooks in the neuropodia. Ihe first hooks are present in setigers 25 to 27 in the present specimens. Occurrence: Western Mexico in bathyal depths.
Scolelepis foliosa (Audouin and Milne Edwards, 1833)
Aonisfoliosa Audouin and Milne Edwards, 1833, pp. 402-403, pi. 8, figs. 9-13. Nerine foliosa: Fauvel, 1927, pp. 34-35, fig. lla-f. Scolelepis foliosa: Pettibone, 1963, p. 92 (in part); Foster, 1971, p. 65 (in part). fiicoiY/.- AD-38, NAD-11 (1). Remarks: S. foliosa occidentalis Hartman (1961, p. 90) appears to be separable from the main form of this species as indicated by Hartman ( 1961 ). Thus, the synonymy suggested by Pettibone and Foster is accepted only in part. Occurrence: Probably widespread, but may have been confused with similar species.
Spiophanella, new genus
Spionid species without branchiae and without enlarged, curved hooks in the first setigers. Anterior parapodia with well-developed noto- and neuropodial postsetal lobes; median and posterior setigers with reduced lobes. Notopodia with capillary setae only; neuropodia in median and posterior setigers with hooded multidentate hooks. Genotype: Spiophanella pallida (Hartman, 1960, as Spiophanes pallidus, pp. 118-119, pi. 10, figs. 1-2). Deep-Watcr Polychaetes jj
Spiophmiella agrees with Spiophanes in tile \¿Lck of branchiae and in the distribution of hooded hooks; it differs most marl;ediy in tlie lacle of enlarged hoolis in the first setigers.
Spiophanella pallida (Hartman, 1960)
Spiophanes pallidus Hartman, 1960, pp. 118-119, pi. 10, figs. 1-2; Hartman, 1969, pp. 187-188 2 figs- Fauchald, 1972a, pp. 199-200. ^ Records: AD-6, NAD-17 (1); AD-33, NAD-21 (1); AD-55, NAD-25 (1); AD-150, NAD-26 (1). Remarks: Fauchald (1972a, p. 199) remarked that the characteristic Spiophanes hooks could not be recognized in specimens from western Mexico. This was thought to be due to the poor condition of the available specimens. An examination of the type material and of additional material from southern California has shown that hooks are consistently absent in this species. Petdbone (1962, p. 85) suggested that this species might be a member oi Prionospio. The lack of branchiae precludes that, but, as suggested by Petdbone, the species definitely does not belong to Spiophanes. Occurrence: Southern California and western Mexico in bathyal depths.
Spiophanella anoculata Hartman, 1960
Spiophanes anoculata Hartman, 1960, p. 118; Hartman, 1969, pp. 179-180 2 figs • Fauchald 1972a n 198. , ,p. Records: AD-6, NAD-17 (1); AD-9, NAD-21 (6); AD-17, NAD-17 (1); AD-41, NAD-21 (2)- AD-139 NAD-24 (4); AD-149. NAD-15 (1); AD-150, NAD-26 (26); AD-154, NAD-26 (24). Remarks: Spiophanes anoculata has long, slender frontal horns and has reduced notopodia in the first setiger. The neuropodial hooded hooks are bidentate. Occurrence: Slope depths off southern California and western Mexico.
Spiophanes berkeleyorum Pettibone, 1962
Spiophanes berkeleyorum Pettibone, 1962, pp. 78-83, figs. 1-4. Record: AD-65, NAD-21 (1). Remarks: Spiophanes berkeleyorum closely resembles S. kroyeri. The posterior and median postsetal notopodial lobes are flat and folióse in .S'. berkeleyorum and are digitate in S. kroyeri. Occurrence: Shelf and bathyal depths off Wasfiington and western Canada.
Spiophanes jimbriata Moore, 1923
Spiophanesfimbriata Moore, 1923, pp. 179-182; Hartman, 1969, pp. 183-184, 4 figs; Fauchald 1972a p 199. Record: AD-148, NAD-12<1). Remarks: Spiophanes fimbriata resembles S. kroyeri closely, but can be separated on the basis of the quadridentate, rather than tridentate, hook.s and by the fact that these hooks first occur from setiger 15 in S. kroyeri and from about setiger 25 in S.fimhrata. The two species are frequently considered synonymous (see Pettibone, 1962, p. 85). Occurrence: Bathyal depths off central and southern California.
Spiophanes kroyeri Grube, 1860
Spiophanes kroyeri Grube, 1860, pp. 88-89, pi. 5, fig. 1; Söderström, 1920, pp. 240-243, figs. 150-152; Fauchald, 1972b, p. 99, fig. 4c-d. Records: AD-6, NAD-17 (1); AD-7, NAD-16 (1); AD-9, NAD-21 (4); AD-11, NAD-15 (1); AD-17 NAD-17 (4); AD-32, NAD-19 (2); AD-65, NAD-21 (1); AD-119, NAD-22 (1); AD-148, NAD-12 (1). Remarks: The present specimens are small and delicate compared to shallow-water specimens reported as this species as are specimens from Norway (Fauchald, 1972b). 5. kroyeri can be separated from 5. berkeleyorum by the shape of the notopodial postsetal lobes in median ancl posterior setigers. These are digitate in S. kroyeri and folióse in S. berkeleyorum. The hooks are quadridentate in S. fimbriata and are tridentate in S. kroyeri. Occurrence: May be widespread, but have been confused with similar species. 24 Allan Hancock Foundation Monograph No. 11
FAMILY CIRRATLUDAE CARUS, 1863
KEY TO SPECIES la. AU setae capillary j b. At least some spines present in addition to the capillary setae 3 2a. Anterior and posterior segments short and crowded; median setigers monilate Tharyx, near monilans b. All setigers shorter than wide; none monilate Tharyx, near multifilis 3a. Neuropodial spines present Irom the first setiger Chaetozone, near corona b. Neuropodial spines present ivom a median setiger Chaetozone setosa
Genera and species of the bipalpate cirratulids are currently under revision. The concepts indicated by the key above are inadequate to cover the number of taxa involved. The consservative approach appears to be best to avoid adding new names to the literature before the revisions have been completed.
Chaetozone, near corona Berkeley and Berkeley, 1941
Chaetozone spinosa var. corona Berkeley and Berkeley, 1941, pp. 45-46. Chaetozone corona: Hartman, 1969, p. 235-236, 3 figs. ß/?fori/.- AÜ-154, NAn-26 (7). Remarks: The present speciinens agree with C. corona in the shape of the anterior end and in the presence of neuropodial spines from the first setiger. However, the spines remain single in all setigers and do not form cinctures around the posterior end as in C. corona.
Chaetozone setosa Malmgren, 1867
Chaetozone setosa Malmgren, 1867, p. 206, pi. 15, fig. 84; - Hartman, 1969, pp. 241-242, 3 figs. Records: AD-6, NAD-17 (5); AD-13, NAD-18 (1); AD-74, NAD-14 (1); AÜ-148, NAD-12 (1); AD-149, NAD-15(1). Remarks: Chaetozone setosa has both notopodial and neuropodial spines present from a median setiger; the spines increase in number until they form cinctures around the posterior end. Notopodial spines start a lew setigers posterior to the neuropodial ones. The spines are interspersed with capillary setae even in far posterior setigers. Occurrence: The northern hemisphere, possibly cosmopolitan.
Cirratuius, species indeterminable
/íírorr/.- AD-7, NAD-16 (7). Remarks: The present specimens have tentacles arising from the same setiger as the first branchiae. The tentacular are cirri liinited to one segment, as is characteristic of the genus Cirratuius. The specimens are too incomplete to allow coitiplete identification.
Tharyx, near monilaris Hartman, 1960 (Plate 2, Figs, a-b)
Thamx monilans Hartman 1960, pp. 127-128, pi. 12, figs. 1-2; Hartman, 1969, pp. 261-262, 2 figs. Records: AD-7, NAD-16 (4); AD-9, NAD-21 (16); AD-16, NAD-13 (1); AD-17, NAD-17 (1); AD-18, NAD-23 (6); AI)-33, NAD-21 (12); AD-43, NAD-22 (1); AD-44, NAD-22 (4); AD-65, NAD-21 (15); AD-86, NAD-21 (38); AD-89, NAD-22A (6); AD-110, NAD-21 (4); AD-119, NAD-22 (30); AD-139, NAD-24 (29); AD-141, NAD-llB (394); AD-149, NAD-15 (14); AD-I50, NAD-26 (15); AD-154, NAD-26 (6). Remarks: Nearly all specimens listed above agree with the illustrations in Hartman (1969) and have serrated setae. Fhe specimen from .í\D-16 resembles the others closely in all characters, but the setae are smooth (fig. b). Both forms agree with T. monilaris in the structure of the anterior end (Fig. a) and in the relative proportions oí the segments. More than one species may be contained in the current material, and perhaps none of these belong to T. monilaris. Deep-Water Polychaetes . -
Thanx, near miiltifilis Moore, 1909
Thm-yx multifilis Moore, 1909, pp. 267-268, pi. 9, figs. 4.S; Hartman, 1969, pp. 263-264, 1 tig. Records: AD-6, NAD-17 (3); Al)-9, \A1)-21 (23); AD-13, \AD-18 (4); AI)-17, \AD-17 (9); AD-32, NAD-19 (1); AD-41, NAD-21 (18); AD-44, NAD-22 (1); AD-86, NAD-21 (10); AD-105, NAD-12 (3); AD-IIO, \AD-21 (10): AD-119, NAD-22 (7); AD-139, NAD-24 (3); AD-148, NAD-12 (2): AI)-149, NAI)-15 (1); AD-150, NAD-26 (6); AD-154, NAD-26 (3). Remarks: Tlmnx multifilis is generally rather large, with a highU arched dorsmn and long, smooth setae. The present specimens agree in that the setae are long, the segments are crowded in all seligers, and the animals are generally larger than those considered above as T., near momlaris. They differ from T. multifilis in that the dorsmn is less strongly arched than usual. More than one species mav he involved.
FAMILY CAPITEl.LIDAK GRUBE, 1862
KF.^• I O SPECIES
la. Thirteen thoracic setigers present; all have pointed setae only Dasybrauchus glabrus b. Eleven thoracic .setigers present; some mav have uncini 2 2a. All 11 thoracic setigers with pointed setae only 3 I). Posterior thoracic setigers with luicini at least in the neuropodia 5 3a. Notopodial tori fused in the mid-line in median and posterior setigers Nntomastus (Xotomastus) abyssalis b. Notopodial lori separate in all setigers 4 4a. Uncini with one major tang and a crest of teeth in a single row; nephridial pores limited to the thorax Xotomastus (CJistomastus) tenuis h. Uncini with one major tang and crests oí teeth in double rows; nephridial pores present on the abdomen as well as on the thora.x Xotomastus (Xotomastus) hnagnus 5a. Thoracic imcini limited to the neuropodia 6 b. Thoiacic uncini ¡^resent in both rami Heteromastus ?filiJormis 6a. First six neuropodia with pointed setae; first seliger with notopodia only . .Xeoheteromastus liueus b. First eight neuropodia with pointed setae only; first seliger biramous Xotomastus (Notomastus) precocis
Dasybrauchus glabras Moore, 1909
Das\tminchus glabras .Moore 1909, pp. 280-281, pi. 9, fig. .58; Hartman. 1969, pp. 371-372, 2 figs. Records: AD-44. NA1V22 (1); AD-141, NAD-lIB (2); AD-149, NAD-15 (2). Remarks: Dasybrauchus glahrus has 13 thoracic setigers; all have capillary setae only. All abdominal setigers have uncini. Branchiae are two- or three-lobed. Occurrence: Central and southern California in shelf and slope depths.
Heteromastus ffiliformis (Claparède, 1864)
Capitella filiformis Claparède, 1864, pp. 509-510, pi. 4, tig. 10. Heteromastus filifimnis: Hartman, 1969, pp. 377-378, 5 figs. Record: AD-89, NAD-22A (1). Remarks: The present specimen agree with Heteromastus in ihal the first five thoracic setigers have pointed setae and the last six have long-handled hooks. The species cannot be identified safely since most of the abdomen is missing. 16 Allan Hancock Foundation Monograph No. 11
Neoheteromustits linens Hartman, 1960
Neolictcromastns Hunts Hartman 1960, p. Iä8; Hartman, 1969, pp. 389-390. 2 figs; Fauchalcl, 1972a, p. 243. Records: AD-6, NAD-17 (4); AD-41, NAD-21 (1); AD-44, NAD-22 (1); AD-65, NAD-21 (1); AD-119, NAD-22 (1); AD-149, NAD-15 (3). Remarks: Xenheterowastiis linens has the IM st setiger incomplete with only the notopodia present. The first seven sctigers have pointed seiae; setiger 8 has nciuopodial imcini and notopodial pointed setae; the remaining three thoracic setigers have imcini in both rami. Fhe anterior part of the body lacks areolalion. Occurrence: Bathyal depths off western Mexico and southern California.
Notomastus (Clistomastus) tennis Moore, 1909
Notomastns tenais Moore, 1909, pp. '¿77-27H, pi. 9, fig. .55. Notomastus (ClistomastiL Notomastus (Notomastus) ahyssnlis Fauchald, 1972 Notomastus (Notomastus) ahyssalis Fauchald, 1972a, pp. 248-249, pi. 51, figs. d-g. Records: NAD-17 (2); AD-139, NAD-24 (9). Remarks: Notomastus (Notomastus) ahyssalis has 1 1 thoracic setigers, all with capillary setae. The abdominal notopodia approach each other and fuse medially in posterior abdominal setigers. L'ncini are present in all rami, but the tori are poorly developed and aie usually barely raised above the surface of the body in the posterior part of the abdomen. Occurrence: Western Mexico in bathyal depths. Notomastus (Notomastus) fmaffnus Hartman, 1947 '. Notomastjis magrrns Hartman 1947, pp. 412-415, pi. 50, figs. 1-6. Notomastus (Notomastns) mai^nns: Hartman, 1969, pp. 401-402, 6 figs. Records: AD-6, NAD-17 (1); Al)-7. NA1)-16 (2). Remarks: These specimens resemble N. (N.) magnus in the structure of the anterior end and in the details of the uncini. They are all posteriorly incomplete, however, so the presence and structureof the branchiae could not be determined. Notomastus (Notomastus) precocis Hartman, 1960 Notomastus precocis Hartman 1960, pp. 139-140. Notomastus (Notomastus) precocis: Hartman, 1969, pp. 403-404; Fauchald, 1972a, pp. 251-252. Records: AD-9, NAD-21 (1); AD-18, NAD-23 (I); AD-65, NAD-21 (1); AD-89, NAD-22A (1); AD-141, NAD-1 IB (12); AI)-148, NAD-12 (8); AD-149. NAD-15 (14). Remarks: Notomastus (Notomatm) precocis has pointed setae in all 11 thoracic notopodia, but uncini are piesent in the last three thoracic neuropodia. The abdominal notopodia approach each other, but remain distinct even in the hindmost setigers. Occurrence: Bathyal depths off southern California and western Mexico. Deep-Water Polychaetes ff¡. FAMILY MALDANIDAE MALMGREN, 1867 KEY TO SPECIES la. Anus dorsal 2 b. Anus terniiiial 6 2a. Anterior iieuropodia with rostrate hooks or setae absent in the anteriormosi neuropodia 3 b. Anterior neuropodia with single acicular spines 5 3a. First setiger with an anteriorly directed, quadrilobate collar Maldane monilata b. First setiger wiihoul a collar 4 4a. .\nal platjue with three fíncate anal cirri Asychis ramosus b. .Anal pla(|ue marginally weakly crenulate, often nearly smooth Maldane cristata 5a. Cephalic plaque present Clymaldane laevis b. C^ephalic placjue absent Notoproctus parifirus ba. A\ least iwo anierior .segments with collars; rostrate hooks in double rows . . . .Rhodine Intorquata b. Onlv one anterioi" segment with collar; rostrate books in single rows Xicomaclie lumbricaiLs Asychis ramosus I.evenstein, 1961 As\fliis ramosus I.evenstein, 1961, pp. 165-166, fig. 9a-e; Fauchald, 1972a, p. 258. Records: AD-9, \AD-21 (2); ?AD-43, \AD-22 (posterior end onlv): ?AD-65, NAI)-21 (posterior end onlv); AD-86. NAD-21 (1); An-1()5, NAD-12 (4); AD-llO, NAD-2Í (1); AD-148, NAD-12 (9); AD-154 NAn-26 (I). Ri'iiKirks: I'he proslomium is posleriorlv covered bv a deep hood, as in memlx-is ol ihe genus .V/í//C/í;>í/', but the median keel is short and poorly developed, as in members oí Asychis. Ihe posterior end has a deep fiuuiel bordered marginally by three usually branched anal cirri. Occurrence: Sea ol Okhotsk and off western Mexico in bathyal depths. Clymaldane laevis Fauchald, 1972 Chmaldane laevis Fauchald, 1972a, pp. 259-260, pi. 53, Figs. a-d. Records: AD-6, NAD-17 (3); AD-I7, NAD-I7 (2). Remarks: Clymaldane laevis has both cephalic and anal plaques. The anus is distinctly dorsal, making C. laevis a member of the subfamily Maldaninac. Fhe first four neuropodia have single thick spines. Other maldanins either lack neurosetae in the first setigers or have normal rostrate hooks. Occurrence: Deep-water areas ofl western Mexico. fLumhriclymene, species indeterminable Record: AD-32, NAD-19 (anterior fragment). Remarks: Fhe present specimen has the long, slender body and relatively short, somewhat expanded anierior end illustrated for Lumhriclymene lineus by Hartman (1969, p. 455), but it cannot be identified safely because the posterior end is missing. Maldane cristata Tie,\úwe\\, 1923 Maldane cristata IVeadwell, 1923, pp. 9-10, Figs. 5-8; Hartman, 1969, pp. 457-458, 4 figs.; Fauchald, 1972a, pp. 262-263. Records: AD-44, NAD-22 (I); AD-149, NAD-15 (1). Remarks: Maldane cristata has a deep pocket formed posteriorly on the proslomium. Fhe keel is high and distinct, and the anterior end completely lacks collars. Occurrence: Deep-water areas from Panama to southern (california. 18 Allan Hancock Foundation Monograph No. 11 Mahl/nie ninniluta Fauchald, 1972 Maldnne moiiilata Faiichakl, 1972a, pp. 263-265, pl. 54, figs. c-g. Records: Al)-6, XAü-17 (1); Al)-32, NAD-19 (:5). Remarks: Maldane monilata resembles M. sarsi Malnigren, 1865 (Hartman, 1969, pp. 461-462, 5 figs) in that the posterior rim of the prostomium barelv covers the prostomial keel and forms at best a shallow pocket. .V/. moil data has a vvcll-de\ eloped, cjuadiipartile anterioi' collar on the first setiger; collars are absent in A7. sarsi. Occurrence: Off western Mexico and Panama in bathval depths. Xicuiiiaclie liindiricalis (Fabriciiis, 1780) Sabella lumbricalis Fabricius, 1780, pp. 874-;í75. Nicomache lumbricalis: Hartman, 1969, pp. 465-466, 3 tigs. Record: AD-6, NAD-17 (8). Remarks: Xicomache lumbricalis has an anal funnel, but a cephalic plaíjue is absent. The first three neuropodia have single spines; the body is reddish brown, without color patterns in life. The species can be .separated from its more conmion Pacific congener, ;V. personata Johnson, 1901 (Hartman, 1969, pp. 467-468, 3 figs.), by the presence of two pre-anal asetigerous segments in the former; the latiei' has onl\ a single pre-anal asetigerous segment. N. personata is also anteriorly mottled with white |)igment. Occurrence: Widespread in the noiihern hemisphere in shelf and upper bathyal depths. Notoproctus pacijicus (Moore, 1906) Lumbriciymene pacifica Moore, 1906, pp. 246-248, pl. 12, figs. 40-42. Xotoproctus pacificas: Hailman, 1969, pp. 469-470, 3 figs. Record: AD-6, \AD-17 (37); AD-7, \AD-16 (14). Remarks: Notoprochis pacijicus builds a stiff tube with a yellowish lining plastered with large sand grains, foraminiferans, etc. The anterior end lacks of cephalic plaque; an anal platpie is present and marginally smooth, with two pie-anal asetigerous segments. Occurrence: From western (Canada and Alaska to southern California in shelf and bathyal depths. Rhodine hitorquata Moore, 1923 Rhodine bitorquata Moore, 1923, pp. 223-225, pl. 18, fig. 30; Hartman, 1969, pp. 483-484, 3 figs.; Fauchald, 1972a, pp. 267-268. Record: AD-148, NAD-12 (1). Remarks: R. bitorquata has rostrate hooks in two rows in most neuropodia and well-developed collars around at least two, often thiee, segments near the anterior end. The lubes, which are not covered with debris or sand grains, are smooth, usually copper-colored and ver\ brittle. Occurrence: From western Mexico to Canada from shelf to bathyal depths. FAMILY OPHELllDAE MALMGREN, 1867 KEY IC) SPECIES la. Body long and slender, with distinct ventral and lateral grooves 2 b. Body grub-shaped, without distinct grooves 3 2a. Anal cone and cirri absent Ophelinu paliida b. Long, cylindrical anal cone present; anal cirri absent Ophelinu sp. A. 3a. Maximally 25 segiers present 4 b. At least 27 setigcrs present 5 4a. Eighteen setigcrs present Travisia oregonensis b. Twentv-four or 25 setigcrs present Travisia brevis 5a. More than 30 setigers present Travisia, near giga'i b. Fwenty-seven setigers present Travisia Joetida Deep-Water Polychaetes 19 Ophelniü ¡mllidü (Hartman, 1960) Amniolnpaiic pallida Harlmaii, 1960, pp. 133-135. pl. 14, l]g. 3: Harlniaii, 1969, pp. 321-322, 1 Fig.; Faiichald, 1972a, pp. 235-236. Records: AD-42. \AD-21 (2); AD-141, NAD-llB (3). HciiKirks: O. píillida lacks anal cirri and an anal cone; branchiae are first present frotn setiger 2 and are absent in the last four crowded setigers. Occurrcurc: Bathyal deptiis oft southern California and western Mexico. Ophelina species A. Record: Al)-«6, \A1)-21 (1). Remarks: The present specimen is rather poorly preserved and cannot be described without additional material. It has a long, closed anal cvlinder which is nearh smooth distallv, and ]jerhaps genth lobed. Branchiae are present from seliger 3 and are present, as far as can be determined, on all subsequent setigers. The sjjecies does not agree with descriptions of any species from the Pacific Ocean. Trai'isia brevis Moore, 1923 Travista brevis Moore, 1923, pp. 220-221 ; I lariman, 1969. pj). 343-344. 1 flg.: Hartman and Reish, 1950, p. 37; Fauchald, 1972a, p. 237. Records: AI)-6, \AD-17 (3); AD-9, Ny\D-21 (2); AD-17, NAn-17 (2); AD-hS, .\AI)-23 (1); AI)-43, NAD-22 (1); AD-44. \AD-22 (1); AD-86, NAD-21 (1). Remarks: Travisia brevis can be separated from the othei species in the genus that are foiuid off central Oregon on the basis of the nimilier of setigers. Fully mature spec imeiis never have more than 24 to 25 setigers. I'he parapodial lappets are poorly developecl even in posterior setigers. I'bere are two asetigerous pre-anal segments. Occurrence: Alaska to western .Mexico in shelf and slope depths. Travisia Joetida Hartman, 1969 Travisia foetida Hartman. 1969, pp. 345-346. 3 figs.; Fauchald. 1972a. pp. 237-23«. pl. 49, fig. d. Records: AD-33, NAD-21 (2); AI)-65, NAI)-21 (1). Remarks: 'Travista foetida has 27 setigers; the first segment is smooth, rather than papillated. The nuchal organs are deeplv embedded in the first setiger and aie attached to the prostomium by two epidermal ridges. In other species of Travisia the.se organs are attached directly to the posterior margin of the prostomium. Occurrence: Western Mexico and southern California in shell and slope depths. Travisia, ne,xr taigas Hartman, 1938 Travisia gigas Hartman, 1938, pp. 103-105, figs. 46-48; Hartman, 1969, pp. 347-348, 3 figs. Record: Àb-96, \AD-4 (2). Remarks: Fhe present specimens agree with Travisia gigas in that they have about 40 setigers, the posterior ones bearing very large parapodial lobes. I'hey differ in that they are completely covered with pustules that are distinctly larger than those of shallow-water specimens of 7'. gigasIS. Travisia oregonensis, new species (Plate 2, Figs, e-f) Records: AD-7, NAD-16 (2 HOLOTYPE, Polv 1154, PARATYPE, Polv 1155); AD-149, NAD-I5 (1, PARATYPE, Poly OOOO). Description: 1 he holotv|)e is a complete specimen. 10 mm long and 3 imii wide excluding setae, it is spindle-shaped and consists of 18 setigers and five asetigerous posterior segments. A narrow peristomial ring is present. All parts of the body apart from the prostomium and pygidium, are covered with large tubercules. Fhe tubercules are distinctly larger in the posterior half of each segment in the anterior half of the bodv, and are in indistinct rows. 20 Allan Hancock Foundation Monograph No. 11 The prostomium (Fig. e) is a short, smooth cone with two posteriolateral extensions over the anterior part (){ the perisioiniuni. The pcristoniiuin is a very short ring, poorly deinarcaled from the first setiger. All setigers are hiramoiis, with slender, marginally dentate capillary setae in both rami. Supra- and subpodial swellings are present in the last three or four setigers, but are notdistinctly set off from the rest oí the segment. Anterior setigers, back to setiger 10, are vaguely two-ringed; all others consist of a single ring onlv. Branchiae are present on all but the first and last setigers so that in all, 16 pairs of branchiae are present. Each is thick, digitate, and distinctiv ringed. The pygidium (fig. f) has a short, cylindrical anal cone that is deeply furrowed dorsally. About five di.stincl furrows can be seen. The distal margin has five or six lobes. Travista oregonensis has fewer .setigers when sexually mature than any other species of Tnii'isia. fhe structure of the pvgidiiim differs tVqm that in any other species oiTrai'isia on the Pacific coast. The genus is inider revision by Dr. .Stuart L. Santos. Occurrence: Two areas off central Oregon in balhyal depths. Details of the type localities can be fbuiid in the Station fist. FAMILY SCALIBREGMIDAE MALMGREN, 1867 Mucibrcgma, new genus Scalibregmids with 1-shaped prostomium and one peristomial segment branchiae and furcate setae absent. Setae include acicular spines and long, slender, pliable capillary setae. Mucihregma resembles the scalibregmids mainly in the structure of the prostomium and the general lack of specializations of the anterior end. The family is rapidly becoming a catch-all for simpiv structured polychaetes (Fauchald, 1977). A revision by Kudenov and Blake (1978) indicates Ú\MMucibregma would tit in their system as a separate subgroup (Cl) under section II of the family. Mucihregma spinosa, new species (Plate 3, Figs, a-b) Record: AD-6, NAD-I7 (1, HOLOTYPE, Poly 1146). Description: The holotype, and only known specimen, is posteriorly incomplete; it consists of 17 setigers and is 8 mm long and 1.7 mm wide, excluding setae. The whole body was encased in a tough nuicus membrane attached anteriorly to large glands on the perislomium. fhe bodv is dorsoventrally flattened, and the parapodia are prominent in the posterior half of the fragment. The prostomium (Fig. a) is f-shaped, with the two frontolateral corners projecting above and distinctly set off from the rest. Ventrally, appears as a flattened triangular plate; dorsallv, it is set off from the peristomiutn by a distinct furrow, and the space between the two is marked by a small lens-shaped depression. The peristomium is very wide and strongh nuiscular. fhe mouth opening is anteriorly directed, with a widely expanded, scoop-shaped, smooth lower lip (Fig. 6). Both the dorsal and ventral sides of the peristomium are covered by a large mucus gland. Ventrally, the gland is interrupted by a smooth, ovate muscular pad on the lower part of the lower lip. Dorsallv, the gland is continued posteriorly as a girdle onto the first setiger. All parapodia are biramous. fhe notopodia are larger than the neuropodia in the first few setigers; farther posteriorly, both blanches are e<|ual in size. Noto- and neuropodia are similar in shape; each is a thick, distallv truncate lobe, and distinct pre- and postsetal lobes are ab.sent. There are no dorsal or ventral cirri in anv setiger. Setae include thick, distally blunt, simple spines in all parapodia; additionally, each noto- and neuropodium has one (maximally two) long, slender, very pliable, simple capillarv seta. Mucihregma spinos can be .separated from other scalibregmids as indicated for the genus. Occurrence: Off central Oregon in bathyal depths. Details of the type locality can be found in the Station List. Scalibregma inflatum Rathke, 1843 Scalibregma inßatum Rathke, 1843, p. 184, pi. 9, figs. 15-21; Hartman, 1969, pp. 313-314, 4 figs. Records: AD-6, NAD-I7 (5); AD-18, NAD-23 (1); AD-33, NAD-21 (1). Deep-Water Polychaetes 21 Rrmarhs: Scfilihregma inflntxim has an inflated anierior end with a T-shaped prostomium and a long, often ragged-appearing, posterior enci. Arborescent branchiae are present on four anterior setigers. Setae incUide capillaries and, basally in each fascicle, a series of short furcate spines. Thick acicular spines are absent. Orrurreiirt': The northern hemisphere, mainly in shallow water, but previous records from bathyal depths have been noted from California. FAMILY PHYLLODOCIDAE, WILLIAMS, 1851 KEY TO SPECIES : la. Anterior neuropodia with large, inllated setae; other neuropodia with slender, composite spinigei's Choetoparid careyi b. .\il neiuosetae slender, composite spinigcrs 2 2a. Two pairs ol tentacular cirri present Liiffia ahyssirola h. Four pairs ol tentacular cirri present 3 3a. Vential tentacular cirri strongly llattened Ftcrocinus iiniijiniai b. Ventral tentacular cirri cylindrical or clávate 4 4a. I'rostomium with four antennae Frotomystidcs orcidentalis b. Prostoiniiini with five antennae .^ 5a. Parapodia iniiiamoirs F.umida, \\fMJu\ign(i b. Parapodia biramous Austropliyllum exisiliuni /I•Xitstrophylliun I'xsiliiim Fauchald, 1972 Aiistniplnilinn <-xsiliiun Fauchald, 1972a, pp. 47-4X, pi. 5, tigs. a-d. RcamLs: AD-6, \A1)-17 (1); AI)-7, NAD-16 (1); AD-9, \A1)-21 (1); Al)-18, NAD-23 (1); Al)-59, NAD-II (I). Remarks: UschakoN (1972, pp. 163-164) reviewed the genus ,4í/.SYíO/;/íV//í/W. As suggested bv Fauchald (1972a, p. 46). Eiildlid spliacrocephula Levenstein (1961) was referred lo this genus. Ihis species closely resembles/4. cxsilium. Fhe two can be separated as (ollows: the inferior frontal antennae are strongly clávate in A. exsilium, and slender in A. sphnerori'l/hala; the dorsuin bas three oi foui- dark transverse bands in A. exsilium, and is grey with a greenish tinge in A. .sphacrorephala. Ocninrnce: Batlivai depths off the tip of Baja California, Mexico. Chaetoparia careyi, new species (Plate 4, Figs, a-g) Rernnl: AD-119, \An-2 (1, HOl.OFYPF, Poly 1143). Description: The holotypc and only known specimen is posteriorly incomplete; it is 2.2 mm long and 0.25 mm wide, excluding setae. It consists of 18 .setigers. The anterior end is truncate, and the body tapers evenly posteriorlv. The parapodia are long compared to the width of the body, so the worm appears ragged. The color is light ochre, with scattered red dots at the bases of the parapodia, especially in posterior setigers. The prostomimn (Fig. a-b) is anteriorly truncate and is bent veniiad. Ibe Iroiital aniemiae are widely separated. Each is subulate and is about as long as the width of the prostomium; the ventral pair is longer than the dorsal pair. The first segment is asetigerous and is represented only by a single tentacular ciri us. 'Fhe cirrosi\k's have mosllv been lost, but since the cirrophores remain, the pattern can be determined. The second segment is setigerous and has a distinct acicular lobe and both dorsal antl \entrai cirri. Fhe third segment has biramous parapodia; the notopodia are very huge and diiet ted lateratl, the tieiuopodia are short and directed ventrad. Ventral cirri are missing on this segment; dot sal cirri are present. A single tentacular cirrirs, representing the dorsal cirrus of the second segment, is present; all other tentacular cirri have been lost. 1 he fii si and secoiul segments are completely (used to the prostomium; the third segment is separated from the rest of the anterior end by a deep incision. 22 Allan Hancock Foundation Monograph No. 11 Normal parapodia (Fig. c) have long, conical acicular lohes: the ventral cirri are digitate on segment 4; the cirri are cla\aie or distalK inllaied on all other segments. 1 he ventral cirrus reaches heyond the ii|) oí the acicular lobe in all segments. I'he dorsal cirri are lost in most setigers; those present are rectangular in outline; thev are attached along one ol the siiort sides and reach to the tip ol the acicular lobe. All setae in the first .seliger (Figs. d-e)areot one kind: each is thick, subdislalh inflated, and has a distinct tab on one side; a distinct, smooth pennon is attached below the le\ci of the tab. Ail setae in the second, biiamoiis, setiger (Fig. I) are similar. Each is ver\ stoiu, distalK sligiillv inllaied, and oblit]iiei\ truncate, with the distal margin disiincth dentale. All oilier setigers have similar compound seiae (Fig. g), wiih long smooth appendages and disiallv dentate shafts; the teeth on the shaft are all of similar si/e. The ^enusC/iarto fin lia is known lor one species onlv,C. nilssoui Malmgren (L'schakov, 1972, pp. 143-145, pi. 14, llgs. 5-10). The |)resent s])ecies agrees with the genus in that it has the fust two setigers completelv fused to the prostomium and the first lew setigers have motlified setae. It dillers fromC'. nilssoui in having the parapodia of the third segmeni greatly enlarged, and in having difiereni kinds of setae in the first and second setigers. The species is named for Dr. Andrew G. ("arey. Jr., who made it possible for us to study these collections. (jeogruphical Occtinoicc: One localil\ oil central Oregon. Eumidii. near lirsigera (Malmgren, 1865) SigcJiisigrrn Malmgren, ISfxT, p. 100, pi. 11, fig. 27. Eiimi/la /iisigem: L'schakov, 1972, pp. 156-157, pi. 10, figs. 8-10. Reconl: AD-17, N AD-17 ( 1 ). Remarks: The present specimen agrees w^ith Eitmida fusigera sensu L'schakov (1972), except ihai the paiapodia aie about twice as long as illustrated by L'schakov (1972, pi. 10, fig. 9). This difference is not significant enough to warrant a sejiarate status lor the single specimen. Ltigiu ahyssirola L'schakov, 1972 I.ugid ainssirola LIschakov, 1972, p[). 116-117, pi. 1. ligs. 1-5. Reaml: AD-9, NAD-21 (1). Remarks: Fhe genus Lugia is characterized by having the First segment modified with tentacular cirri, whereas the second segment has a normal neuropodial lobe with setae and ventral cirri, and only the dorsal cirri are modified as tentacular cirri. It dillers from Mystifies in having dorsal cirri on the third segment. Orrurrenee: Abyssal depths olT Japan. Priitotnystides occidoitalis (Ditlevsen, 1917) Mystides occidentalis Ditlevsen, 1917, p. 62, pi. 4, figs. 8, 11, 15. Protomystides orcidcntalis: LJ.schakov, 1972, p. 126, pi. 3, figs. 7-9. Records: AD-86, NAD-21 (2); AD-110, NAD-21 (1). Remarks: The two specimens from AD 86 were found together in the empty tube oï a Spiochaeto/Jterus species. Occurrence: The Davis Strait off Greenland in batlival depths. ¡'terocirrus nnajimai L'.schakov, 1972 Pterocirrus imajimai L'schakov, 1972, pp. 162-163, jjl. 9, figs. 11-14. Records: AD-6, NAD-17 (2): ?AD-89, NAD-22A (1); :--AD-149, NAD-15 (1). Remarks: The genera of phyllodocids with flattened ventral tentacular cirri were reviewed bv L'schakov (1972). There ma) be some doubt as to some of the conclusions drawn, but it appears to be best for the lime being to follow Uschakov's definitions. The speciinens from AD-6 agree fully with this species; they have elongated, pointed dorsal cirri, and the median aiUenna is aiiadied al the posterior end of the prosiomiimi. The specimens from .'\D-89 and AD-149 had lost all dorsal cirri, but otherwise closely resembled the preseni species. Occurrence: Off Japan and Baja California in abyssal depths. Deep-Water Polychaetes «» FAMILY POLYNOIDAE MALMGREN, 1867 Ewmiitit (inoniliita (Moore, 1910), new combination Antiitoe anoiitlatn Moore, 1910, pp. 3,58-3()l, pi. 30, figs. 34-40. Antiiioelld miorulata: Ilarinian. 1968, pp. 39-40, 7 figs. Records: AD-148, .NAD-12 (2); AD-149, NAD-15 (1). Remarks: The genus Eucranta .Malmgren, 1865, was erected for a single species, E. villnsa One species from the AiitarclK, /•:. mollis (Mclnlosh, 186; see Monro, 1936, pp. 100-101), originally described in Eujmlym,,; was mcncd lo Eucranta by ßergström (1916, pp. 294-295). The pre.sent species agrees with Eucranta m that it has 15 pans of elytra; prostomial peaks are present but are poorly developed; the notosetae are at least as thick as ihe neurosetac, and the latter are partially deeply and equally cleft at the tip, so that the proximal and distal teeth of die bidentate setae are of api)roximalely the same length E. villosa Malmgren ( 1865) aiul E. villosa uotialis Monro ( 1936, p. 901. fig. 1 1 a-h) have long slender elvtral papillae on the posterior lialf of eac h elyi ron. /•.'. mollis and E. auoculata have only short tubercles or papillae. E. mollis has two pairs of eyes; E. auoculata has no eyes. Occurreiicf: Southern and central California in bathval depths. Harmothoe, species indeterminable Records: AD-6, NAD-17 (1); AD-7, \AD-16 (3). Remarks: These specimens are incomplete and lack elytra; they agree with Harmothne in the structure of the prostomium and the setae, but cannot be completely identified. They all belong to one species. FAMILY .SIGAI.IONIDAE MALMGREN, 1867 KEY TO SPECIES la. A single median antenna present Pholoe' caeca b. Three antennae present 9 2a. Notopodial fimbriae in a group of six near the base on the posterior face . . .\eoleaiiira racemosa b. Four or five fimbriae in a row at the distal end of each notopodium S'eoleanira areolata Xeoleaiiira areolata (Mclntosh, 1885) Leanira areolata Mclntosli, 1885, p. 151-153, pi. 21, fig. 3, pi. 25, figs. 8, 9, pi. 13A, fig. 1. Neoleanira areolata: Pettibone, 1970, pp. 372-376, figs. 5-6. Sthenolepis areolata: Fauchald, 1972a, pp. 33-34, pi. 1, fig. i. Records: AD-33, NAD-21 (I); AD-64, NA1)-21 (I). ' Remarks: The present specimens agree with A', areolata as reviewed by Pettibone (1970). Occurrence: Bathyal depths from .southern (]alifi)rnia to Japan. Neoleanira racemosa (Fauchald, 1972) Sthenolepis racemosa Fauchald, 1972a, p. 34-36, pi. 2, figs. a-d. Records: AD-9, \AD-21 (11); AD-17, NAD-17 (1); AD-18, NAD-23 (1); AD-32 NAD-19 (!)• AD-4I NAD-21 (1); AD-110, NAD-21 (3); AD-119, NAD-22 (1). Remarks: The median antenna, not mentioned in the original description, is as long as that of .V. areolata. Occurrence: Western .Mexico in bathyal depths. Pholoe caeca Uschakov, 1950 (Plate 3, Figs, d-f) Pholoe minuta caeca Uschakov, 1950, p. 166; Uschakov, 1955, p. 165. Records: AD-6, NAD-17 (1); AI)-I7, NAD-17 (I); AD-141, NAD-llB (I). 24 Allan Hancock Foundation Monograph No. 11 Remarks: Pholoe caeca has never been illustrated; illustrations of the anterior end and setal structures (Fiíís d-f) are here added. In addition to the lack ot eves indicated by U.schakov (1950), the species also dilfcrs fVoni P. minuta (Fabricius, 1780; see Hartman-Schröder, 1971, pp. 79-80, fig. 24) in the proportions of the anterior end. The prostoniium is considerably more elongated than as illustrated for P. minuta (Fauvel, 1923, fig. 44c; Hartman-Schröder, 1971, fig. 24a). The differences are here considered to be of specific value. Occurrence: Sea of Okholsk in bathyal depths. FAMILY HESIONIDAE SARS, 1862 (iyf)tis hians, new species (Plate 5. Figs, a-e) Reconis: AD-22, NAD-11 (2, HOLOTYPF, Polv 1144, PARAFYPE, Poly 1145); AD-29, \AD-11 (2); AD-1S9, \AD-24 (3). Description: Ihe holotype is an incomplete .specimen with 27 setigers; it is 10 mm long and 2.n mm wide, excluding setae. It is yellowish and lacks color patterns; eyes are absein. The prostoniium (Fig. a) is wider than long in both type specimens, but it is probably distorted by ihe everted phai \ nx. It has a pair of smooth, slender, lateral antennae and a short, nearly button-shaped median antenna, attached well behind the frontal margin. The palps are about the same length as the lateral margin. Ihe palps are about the same length as the lateral antennae; each is biarticulated and is about twice as stout as the frontal antennae. Eight pairs of tentacular cirri are present. Fhe styles have been lost in most; those that are present arc slender and smooth. Paiapodia (Fig. d) are greatly elongated and are large compared to tiie size of the body; both rami are well developed. The nolopodium has a pointed acicular lobe with the lip superior to the end of the aciculum; the dorsal cirrus is longand smooth. The neuropodial acicular lobe is longer than the notopodial one and has a distinct, inferior digitate prolongation; a presetal pocket is present on the inferior edge oi the lobe. The ventral cirrus is long and slender and reaches beyond the tip of the acicular lobe. All notosetae (Figs, b-e) are long, slender, marginally dentate setae with the teeth in two rows; each seta is internally camerat'ed. Xeurosetae (Fig. e) are long, slender composite fakigers. Each ai)pendage is marginally serrated and has a short hood. 1 he tip is curved and is distinctly unidenlate in all setigers. The everted pharynx is basally smooth; distally it is widely flared (thus the specific name) and terminates in numerous papillae. Gyptis hians resembles C'. hrunnea (Hartman, 1961, pp. 69-70, pi. 5, figs. 1-4) Irom California. Both have well-developed notopodial rami with numerous setae. Both have the anterior margin of the eversible pharynx studded with numerous papillae, and both have the same general body shape. G. hians, however, has marginally serrated, camcrated notosetae, and parapodial rami that are distally prolonged into digitate tips. Furthermore, the neurosetae are distallv entire rather than bidentate as in G. hrunnea. Occurrence: Bathyal depths off central Oregon. Detailed data tor the type locality can be found in the Station List. Ophiodrotiius, species indeterminable Record: AD-6, NAD-17 (1). . . « Remarks: I he specimen is poorly preserved: it appears to differ from the commonly occurring shallow- water species, O. pugettensis. Deep-Water Polychaetes 25 FAMILY FILARGIIDAE SAINT-JOSEPH. 1899 KKVrO SPECIES la. Emergent notopodial spines siraighi Syiii'lmis near klatli 1). Emergent noioixKUal spines curved . . . ' 9 2a. Antennae distinctly longer than the ¡xilps 3 h Antennae shorter than the palps - •Sa Median antenna distinctlv longer than the lateial ones; notosetae 'iú,scin ^^^Si¡ranilmün¡tacnÍr Aucislrosylli.s l)rei'i(ej)s Hartman, 19(33 (Plate :\. Fig. c) Ancistrosyllisbreviceps Hartman, 19(").'5. pp. 13-16, fig. la-d; Pettibone, 1966, p. 168; Fig. l,a-(l; Hartman 1968. pp. 37.'5-376. 1 figs. ReamU: AI)-42, NAD-21(1); AD-139, \AI)-24 (I); AD-141. NAD-1 IB ( 1 ). Remarks: One of the present specimens has the pharynx everted (fig. c). It consists of an anterior part with about 14 thick marginal lobes and a papillose posterior part. These papillae are in five to six irregular rows on the ventral side of the phar\nx and are irregularly dispersed on the dorsal side. Ociiirrenre: Bathyal and deep-shelf areas of f southern California. Aiiri.strusyllis. ncAV groeiil/intlica Mclntosh, 1879 Ancistrosyllisgroenlundini Mclntosh. 1879, p]). 502-503, pi. 65, figs. 3. 20; Pettibone, 1966, pp. 16-168, fig. 3, a-l. Records: Al)-6, \AD-17 (1); AD-9, NAD-21 (1); AD-41. NAD-2I (3) AD-110, NAD-21 (2)- "-AD-n9 NAD-22 (I juvenile). Remarks: The present specimens agree with Ancistrosyllis groenlandica sensu Pettibone (1966) in most characters; however, the integument is smooth or wrinkled rather than papillo.se, and the first hooks are present on segnients 3 to 7, depending on the si/.e of the specimen, rather than from 4 to 6 as in A. groenlandica. The differences are small, and the material is poorly preserved. Better material will be needed in order to document the presence of this species off Oregon. Sigamhra setosa Fauchald, 1972 Sigambra setosa Fauchald, 1972a, pp. 62-64, pi. 7, figs. a-c. Records: AD-7, NAD-16 (1); AD-139, NAI)-24 (1); AD-149. NAD-15 (1). Remarks: Sigamhra setosa belongs to a group of s|)ecies that has hooks jiresent from anterior setigers; it differs from other members of this group in that notosetae are jjieseni in all setigers and the median antenna is distinctly lotiger than the lateral ones. Occurrence: The upper end of the Gulf of California in bathyal depths. Sigambra tentficnlata (Ire-.idwvU, 1941) Ancistrosyllis tentaculata Tread well, 1941, pp. 1-3, figs. 1-3. Sigambra tentaculata: Pettibone. 1966. pp. 182-186.'figs. 14-15; Hartman 1968 np 391-39'> 3 fies Record: AD-7, NAI)-16 (1). ' • t 1 • • « • Remarks: Fhe piesenl specimen lacks notosetae; hooks are present from .seliger 4; and the median anemia is distincily longer than the lateral ones. Occurrence: Known from a wide depth range down to bathyal depths, from both the Atlantic and Pacific coasts of North and .South .America. 26 Synelnm, neTir klatti (Friedrich, 1950) Chphohesione klatti Friedrich, 1950, pp. 171-173, ilgs. 1-2. Synelnos klatti: Pettibone, 1966, p. 191; Hartman-Schröder, 1971, pp. 144-14n, fig. 49a-e. /{m;íY/.- AD-154, NAD-26(1). ,,nA«^I, Remarks: The present specimen agrees with Synelmis as this genus was detmed by Pettibone (19bb). It resembles S. klatti, but is too poorly preserved tor complete identification. FAMILY SYLLIDAE GRUBE, 1850 Sphaerosyllis, near californiensis Hartman, 1966 Sphaerosylüs califormemis Hartman, 1968, pp. 453-454, 7 figs. ßicorflf.-'AD-6, NAD-17 (1). Remarks: The present specimen resembles Sphaerosyllis ralifnmiensis; however, the eyes are less well developed and are not lensed, and the serrated anterior falcigers are considerably more .slender and less distinctly serrated than in S. californiensis. FAMILY NEREIDAE JOHNSTON, 1845 Ceratocephale loveni Malmgren, 1867 Ceratocephale loveni Malmgren 1867, pp. 61-2, pi. 5, fig. 33; Hartman, 1952, p. 15; Hartman, 1960, p. 95. Record: AD-7, NAD-16 (1). Remarks: The specimen belongs to the stein species, rather than to the subspecies/?rtrí/írrt Hartman, 1960, in that the jaws have 11 teeth and the furcate ventral cirri are absent on at least the first setiger. Furthermore, the posterior para podia are relatively shorter than in the subspecies and are more as they are in the main form. Occurrence: May have been confused with similar related species. Is widely dispersed in the northern hemisphere in lower shelf and bathyal depths. FAMILY GLYCERIDAE GRUBE, 1850 KEY TO SPECIES 1 a. Parapodia with two postsetal lobes Glycera robusta b. Parapodia with a single postsetal lobe Glycera projundi Glycera profundi Chamberlin, 1919 Glycera profundi Chamberlin, 1919, pp. 350-352, pi. 64, figs. 2-6; Fauchald, 1972a, pp. 103-104, pi. 22, figs. a-g. Records AD-7, NAD-16 (1); AD-9, NAD-21 (1); AD-65, NAD-21 (1); AD-I41, NAD-IIB (1); AD-149, NAD-15 (2). Remarks: G. profundi was redefined by Fauchald (1972a) and distinguished from the very snnilar G. capitata Orsted, Í844 and G. branchiopoíla Moore, 1911. It has a single postsetal lobe in all setigers; the pharyngeal organs are smooth and are either tall and slender, or wide and folióse. The posterior parapodia have strongly prolonged parapodial bases. Occurrence: Gulf of California in bathyal depths. Deep-Water Polychaetes 27 Glycera robusta Ehlers, 1868 Glycera robusta Ehlers, 1868, pp. 656-658, pi. 24, figs. 31-32; Hartman, 1968, pp 627-628 4 figs • Hartman and Reish, 1950, p. 20. i r . g -, Record: AD-9, NAD-21 (1). Remarks Glycera robusta has two postsetal lobes; the piiaryngeal organs have transverse ridges and the branchiae, which are not retractile, form small blisters on the parapodial bases. Occurrence: ]A\yâ.n to southern California in shelf depths. FAMILY GONIADIDAE KIN'BERG, 1866 KEY TO SPECIES la. Prostomium smooth Bathyghcmde cedroerisis b. 1 rostommm distinctly annulated G..»«./a bruunea Bathyglycinde cedroensis Fauchald, 1972 Bathyglycinde cedroensis Fauchald, 1972a, pp. 107-108, pi. 23, figs h-n Records: AD-9, NAD-21 (2); AD-33, NAD-21 (2). Remarks: The genus Bathyglycinde was defined by Fauchald (1972a) for three species with smooth prostomia, and with pharyngeal organs similar to those in Ghcmde but missing f lom areas I and VI as these areas were defined by Hartman ( 1950, pp. 45-47). The notosetae are all fine capillaries. B. cedroensis differs f^rom B. lindberg,(\]^ch,ikox, 1955, p. 176, fig. 49g-l) in that it has large and folióse rather than small and digitate ventral cirri and the postsetal lobes are longer than the presetal ones mB. cedroensis and of the same length in ß. lindbergi. Occurrence: Bathyal depths off Baja California, Mexico. Goniada brunnea Treadwell, 1906 GonoíWrtftrMwwctíTreadwell, 1906, p. 1174, figs.67-70; Hartman. 1950, pp. 17-19 pi 1 figs 1-6 pi 4 fig I, text fig. 1; Hartman and Reish, 1950, p. 21; Fauchald, 1972a, pp. 111-112 ' 't^ - 8- Records: AD-6, NAD-17 (3); AD-7, NAD-16 (2); AD-9, NAD-21 (3); AD-17 NAD-17 (1)- AD-33 NAD-21 (2); AD-44, NAD-22 (1); AD-89, NAD-22A (1); AD-110, NAD-21 (1); AD-141 \AD-11B (1)' AD-148, NAD-I2(I); PAD-149, NAD-I5(1); AD-154, NAD-26(1). " Remarks: The species is accepted here as redefined by Hartman (1950). The number of micrognaths varies as indicated by Hartman, and the size varies as well. Occurrence: Widely dispersed horizontally and vertically in the eastern Pacific Ocean. FAMILY NEPHTYIDAE GRL'BE, 1850 KEY TO SPECIES la. Interramal cirri involute; palps simple Aglaophamus, near eugeniae b. Interramal cirri recurred; palps double Nephtys comuta Aglaophamus, near eugeniae Fauchald, 1972 Aglaophamus eugeniae Fauchald, 1972a, pp. 82-84, pi. 14, figs a-e Record: AD-139, NAD-24 (1). Remarks: The present specimen agrees with A. eugeniae in the structure of the parapodia and in the absence of notopodial cirri on setiger 1. It differs in that the prostomium is square rather than rounded and in that the first interramal cirri are present from setiger 13 rather than from setigers 8 to 10. The shape of the prostomium is somewhat variable in the nephtyids, but is usually not as variable as seen here. 2g Allan Hancock Foundation Monograph No. 11 Siiiiilarlv the first occurrence of the interramal cirri may vary somewhat, unless they first occur in one of the fnst't'hree or lour setigers, but they usually do not vary as much as would be suggested by considering the present specimen a member of A. eMgfnJai. Nel>litys cnrttuta Berkeley and Berkeley, 1945 Nehth^^ cormita Berkeley and Berkeley, 1945, pp. 328-330, figs. 2-4; Fauchald, 1972a, p. 90. Records: AD-16, NA1)-13 (4): AD-141, NAD-1 IB (6); AD-149. NAD-15 (5). Remarks- The subspecies//•«».-nsr««« described by C:iark and Jones (1955) may not be separable from the main form, as indicated by Hartman (1968, p. 581). Ihe species is characterized by its bifid palps; however, the parapodial lobes have been poorly described, and the variation in structure was not noted. It appears best to retain the two concepts separately for the time being. Occurrence: Shelf and balhyal areas in the northeastern Pacific Ocean. FAMILY rOMOlTERIDAK GRUBE, 1848 I omopteridae, indeterminable Aícorí¿.- AD-139, NAD-24 (2); AD-I41, NAD-1 IB (fragments). Remark.',: Fhese si)ecimens and fragments of pelagic polychaetes must either have been caught by the gear on its way up or down ihiougl'i the water or have come in with the water used for screening the samples. FAMILY AMPHINOMIDAE SAVIGNY, I8I8 F ara m t>h i name ¡¡ac'ifica new species (Plate 6, Figs, a-d) Records- AI)-6, NAD-17 (1); AD-7, NAD-I6 (6); AD-18, NAD-23 (I); AD-19, NAD-22 (I); AD-65, NAD-2I (4); AD-86, NAD-2I (1); AD-89, NAD-22A (I); AD-90, NAD-22 (I, HÜLOTYPE, Poly 1157), AD-IIO, NAD-26 (I); AD-119, NAD-22 (7), AD-I39, NAD-24 (8); AD-150, NAD-21 (7). Description: I he type is a complete specimen with 37 setigers. It is 11 mm long and 2 mm wide, excluding setae. The anterior part of the body is inflated. I'hc anterior end is truncate, and the body tapers evenly to the posterioi end. The body is yellowish and lacks color patterns; there are no eyes. The prostomium (Fig. a) has a quadrangular posterior part condnued posteriorly as a ridge. The anterior \y,\v\ of the prostomium is semicircular and has two pairs of antennae; the medial pair is digitate and sessile, and the outer i)air is somewhat longer. Each antenna is attached to a short ceratophore. The first setiger, which is complete, has very long dorsal and ventral cirri. All the other parapodia (Fig. d) are similar. In each, the two rami are well separated; the notopodial lobe is short and blunt and has a long, posteriorly attached dorsal cirrus; the neuropodium is somewhat more prominent, usually truncate in outline, and'the ventral cirrus is attached distinctly ventrally. Dorsal and ventral cirri are both slender. Branchiae include five or six pairs, stardng on setiger 3; each is dichotomously branched. I he very smallest specimens had three pairs of branchiae, but none of the 36 specimens examined had any more than six pairs. , Notosetae include a superior fascicle of very stout spines with fine reverse dentition, and an interior fascicle of longer, somewhat slenderer, completely smooth spines. Neurosetae (Figs, b-e) include a few thick, subdistally infiated spines, with a spur and a denate portion, distally ending in a curved, thickened knob! Additionally, each fascicle contains many long, slender, marginally dentate setae and one or two long, slender, smooth setae in addition to the arrow-headed, thick acicula. Deep-Water Polychaetes •q Paraphinome pacifira resembles P. jeffrpysii (Mclntosh, 1867; see Hartman-Schröder, 1971, p. 31). It differs most markedly in having four rather than five, anteiniae, and even fiilly grown forms never have more than six pairs of branchiae. P. jejjreysn has up to 12 pairs of branchiae.' Occurrnur: Several stations in depths ranging from 1800 to 2900 m off central Oiegon. Detailed data for this type locality can be found in the Station List. FAMILY OM'PHIDAE KI.\BF.R(;, 186.5 KEY TO SPECIES la. Peristomial cirri absent; tube hyaline and quill-like Hyalinoeda strirta b. Peristomial cirri present; tube other than hyaline and quill-like 2 2a. At least some branchiae pectinate •^ b. Branchiae absent or, if present, simple and strajT-like 4 3a. Simple tridentale hooks present in some anterior setigers Onuphis vexillaria b. Simple tridentate hooks absent Onuphis pw/undi 4a. Branchiae absent Mothria mixta b. Branchiae jjresent 5 5a. Branchiae present from setiger 1 Mothria iridescens b. Branchiae first jjiesent from a later setiger 6 6a. Composite hooded hooks tridentate 7 b. Composite hooded hooks bidentate 9 7a. Oratophores nearly smooth Nothria mexicana b. Ceratophores distinctly annulated 8 8a. Distal tooth of coinposite hooks distinctly longer than the other teeth; branchiae from setiger 5 Nothria geophiliformis b. Distal tooth of composite hooks approximately as long as the other teeth; branchiae from setiger 4 J\'othria pallidn 9a. Tube flattened and covered with shell fragments; branchiae not present before setiger Ï0 Xothria cunchylega occidentalis b. lube cylindrical and covered with fine iriud particles; branchiae present before setiger 10 Nothria lepta Hynlinoeria stricta Moore, 1911 Hyalinoeda tubicola stricta Moore, 1911, pp. 280-282, pi. 18, figs. 96-97. Hwlinoecia stricta: I-auchald, 1968, pp. 16-17, pi. 3, figs, f-k; Fauchald, 1972a, p. 120. Record: AD-UO, NAD-21 (1). Remarks: The present speciinen agrees with the species as reviewed by Fauchald (1972a). It is the first record of the species north of Point Conception, California. Occurrence: Deep water from southern California to Panama. Nothria conchylega occidentalis Fauchald, 1968 Nothria conchylega occidentalis F"auchald, 1968, pp. 20-21, pi. 5, figs. a-n. Record: AD-3Ü, NAD-ll (I). Remarks: The subspecies differs from the main form in that the anterior hooded setae are simple rather than composite, and the postsetal lobes are auricular in two setigers rather than in onlv one. Occurrence: From southern California to Colombia in shelf depths. Nothria geophiliformis (Moore, 1903) • Northia geophilijormis .Moore, 1903, pp. 445-448, pi. 25, figs. 57-59. Nothria geophiliformis: Fauchald, 1968, p. 22, pi. 6, figs. a-d. Record: AD-I6, NAD-I3 (1). 30 Allan Hancock Foundation Monograph No. 11 Remarks: Nothria geophtliformis has branchiae Irom setiger 5, the first setiger has a low, transverse presetal lobe and on the luioded hooks the distal tooth is greatly prolonged and projects beyond the edge of the hood. Records of this species from Spain (Ibai-iez, 1972 and 1973) need to be confirmed, especially since the population appears to have a different habitat from the one previously reported. Occurrence: Japan to Baja California, Mexico in shelf and slope depths. May be present in interddal sand flats in Spain. Nothria iridescens ]o\\n%on, 1901) Nothria iridescens ]ohn%on, 1901, p. 408, pi. 8, figs. 86-87, pi. 9, figs. 88-92. Nothria iridescens: Fauchald, 1968, p. 24, pi. 7, fig. a (in part); Hartman and Reish, 1950,p. 2?>: Pauchald, 1972a, pp. 124-125. OnufMs iridescens: Hobson, 1971, pp. 533-535, fig. 3a-d, 4a-c. Record: AD-148, NAD-12 (1). Remarks: Nothria iridescens is here accepted as restricted by Fauchald (1972a) and Hobson (1971). Occurrence: Shelf and slope depths from Britkh Columbia to western Mexico. Nothria lepta (Chamberlin, 1919) Onuphis lepta Chamberlin, 1919, pp. 290-295, pi. 45, figs. 1-7. Nnthria lepta: Fauchald, 1972a, pp. 125-126. Records: AD-9, NAD-21 (1): AD-18, NAD-23 (3); AD-44, NAD-22 (3); AD-53, NAD-26 (1); AD-llO, NAD-2I (1); AD-119, NAD-22 (1); AD-139, NAD-24 (1); AD-154, NAD-26 (1). Remarks: The present specimens agree with Nothria lepta in the strucure of the anterior end, the first occurrence of the subacicular hooks, and the construction of the first setigers. They differ from the species as reviewed by Fauchald (1972a) in that the first branchiae are present on setigers 7 to 10 rather than on setiger 6 as described. Occurrence: Slope depths off Panama and Baja California. Nothria mexicana Fauchald, 1968 Nothria mexicana Fauchald, 1968, pp. 25-26, pi. 7, figs, b-e; Fauchald, 1972a, pp. 126-127. Records: AD-6, NAD-17 (1); AD-17, NAD-17 (3); AD-105, NAD-12 (3); AD-148, NAD-12 (2); AD-150, NAD-26 (2); AD-154, NAD-26 (1). Remarks: Nothria mexicana resembles A', geophiliformis in most characters, but differs in having long, nearly smooth occipital ceratophores and capillary setae that are finely pilose, rather than limbate. Occurrence: The central American Trench and off Baja California, Mexico in slope depths. Nothria mixta, new species (Plate 7, Figs, a-f) Records: AD-139, NAD-24 (3, HOLOTYPE, Poly 1150, PARATYPES, Poly 1151). Description: The holotype is an anterior fragment of 19 setigers; it is 8 mm long and 1 mm wide, excluding setae. The other specimens are longer, but are less well preserved. The anterior part of the body is cylindrical; the body is very strongly flattened dorsally from seliger 6. The specimens are yellow and lack color patterns; there are no eyes. The prostomium ( Fig. a) is distincdy wider than it is long, with a pair of short, ovate frontal antennae that are directed ventrad. There are five short occipital antennae. The inner lateral ones are the longest and reach setiger 3; the median and outer lateral ones reach setiger 2. The ceratophores have approximately four distinct annulations, of which the distal-most is the longest. All ceratostyles are tapering. The first parapodia (Fig. d) have rounded acicular lobes; the presetal lobes are short and truncate; the digitate poslsetal lobes are broadly attached. The dorsal and ventral cirri are digitate; the dorsal cirrus is somewhat longer than the ventral one. A distinct cirrophore was not visible. Ventral cirri are cirriform in the first three setigers and pad-shaped from setiger 4. Deep-Water Polychaetes 31 Anterioi- setiffers ha\e hooded pseudocomposite hooks (Figs, b-c): each has a shatply pointed hood dislally and is bi- or li ¡tiéntate. The teetii decrease evenly in si/e from liie most distal one and iorm a wide angle with the shaft. Subacicnlar hooks (Fig. e) are present from setiger 12 in all specimens; each hook is distally bidentate, with the subdistal tooth at a nearK right angle to the shaft of the seta. IVdinaie seiae (Fig. I) aie jjresent in median setigers. Each is dislally sligluK <)bli<|ueand hasaboiii 2()ieeiii; the marginal teeth are not markedly longer than the others. Branchiae are ai)sent. The jaw apparatus was not dissected on the few available sjjecimens. Abranchiate species of AWir/rt were reviewed by Faiichald (1968, p. 27; 1974, p. 18). Of the species listed by Faiichald (1968), none lias both bi- and iridenlale hooks in anierioi- seligers. as does ,V. wixta. It most closely resembles species in the group with iridenlale hooded hooks and can be separated from ihem as indicated below. The pectinate setae have 10 leeili in A', notialis (Monro, 1930, ])p. 129-131, fig. 48) and in A', py^idialis Faiichald (1968, pp. 26-27, pi. 7, figs, f-m), and 20 or more teeth m \. fiordica Faiichald (1974, pp. 1.5-18, fig. la-f) and in .V. inixla. In S. fiordica subacicnlar hooks are present from setiger 16 and each hook has the teeth at an obtuse angle witii the shaft. In A', wixla subaci« ular hooks are present from setiger 12, and each hook has the teeth at right angles to the shaft. I'he species name refers lo the presence of both bi- and iridentale anterior hooks. Occurrence: One locality off central Oregon. Details can be found in the Station List. Nothria Imllida Moore, 1911 Nothria tmllidd Moore, 1911, pp. 2.')6-259, pi. 15, figs. 24-28, figs. 35-37; Fauchald, 1972a, p. 127. Records: AD-6, XAD-H (1); AD-7, NAD-16 (3): AD-17, \AD-17 (1); AD-149, \AD-15 (1); Al)-150, NAD-26(1). Remarks: Nothria ¡mllida has branchiae f lom setiger 4; it has irideiitate hooded hooks, and the ventral cirri are pad-shaped from setiger 5. Occurrence: Southern and centra! ('.alilornia in slope depths. Onu/j/iis praftindi Fauchald, 1968 Onuphis profuudi Fauchald, 1968, pp. 40-41, pi. 10; Fauchald 1972a. pp. 133-134, pi. 26, fig. a. Records: Al)-6, NAD-17 (3); AD-18, NAD-23 (1); AD-33, NAD-21 (1); AD-88, NAD-22A (1); AD-89, NAD-22A (1); AD-139, \AD-24 (1); AD-150, NAl)-26 (1). Remarks: Onuphis ¡rrofundi has branchiae from setiger 6 or 7. Subacicnlar hooks are first i^resent from setigers 17 to 22 (in the present specimens, most f recpiently from setigers 20 to 22). Fhe composite anterior hooded hooks are iridenlale and the ventral cirri are pad-shaped from setiger 8. Occurrence: Slope dei)ths off western Mexico. Onuphis vexillaria Moore, 1911 Onuphis vexillaria Moore, 1911, pp. 266-269, pi. 17, figs. 69-76; Fauchald, 1968, p. 43; Fauchald, 1972a, p. 134. Records: AD-9, \AD-21 (2); AD-19, NAD-22 (1); AD-44, NAD-22 (2); AD-53, NAD-26 (1); AD-55, NAD-25 (3); AD-65, NAD-21 (1); AD-86, NAD-21 (4); AD-88, NAD-22A (1); AD-110, NAD-21 (3); AD-145, NAD-23 (2); AD-1.54, NAD-26 (1). Remarks: O. vexillaria has a lew large, tridentate simple hooks on some of the anterior setigers in addidon to the usual composite hooks. These siinjile hooks, called "acicular hooks'" by Hartman (1944a, pp. 68-69), are about twice as thick as the other hooded hooks and usually appear to be somevvhai inflated compared with the other hooks. They are usually present on only a few setigers. In the present material they are regularly present on setigers 5 and 6 and, in most specimens, also on setiger 7. I'liey are always absent from seliger 8 and later. Branchiae are first present from setiger 5; subacicnlar hooks are present from setiger 50 in fully grown specimens; the ventral cirri are pad-shaped from setiger 8 or 9. Occurrence: Southern California and western Mexico in slope depths. 32 Allan Hancock Foundation Monograph No. 11 FAMILY EUNICIDAE SAVIGNY, 1818 KEY TO SPECIES la. Maximally 8 branchial filaments in each branchia; distal tooth of composite hooks curved Eunice kobiensis b. Maximally 16 branchial filaments in each branchia; distal tooth of composite hooks straight.... Jiunice segregata Eunice kobiensis Mclntosh, 1885 Eunice kobiensis Mclntosh, 1885, pp. 278-280, pi. 38, figs. 12-13, pi. 2ÜA, figs. 1-3; Fauchald, 1969, pp. 4-6, fig. 2. Eunice longicirrata: Hartman and Reish, 1950, p. 23 (not Webster, 1884). Recorch: AD-6, NAD-17 (1); AD-7, NAD-I6 (1); AD-13, NAD-18 (1); AD-86, NAD-21 (1). Remarks: Eunice kobiensis was separated from other similar species by Fauchald ( 1969), who demonstrated that what had been called £". kobiensis from the eastern Pacific Ocean in fact belonged to three species. The present specimens have branchiae from seliger 3 to setigers 26, 27, 37 and 37, respectively, and have a maximum of seven branchial filaments. A very small specimen has subacicular hooks present from seliger 13; the other specimens have these hooks present from .setigers 24 to 34. The structure of the anlerior appendages and of the dorsal cirri closely resembles the conditions in the type specimen. (iustus (1972) attempted to demonstrate that the characters used to separate the species listed by Fauchald (1969) were invalid because of too much intra- and interpopuladon variation. Her statistical argument is unconvincing due to inadequate materials, and I he species are here considered separable. Occurrence: Japan and northeastern part of the Pacific Ocean in shell and slope depths. Eunice segregata (Chamberlin. 1919) Leodice segregata Chamberlin 1919, ])p. 237-240, pi. 54, figs. 1-5, in ])art. Eunice segregata: Fauchald, 1969, pp. 6-8, fig. 3; Fauchald, 1972a, pp. 143-144. Records: AD-6, NAD-17 (1): AD-17, NAD-17 (1). Remarks: The present specimens have branchiae from seliger 3 to setigers 38 and 42, respectively. The maximal number of branchial filaments is 16. Subacicular hooks are fust present from setigers 38 and 46, respectively. Occurrence: Southern California to Panama in slope and bathyal depths. FAMILY LUMBRINERIDAE MALMGREN, 1867 KEY TO SPECIES 1 a. Branchiae present 9 b. Branchiae absent 2 2a. Composite hooded hooks present 3 b. Composite hooded hooks absent 6 3a. Acicula black 4 b. Acicula yellow fi 4a. A single median nuchal papilla present; both pre- and postsetal lobes prolonged in posterior setigers Lumbrineris eugeniae b. Paired lateral nuchal pits present; only postsetal lobes prolonged in posterior setigers Lumbrineris index 5a. Both pre- and postsetal lobes prolonged in posterior setigers Lumbrineris cruzensis h. Neither pre- nor postsetal lobes prolonged in any setiger Lumbrineris, near latreilli 6a. Acicula black 7 b. Acicula yellow 8 7a. Both pre- and postsetal lobes prolonged in posterior setigers Lumbrineris bicirrata b. Neither pre- nor postsetal lobes prolonged in any setiger Lumbrineris moorei Deep-Water Polychaetes 33 8a. Simple hooks jjiesent from seliger 1; both pre- and postsetal lobes prolonged in posterior sciigers I.iiinbrineris, near Uigiina b. SiiniJle hooks first present from seliger 21; neither pre- nor postsetal lobes prolonged in any seliger Lumhrineris, near nhyssicola 9a. Branchiae branched Ninoe loiigibranchia b. Branchiae simple, short postsetal knobs 10 lOa. Hooded hooks first ])resent from a median setiger (setigers 45-74) ù^inoe fusca b. Hooded hooks first present from setiger 1 Ninoe fuscoides Lumhrineris, near abyssicola Uschakov, 1950 Lumhrineris ahyssirola Uschakov, 1950, p. 195, fig. 27. Record: AD-65, NAD-21 (1). Remarks: The |)reseni specimen has short parapodial lobes in all setigers. The aciciila are yellow, and simple hooded hooks arc present from setiger 21. The setae are very long compared with the width of the bodv. The sjiecimen differs from /.. nhyssicola in that the latter lias hooded hooks present from one of the first setigers. The anterior end has been crushed, so the jaw structure could not be studied. Lumhrineris hirirratn Treadwell, 1929 Lumhrinereis hicirrata Treadwell, 1929, pp. 1-3, figs. 1-7. Lumhrineris hicirrata: Fauchald, 1970, pp. 77-78, pi. 10, figs, e-g; Hartman and Reish, 1950, p. 24; Fauchald, 1972a, p. 147. Records: AD-149, NAD-15 (2); AD-150, \Al)-2tt (1). Remarks: Lumhrineris l>icirrata has imidentate maxillae III, prolonged posterior pre- and postsetal lobes and black acicula; composite setae are absent. Occurrence: Shelf depths from Washington to western Mexico; one bathyal locality off Baja California, Mexico. Lumhrineris cruzensis Hartman, 1944 Lumhrineriscruzensis Hartman, 1944, pp. 165-166, pi. 17, figs. 263-269; Fauchald, 1970, pp. 83-84, pi. 12, figs, g-j: Fauchald, 1972a, p. 149. Records: AI)-1 11, NAD-llB (2); PAD-149, \A1)-15 (I). Remarks: Lumhrineris cruzensis lias a rounded prostomium, |)rolonged pre- and postsetal lobes in posterior setigers, composite setae in some anterior setigers and yellow acicula. Maxilla HI is unidentate. Occurrence: British Columbia to western Mexico in shelf and bathyal depths. Lumhrineris eugeniae P'auchald, 1970 Lumhrineriseugeniae Fauchald, 1970, pp. 87-89, pi. 13, figs, c-f ; Fauchald, 1972a, pp. 149-150, pi. 29, figs, a-b. Records: AD-6, NAD-17 (1); AD-9, NAD-21 (1). Remarks: Lumlnineris eugeniae has a single median eversible nuchal papilla. The pre- and postsetal lobes are prolonged in posterior setigers; acicula are black. Composite setae are present in some anterior setigers. Fhe setae are strongly prolonged. The prostomium is very long, with nearly parallel sides and a bluntly truncate anterior end. Occurrence: Bathyal depths off western Mexico. Lumhrineris index Moore, 1911 Lumhrinereis japónica index Moore, 1911, pp. 288-289, pi. 19, figs. 119-127. Lumhrineris index: Hartman, 1944, pj). 162-163, pi. 12, figs. 254-256; Fauchald, 1972a, p. 150. Record: AD-30, NAD-11 (3). Remarks: Ibis species has black acicula, prolonged poslsetal lobes in posterior setigers, and composite setae in the first setigers. Maxilla 111 has two teeth. Occurrence: Central and southern California in shell and bathyl depths. 34 Allan Hancock Foundation Monograph No. 11 Lumbrineris, near lagunae Fauchald, 197Ü Lumhrineris lagnnae Fauchald, 1970, pp. 92-94, pl. 15, tigs. a-e. Records: AD-llO, NA1)-21 (1); AI)-139, NAI)-24 (1). Remarks: The present syiecimens agree with L. Inginuie in that they have yellow acicula, and simple hooded hooks are present from the first setiger. The pre- and poslsetal lobes may l)e |)rolonged in posterior seiigers. Maxilla III has a single tooth. Ihe postsetal lobes in anterior setigers are triangular, not broadly rounded as in /,. laguune. The degree of prolotigation of the pre- and postsetal lobes in posterior setigers could not be assessed fully, since both speciiuens were incomjilete |)osteriorly. Lumbrineris, near latreilli Audoiiin and Milne Edwards, 1834 Lumbrinereis latreilli Audouin and Milne Kdwards, 1834, pp. 168-170, ])1. 3B, figs. 13-1.^. Lumbrineris latreilli P'auchald, 1970, pp. 94-97, pl. 1.^, figs. f-h. Records: AD-U, NAD-15 (1); AD-141, XAD-llB (1); AD-149, NAD-15 (1). Remarks: Fhe present specimens agree with L. latreilli sensu Fauchald (1970). As pointed out by Fauchald ( 1974, p. 24), species in this complex have been conf used and are difficult to separate. The present material is neither extensive enough nor well enough preserved to permit a more complete description. Lumbrineris moorei Hartman, 1942 Lumbrineris moorei Hartman, 1942, pp. 116-118, fig. 12a-b and g; Fauchald, 1970, p. 102; Fauchald, 1972a, p. 153. Records: AD-6, NAD-17 (1); AD-9, NAD-21 (3). Remarks: Lumbrineris moorei has prolonged anterior setae. Simple hooded hooks are preseiu liom about setiger 25, and the po.slerior parapodial lobes are short. Acicula are black, and maxilla 111 has a single tooth. Occurrence: Bathyal depths off soiuhern California and western Mexico. Ninoe fusca Moore, 1911 Ninoe fusca Moore, 1911, pp. 285-288. pl. 19, figs. 110-118; Hartiuan, 1968, pp. 781-782, 7 figs • Fauchald, 1970, pp. 116-117; Fauchald, 1972a, p. 156. Records: AD-6, NAD-17 (1); AD-33, NAI)-21 (2); AD-44, NAD-22 (1); AD-65, \AD-21 (3). Remarks: Ninoe fusca has a central nuchal pocket. Branchiae are present from the first setiger and are limited to some (usually about 40) anterior setigers. Hooded hooks are present from a median setiger, between setigers 45 and 75. The acicula arc black, and the bases of the setae are dusky. Occurrence: Southern California and western Mexico in bathyal and abyssal depths. Xinoe fiiscoides Fauchald, 1972 Ninoe fuscoides Fauchald, 1972a, pp. 156-158, pl. 31, figs. a-f. Records: AD-6, NAD-17 (2); AD-9, NAD-21 (2); AD-16, NAD-13 (3); AI)-17, NA1)-17 (6); AD-32, NAD-19 (2); AD-89, NAD-22A (1); AD-139. \An-24 (I); Al)-154, NAD-26 (I). Remarks: Ninoe fuscoides has branchiae and branchial distribution similar to that found in .V. fusca. However, the hooded hooks are present ÍVom the first setiger, or at least before setiger 5, and maxilla 11 has foin-, rather than two, teeth as in N. fusca. The acicula are black and the setae are basally dusky. Occurrence: Central American trench and off Baja California, Mexico, in bathyal and abyssaldepths. Ninoe longibranchia Fauchald, 1972 Ninoe longibranchia Fauchald, 1972a, pp. 158-160, pl. 32, figs. a-g. Records: AD-6, NAD-17 (1); AD-7, NAD-16 (1); AD-13, NAD-l's (2); AD-16, NAD-13 (2); AD-141 NAD-llB (3); AD-149, NAD-15 (5). Remarks: N. longibranchia has branchiae only on a few anterior setigers, and the superiormost filament is cirriform and nearly twice as long as all others. The maximum number of branchial filaments is six. N. gemmea .Moore, 1911, with which this species has been confused, has branchiae from setiger 4 to about setiger 50 and maxiiually has three branchial filaments (Hartman, 1968, p. 783; Fauchald, 1970, p. 117). Occurrence: Bathyal depths of I western Mexico. Deep-Waler Polychaetes 35 FAMILY ARABELLIDAE HARTMAN, 1944 Drilnnercis fdlcata Moore, 1911 Drdoncreislakata Moore, 1911. pp. 298-299, pi. 20, figs. 150-154; Fauchald, 1970, pp. 135-136, pi. 21, fijr. g; Fauchald, 1972a, p. 161. Records: AD-6, XAD-17 (1); Al)-16, \A1)-13 (1); AI)-141, NAD-llB (I). Remarks: In Drilonereu falcata, maxilla I is proximally dentate, and mandibles are present. Orniirrnri': Siielf areas oil (California and batliyal de|Jlhs off western Mexico. FAMILY DORVILLEIDAE CHAMBERLIN, 1919 Dortnllea batia Jumars, 1974 Duniillea batia Jumars, 1974, pp. 115-117. fig. 6. Record: AD-149. \AD-15 (1). Remarks: I), batia has poorly developed anterior appendages. It has smooth setae which are distally spinigerous rather than lalcigerous. as is usual in Dorvillea. Occurrence: San Diego Trough, California in bathyal depths. FAMILY STERNASPIDAE CARUS, 1863 Sternaspis fossor Stimpson, 1853 Stermisf)is fossor Stim|)son 1853, p. 29, fig. 19; Hartman, 1969, pp. 351-352, 1 fig. Sternaspis sciitata; Hartman and Reish, 1950, ]). 38. Records: AD-9, NAD-21 (1); AD-32, NAD-I9 (1); AD-33, NAD-2I (7); AD-64, NAD-2I (I); AD-110, NAD-21 (2); AD-119, NAD-22(1); AI)-141, NAD-llB (2); AD-148, NAD-12(1); AD-I49, NAD-15(3). Remarks: Steniaspis fossor is hardly separable from the older .S. sciitata, as noted by Hartman (1969). The present specimens have the skin papillae concentrated on posterior setigers. The number of setal bundles associated with the shield varies from nine to 14; the lower numbers are on two small specimens. Above a certain size level, it appears that the number of setal bundles varies independently of the size of the specimen, as remarked by Fauchald (1972a, p. 238). Occurrence: Both sides ol the Americas is shelf and bathyal depths. FAMILY OWENIIDAE RIOJA, 1917 Myriochele, species indeterminable Records: Al)-44. NAD-22 (no specimen recovered); AD-74, NAD-I4 (1). Remarks: Fhc siJctimen from station .A,D-44, previously identified as Myriochele heeri, was not recovered; the othei specimen is too poorly preserved to allow complete identification. 36 Allan Hancock Foundation Monograph No. 11 FAMILY Fl.ABELLIGERIDAE SAINT-JOSF.PH, 1894 KEY TO SPECIES la. Large simple hooks present in posterior parapodia Uncopherusa hifida b. Posterior parapodia without large simple hooks 2 2a. Body anteriorly inflated with a tapering posterior end; all setae capillaries of similar thickness . Diplocirrus micans h. Body more or less cylindrical; neurosetae distinctly thicker than the notosetae Brada pluribranchiata Brada pluribranchiata (Moore, 1923) Stylarioides pluribranchiata Moore, 1923, pp. 222-223. Brada pluribranchiata: Hartman, 1969, pp. 279-280, 7 figs; Fauchald, 1972a, p. 216. Record: AD-9, NAD-21 (2). Remarks: Brada pluribranchiata has a poorly developed setal cage. It is covered with large pajjillae with long, slender tips. The neurosetae are characteristically abruptly tapered along the mid-length, with a long, slender, usually flexible tip. Occurrence: Deep shelf and balhyal depths off southern California. Diplocirrus micans Fauchald, 1972 Diplocirrus micans Fauchald. 1972a, pp. 218-219, pi. 44, figs. a-e. Record: AD-7, NAD-16 (1). Remarks: In the pre.sent specimen the noto- and neurosetae are siinilar throughout the body, with the exception of the prolonged setae of the cephalic cage. The dorsum has a very sparse covering of papillae, most of which are so small as to be visible only microscopically. Occurrence: Bathyal depths off western Mexico. Pherusa, species indeterminable Records: AD-41, NAD-21 (1); AD-148, NAD-12 (fragment). Remarks: The specimen from station AI)-41 is large and complete, biu all setae are broken so it cannot be identified. The posterior fragment from AD-148 has the bidentate setae of certain species oi'Pherusa and the papular investment characteristic of members of this genus. Uncopherusa, new genus Cephalic cage formed from one setiger; setae include smooth capillaries, bifid hooded hooks, and greatly expanded, curved hooded hooks in the posterior end. Number of pairs of branchiae unknown; fjody partly covered with papillae. The genera resembling Pherusa were reviewed by Fauchald ( 1972a). Uncopherusa belongs to this group of genera, but differs sharply in the setal distribution and in the kind of .setae present. The very large hooks on the posterior end appear to be unique in the family. Hartman (1965, pp. 154-155, pi. 29) described the posterior end of what she thought might be a spionid. The structure of the setae closely resembles that found in Uncopherusa. Uncopherusa bifida, new species (Plate 6, Figs, e-h) Record: AD-89, NAD-22A (1, HOLOTYPE, Poly 0000). Description: The holotype and only known specimen is complete with 30 setigers. It is 4.5 mm long and 0.3 mm wide excluding setae. The body is cylindrical and somewhat thickened anteriorly and posteriorly. The first few segments are crowded, the median segments are elongated, and the posterior segments are again crowded. Dorsally, the peristoinium is produced into a somewhat flattened lip that projects over the retractable anterior end. The anus, which is terminal, is guarded by four short anal cirri. Deep-Water Polychaetes o« The first seliger (Fig. e) forms tiie cage. The iiotopocha are wholh dorsal and approach each other medially. The neiiro])odia are anteroveiilral lo the noto])odia and are directed forward. The iiotopodia are completely reduced in the second setiger. All other setigers have complete parapodia. The neuroixxlia have a distinct poslsetal lip which is especially pronounced in anterior parapodia. In setigers 3 to 5, the notopodia are .somewhat dorsal in position; oiher jjarapodia excejit the far jjosterior ones are strictly lateral. Far posterior parapodia are very low and are essentially marked only by the emergence of the setae. In this region the body (Fig. I) is somewhat pustulate, and the neuropodia are directed veiurad. and the notopodia dorsad. The anterior part of the body is encrusted with sand covering short ]iapillae; such encrustations and papillae are also present on the parapodial bases in the remainder of the body and on the whole posterior end. The notojjodial cage setae are short and conical and form a spreading fascicle over the posterior part of the peristomiiuri. The neuropodial cage setae are about one half as thick as the notopodial ones and at least three times as long, and they fonu a close fa.scicle on either side of the anterior end. Setigers 2 to 10 have only smooth, tapering ca])illary setae. Anterior fascicles are dense, including more than 20 setae per fasicle; [Kjsteriorly they become increasingly sparse. Each neuroseta is curved, nearly genicidate; notosetae are straight. The remainder of the notopodia contain these sinooth, straight, capillary setae in decreasing numbers so that in the last few segments only one or two setae are present. Neurojjodia from setiger 11 on contain one or a few bent, distallv bifid hooded hoks (Fig. g). These are usually accompanied by one or a few simple capillary setae. From setiger 24 they are replaced by a single, very large; double-cinved bidentate hooded hook (Fig. h) in each segment; the last hook, in setiger 30, is about one half the size of the others and is considerably less curved. Ihese hooks are all directed anterioventrad. Uncnpherusa bijida differs from other members of the family as indicated above for the genus. Orniirriire: One locality off central Oregon in bathyal depths. Details of the type locality can be found in the Station List. FAMILY FAUVELIOPSIDAE HARTMAN, 1971 KEY TO SPECIES la. Body with 41 setigers Fauveliopsis magna b. Body with 33 setigers Fauveliopsis glabra c. Body with 26 setigers Fauveliopsis armnta Faux'eliopsis armata, new species (Plate 7, Figs, g-i) Records: AD-6, NAD-17 (5, HOLOTYPE, Poly 1 160; PARATYPES, Poly 1161); AD-7, NAD-16 (8); AD-87, NAD-21 (2); AD-89, NAD-22A (I). Description: The holotype (Fig. g) is a complete specimen with 2(i setigers; it is 4.5 mm long and 1 nun wide at the widest. The body is generally cylindrical, increasing slowly in width to the posterior one-fourth; anterior and posterior ends are abruptly tapered. The anterior end (F'ig. i) has a short median lobe projecting barely beyond the rest of the anterior end; the peristomiiun forms a raised rim laterally on either side of this prostomial lobe. Fhe first seliger is wholly lateral, but the setae are directed anteriad. The first 9 or 10 setigers are marked by slightly raised, smooth welts on either side. The notopodia are more or less dorsal; the neuropodia are fully lateral. More posteriorly the parapodia are inarked only by the emergence of the setae. Each parapodium consists of a single spine and a capillary seta in each ramus; between the rami, usually close to the base of the notopodium, is a single large, pear-shaped papilla. Far anterior and posterior hooks (Fig. h) are strongly cinved and about twice as thick as those in median setigers. The capillary setae are relatively thick and strongly tapered in all setigers. The number of setigers in the fully adult specimens (eggs could be seen in some) varies between 23 and 26 in the present specimens; smaller specimens (less than 3 mm in length) have fewer setigers. The anterior enci of the body, including the first 10 setigers, has a rugose epithelium; the parapodial welts are smooth. Fhe rest of the body is completely smooth and nearly translucent. 38 Allan Hancock Foundation Monograph No. 11 The species oï F(ntveliol)sis can be grouped on the maximal number of setigers present in the adults. FauveUopsis cliallengcriae Mclnlosh (1922, pp. .5-7, pi. 2, figs. 1-8, pi. 3, fig. 2) and /•'. glabra (Hartman, 1960, pp. 129-130,pi. li.ligs. 1-2) have 3:5 setigers./•"./m'Tv/Wa 1 larlman (1971, p. 1122) has 28,/'". «iwr//« has 23 to 36 setigers. Finally,/•". hrei'ii (Hartman, 1965, p. 172), /•". Iiaitmani Levenstein ( 1970), 229, lig. 2a-b) andF. brattegardi Fauchald (1972b, p. 101, fig. 4a-b) have 16 setigers. One species, F. scabra Hartman and Fauchald ( 1971, pp. 117-118, pi. 17, figs, a-b), reported with varying numbers of setigers from 25 to 32 has been fbuiifl in material from the deep Adainic Ocean. I his species differs from the others in that the anterior rather than the posterior end is inflated, and tlie whole epithelium is rugose and warty. I he other species are generally smooth, only partially or rarely rugose, and tlie posterior end is inflatefl. FauveUopsis armata most closely resembles F. brevijwda in nimiber of setigers. In F. brevipoda (as F. brevis in Hartman, 1967, p. 123, pi. 37, tigs, a-b) the notopodial spines are slenderer than theneuropodialonesin all setigers; the spines are of similar thickness in both rami in F. ármala. Occurrence: Bathyal depths off central Oregon. Detailed data for the type locality can be foiuui in the Station List. FauveUopsis glabra (Hartman, 1960) Brada glabra Hartman, 1960, pp. 129-130, pi. 14, figs. 1-2. FauveUopsis glabra: Hartman. 1969. pp. 283-284, 2 figs. Records: AI)-74, NA1)-14 (3); ?AI)-110, NAI)-21 (1).' Remarks: FauveUopsis glabra has been distinguished above from other species from this area. The specimen from station AD-l 10 is fragmentary and cannot be completely identified. Occurrence: Southern California in deep shelf and bathyal depths. FauveUopsis magna, new s[)ecies Record: AD-9, NAD-21 (1, HOLOTYPE, Poly 1149). Description: The holotype and only known specimen is complete (41 setigers), and is 11 mm long and about 1.2 mm wide. Fhe body is taeniate and abruptly truncate anteriorly. It is the widest near the posterior one-fourth and tapers evenly to a short, unadorned pygidium. 1 he anal opening is terminal. All parapodia are biramous; the first two are directed forward; all others are lateral and have the two rami well separated. An ovate interramal papilla is present in all setigers. Each of the first 30 parapodia has a single curved spine and a single capillary seta in each ramus. The spines in the first two setigers are strongly curved and directed forward. The other spines are gently curved. The last 11 setigers have increasing niunbers of setae so that in the most ¡posterior setigers a total of three spines and two or three capillary setae may be present in each ramus. The surface of the whole animal is covered with very small, trim, papillae in scattered arrangement, making the surface appear velvety rather than glistening, as is usual in meiTibers of this genus. FauveUopsis magna is characterized by a very large luunber of setigers, 41, com])ared with the other species (see listing above in the discussion of /•'. armata) and by the presence of increasing numbers of setae in far posterior setigers. Occurrence: One locality in bathyal flejJths off central Oregon: complete station data can be found in the Station List. FAMILY SABELLARIIDAE JOHNS ION. 1865 Phalacrostemrna, species indeterminable Record: AD-6, NAD-l7 (1). Remarks: This specimen has the two prolonged ojjercular penducles characteristic of the genus Phalacrostemma. Fhe paleae are in a single row. The specimen is incomplete and cannot be further identified. Deep-Water Polychaetes 39 FAMILY PECTINARIIDAE QUATREFAGES, 1865 Cistenides, species indeterminable Record: Al)-6, NAD-17 (1). Rewurks: The present specimen is very poorly preserved and cannot be further identified. FAMILY AMPHARETIDAE MALMGREN, 1867 KEY 10 SPECIES la. At least some of the first setigers with needle setae 2 b. Needle setae absent 4 2a. Nuchal hooks present Melinna heterodonta b. Nuchal hooks absent 3 3a. Dorsal glandular ridge on setiger 4 present MeUnnampharete gracilis b. No glantlular ridge on setiger 4 Amelinna ohyssalis 4a. First abdominal segment with large muscidar dorsal valve Jngamjihicteis paleata b. First abdominal segment without dorsal modification 5 5a. Lower lip deeply crenulated 6 b. Lower li|:) smooth or irregularly wrinkled 7 6a. Paleae absent Amfihisamytha hiorulata b. Paleae |)resent Lysippe annectens 7a. Fifteen thoracic setigers present; of these, 12 are uncinigerous 8 b. Eighteen thoracic setigers present; of these, 14 are imcinigerous Amphicteis mucronata 8a. The third from last noiopodia elevated, with pilose setae Auobutlirus gracilis b. None of the notopodia elevated, all notosetae limbate and smooth Ampharete acutifrons Amelinna abyssalis Hartman, 1969 Amelinna abyssalis Hartman, 1969, pp. 533-534, 5 figs. Record: AD-155, NAD-24 (1). Remarks: Amelinna abyssalis has the anterior structure of a member of the Melinninae in that the first neinopodia contain needle-like small spines. Nuchal hooks are absent. The oral tentacles are of two kinds: a single giant tentacle and numerous smaller ones. Occurrence: Bathyal depths off southern California. Ampharete acutifrons {Gruhc. 1860) Amphicteis acutifrons Grube, 1860, pp. 109-1 10, pi. 5, fig. 6. Ampharete acutifrons: Hartman, 1969, pp. 537-538, 4 figs. Remarks: Ampharete acutifrons has a sharply pointed to bluntly triangular prostomium. Tiie branchiae are in.serted in two rows of two each and the imcini have teeth in double rows. Occurrence: Common in North Atlantic waters; reported as far south as Monterey Bay in the eastern Pacific Ocean from shelf to bathyl depths. Amphicteis mucronata Moore, 1923 Atnphicteis mucronata Moore, 1923, pp. 203-206; Hartman, 1969, pp. 547-548, 1 fig.; Fauchald, 1972a, pp. 284-285, pi. 58, fig. a. Records: AD-6, NAD-17 (2); AD-141, NAD-llB (2); AI)-149, NAD-15 (2). Remarks: The long muerons on the paleal setae distinguish this species from all related species in the Eastern Pacific area. The mid-superior part of the prostomium is usually small and three-pronged. Occurrence: Western Canada to western Mexico in shell and bathyal depths. 40 Allan Hancock Foundation Monograph No. 11 Amphisamytha hiocnlata (Moore, 1906) Samytim hiorulata Moore, 1906, pp. 253-253, pl. 12, figs. 52-53. Amphisamytha hiocnlata: Hartman, 1969, pp. 551-552, 5 figs. Records: AD-5, NAD-10 (1); AD-88, NAD-22A (1); AD-149, NAD-15 (31). Remarks: Amphisamytha hiocnlata has a deeply and distinctly crenulated lower lip. The only other am- pharetid from the eastern Pacific with this feature is Lysippe annectens (see below). The two are easily separated, since L. annectens has well-developed paleae and .4. hiocnlata lacks setae in this segment. Occurrence: Western C.anada and southern California in shelf and bathyal depths. Anobothrus gracilis (Malmgren, 1886) Ampharete gracilis Malmgren 1866, p. 365, pl. 26, fig. 75. Anobothrus gracilis: Hartman, 1969, pp. 553-554, 2 figs, (in part); Hartman and Reish, 1950, p. 42; Holthe, 1975, p. 25, fig. 2g-i, fig. 5a (not Hartman, 1965, p. 216, nor Hartman and Fauchald, 1971, p. 156). Records: AD-6, NAD-17 (1); AD-16, NAD-13 (2); AD-50, NAD-IO (2); AD-149, NAD-15 (17). Remarks: Anohothriis gracilis has the tenth pair of notopodia elevated and modified with pilose setae. The three anterior branchiae are in a straight line, and the fourth branchia is behind and medial to the others, leaving very little space between the two groups. The species has been confused with similar species with modified notopodial structures; the records from many areas are unreliable. Hartman (1965, p. 216) and Hartman and Fauchald (1971, p. 156) described specimens from the deep .Atlantic Ocean with 13 rather than 12 thoracic uncinigers; these specimens are no longer considered as belonging to this species. Hartman (1969, p. 553) described specimens from California with 13 uncinigers; this is a lapsus calami. California specimens have 12 luicinigers, as originally described for this species. Occurrence: North Atlantic areas and the Arctic Ocean basin; the eastern Pacific records are in part unreliable, biu the species has been found in California in bathyal areas. Jugamphicteis, new genus Ampharetin with 18 thoracic setigers, including the paleal segment; 14 of these uncinigers. Paleae well cleveloi)ed. Four pairs of branchiae arranged in two rows oí two branchiae each, only the rnedioposterior pair being distinctly associated with a segment. First abdominal segment with medially fused notopodial structures forming a valve between the thorax and the abdomen. Distinct notopodial rudiments present in some anterior abdominal segments. Thoracic uncini with teeth in single rows, abdominal ones with teeth in double rows. Genotype: Amphicteissibogae Caullery, 1944, pp. 82-83, fig. 66a-h. Additionally.y. paleata, described below, belongs to this genus. Two other species aï Amphicteis, A. vega Wirén, 1883, pp. 415-417, pl. 32, figs. 3-4, and A. vestis Hartman, 1965, pp. 215-216, pl. 46, show modified notopodial structures on anterior abdominal setigers. A. vega has expanded, lobate notopodia on the first three abdominal setigers; A. vestis has bilobed folióse noU)podia on the first abdominal setigers. Neither of these two species has the characteristic valve-like structure present in /. sihogae and /. pcdeata, and they are here not considered congeneric with the latter two species. Jugamphicteis paleata, new species (Plate 8, Figs, a-g) Record: AD-33, NAD-21 (1, HOLO'FYPE, Poly 1148). Description: The holotype and only known specimen is 37 mm long and 3.5 mm wide and consists of 33 setigers, of which 15 are in the abdoitien. Ihe body truncate anteriorly and widest near the anterior one-fifth; it tapers evenly posteriorly. The last two setigers (Fig. c) are fused with the pygidium and are distinctly inflated; two slender anal cirri are present. The prostomium (Fig. a) is divided into two distinct parts. The mediosuperior part is deeply divided anteriorly; each part is truncate. Posteriorly, the mediosuperior part is spiralled; the edge of each spiral is formed by glandular ridges. The prostomium is distinctly depressed between the two glandular promi- nences. Fhe interioposterior part ol the ¡irostomium is a rounded cushion. Dorsally, the peristomium forms a transversely wrinkled ring. On the ventral side, the mediosuperior part of the prostomium (Fig. b) Deep-Water Polychaetes 4| is visible anleriorly as two blunt prominences. The inferioposterior part forms two lateral cushions- it has been set off medially as a distinct glandular cushion. The ventral part of the peristomium forms the anterior and lateral lips. The anterior lip is deeply crenulated; the lateral lips are small but distinct cushions on either side of the mouth. I be anterior cdgeof the firstsetiger(paleal segment) forms the posteriorlip-it IS distinctly crenulated. In the roof of the mouth there is visible a distinct transverse ridge which curves postenoventrally at the corners of the mouth. The paleal segment is expanded and directed anteriad; the setae are supported by a low postsetal lip and a higher, rounded presetal lip. The first two poslpaleal setigers have small reduced notopodia hidden below the bases of the branchiae on both sides. The third post[)aleal setiger has a well-developed cylindrical, distally truncate notopodiuin resembling tho.se in the remainder of the thorax. Neuropodia are first present from the fourth postpaleal setigers; each thoracic neuropodium is a low, transverse fold Dorsally, the first abdominal setiger has a large valvular structure formed bv the fusion of a notopodial fold from either side (figs, d and g). In dorsal view, each fold can be identified as a cre.scent attached to the dorsolateral sides oí each segment and with a posteriorly directed free edge. This edge is produced into" about 15 digitate, blunt projections; these increasing evenly in size from the smallest, (located laterally at the base of the neuropodium) to the largest, (located dorsomediallv). The folds are muscular, and'the dorsomedial and jjosterior parts appear to be weakly sclerotinized. The two folds from either side are connected by a thin, nonmuscular membrane that is connected only to the two folds and leaves the medial portion of the dorstim free. Other abdominal segments have flattened, distally truncate neuropodia that become increasingly prolonged and narrowed in the far posterior setigers. The first three setigers following the modified one have distinct notopodial cirri; all others lack this features. The paleal setae arc golden, stout, and distally tapered to a fine curved tip. Notopodial setae are narrowly limbate. straight, and smooth. Thoracic uncini (fig. j) are flattened and have a distinct posterior end; each has a rounded base and has six teeth in a .single row along the cutting margin. The abdominal uncini (fig. e) are about half as big as the thoracic ones; each has a double row of five teeth on the cutting margin, so each uncinus has 10 teeth. The base is less curved, and the posterior bend is somewhat deeper than in the thoracic uncini. Tube was absent. Jugamphicteis paléala differs from the genotype,/, sibogae (Caullery, 1944, for complete references, see above) in the structure of the anterior end./, sihogoe has the anterior edge of the posterior part of the proslomium as four crescentic ridges, each crescent opening posteriorly./, palmta has two spiralled, ramshorn-like ridges forming the anterior edge of the posterior part of the prostomium. Occurrence: One locality in bathyal depths of f central Oregon; complete details can be found in the Staton List. Lysippe annectens Moore, 1923 Lysippe annectem Moore, 1923, pp. 201-202, pi. 17, figs. 11-13; Hartman, 1969, pp. 563-564, 6 figs.; Fauchald 197a, p. 299. ^ Records: AD-9, NAD-21 (3), AD-16, NAD-13 (2); AD-65, NAD-21 (2); AD-110, NAD-2I (!)• AD-148 NAD-12 (3). Remarks: Lysippe annectens was distinguishd in the key from the only other ampharetid known from Oregon with a crenulated lower lip. The four branchiae in Lysippe are in a single row and the uncini have several rows of teeth, both in the thorax and the abdomen. Occurrence: Southern California and western Mexico in shelf and bathyal depths. Melinna heterodonta Moore, 1923 Melinna cristata heterodonta Moore, 1923, pp. 212-213, pi. 17, fig. 25. Melinna heterodonta: Hartman, 1969, pp. 565-566, 6 figs.; Fauchald, 1972a, pp. 303-304. Records: AD-9, NAD-21 (2); AD-33, NAD-21 (I); AD-41, NAD-21 (1); AD-43, NAD-22 (1)- AD-44 NAD-22 (2); AD-61, NAD-17 (1); AD-65, NAD-21 (2); AD-86, NAD-21 (1); AD-89, NAD-22A (1); AD-9o' NAD-22 (1); AD-154, NAD-26 (1). Remarks: Melinna heterodonta has 18 thoracic setigers, including those with the needle setae anteriorly. The number of lobes in the transverse membrane is variable. The membrane usually has about a dozen lobes; most frequent numbers in the present collection are 11, 12, and 14. Occurrence: Western Mexico to central California in shelf and bathyal depths. 42 Allan Hancock Foundation Monograph No. 11 Melinnampharete gracilis Hartman, 1969 Melinnampharete gracilis Hartman, 1969, pp. 569-570, 7 figs.; Fauchald, 1972a, p. 308, pl. 63, fig. c. Rí-corí/i.- AD-6, NAD-17 (2); AD-149, NAD-15 (7). Remarks: MeUnnamjjharete granlis belongs to the Melinninae, in that it has needle setae m anterior setigers; it differs from Melinna in that it lacks the transverse membrane and nuchal hooks present in the latter genus. An inflated ridge is present across the dorsum of setiger 4. Occurrence: Western Mexico and southern California in bathyal depths. FAMILY TEREBELLIDAE MALMGREN, 1867 KEY TO SPECIES la. Lower lip forms a large, permanently everted proboscideal organ Artacama conifert h. Lower lip does not form a proboscideal organ 2 2a. Branchiae stalked; at least some anterior thoracic uncini long-handled 3 b. Branchiae sessile filaments; all uncini short-handled Thelepus setosus 3a. All thoracic uncini long-handled Pista fasciata b. Posterior thoracic uncini short-handled Pista ? cristata Artacama coniferi Moore, 1905 Artacama coniferi Moore, 1905, pp. 853-855, pl. 44, figs. 11-13; Hartman, 1969, pp. 585-586, 4 figs.; Hartman and Reish, 1950, pp. 43-44. Record: AD-86, NAD-21 (1). Remarks: Artacama coniferi has a large everted lower lip that is covered completely with fine papillae and resembles a pine cone in shape. The notopodial rudiments in the abdomen are large and folióse, as are the neuropodial lobes. Occurrence: From western Mexico to Alaska in shelf and bathyal depths. Pista ? cristata (C).F. Müller, 1776) Amphitrite cristata O.F. Midler, 1776, p. 216. Pista cristata: Hartman, 1969, pp. 615-616, 3 figs.; Holthe, 1975, p. 28, fig. 3k and 5n. Record: AD-Il, NAD-15 (1). Remarks: The present specimen agrees with P. cristata in the structure of the branchiae and the distribution of the long-handled and short-handled uncini. The specimen is rather small, and the lateral lappets are not as well developed as usual in this species. Pista fasciata (Grube, 1870) Terebella (Phyzelia) fasciata Grube, 1870, pp. 513-514. Pista fasciata: Hartman, 1969, pp. 621-622, 2 figs. Record: AD-88, NAD-22A (1). Remarks: The present specimen is small, but agrees with P. fasciata sensu Hartman (1969). Several distinct morphs have been subsumed under the name, and it is not clear how many distinct taxa may be involved in the current concept. Occurrence: Possibly cosmopolitan. Thelepus setosus (Quatrefages, 1865) Phenacia setosa Quatrefages, 1865, pp. 376-377. Thelepus setosus: Hartman, 1969, pp. 649-650, 6 figs. Record: AD-149, NAD-15 (1). Remarks: The sessile tufted branchiae are on three successive segments, with the first setae present from the second branchial segments. The uncini are in straight rows in all thoracic setigers. Occurrence: Cosmopolitan down to bathyal depths. Deep-Water Polychaetes FAMILY TRICHOBRANCHIDAE MALMGREN 1866 KEY TO SPECIES la. Branchiae cirriform o b. Branchiae lamellate ••^'•'•'•'.v;;:;;;;;;;;;:;;;::;;;;;;;; jVr;,¿;/i,¿,;-,^,;•¿¿ 2a. 1 wo pairs oí branchiae FiUhranchus rosens b. Three pairs of branchiae Artacamella hancockt Artacamella hancocki Hartman, 1955 Artacamella hancocki Hartman, 1955, pp. 49-50, pi. 3, flss. 1-6; Hartman, 1969 np 587-588 6 fie-s • Holthe, 1977, pp. 35-37, figs. 1-2. ' h - Record: AD-96, NAD-4 (2). Remarks: Artacamella hancocki was originally described in the family Terebellidae. Holihe (1977, pp. 35-37) pointed out that it strongly resembles Trichobranchus and that the similarity to Artacama empha.sized by Hartman (1955) is superficial. While Holthe's analy.sis as to direction of apomorphies and plesiomor- phies in the Hennigian terminology appears dubious without a more complete review of polychaete phylogeny, the general conclusion appears valid. . . Occurrence: Southern California in shelf and upper slope depths. Filihrnnchiis roseiis Malm, 1874 Filibranchns rosens Malm, 1874, pp. 99-100, pi. 1, fig. 9; Hartman, 1965, p. 227. Trichobranchus rosens: Hartman-Schröder, 1971, pp. 493-494. Record: AD-101, NAD-10 (1). Remarks: FiUhranchus roseas has two pairs of cirriform branchiae, and the first segment has large paired lateral lobes that extend toward the ventral side, but remain separated ventrally. Hartman (1965) kept the genus FiUhranchus separate from the genus Trichobranchus, with which it is usually synonymized. The separation appears to be justified. The type species is poorly known, and it is possible that both the specimens from the deep Atlantic reported by Hartman and the present specimen should be considered distinct species. Occurrence: Shallow water off Sweden and bathyal depths in the Atlantic Ocean off New England. Terebellides stroemii Sars, 1835 Terehellides stroemii Sars, 1835, pp. 48-50, pi. 13, fig. 31; Hartman, 1969, pp. 653-654, 7 figs.; Hartman and Reish, 1950, p. 44; Fauchald, 1972a, pp. 324-325. Records: AD-6, NAD-17 (4); AD-7, NAD-16 (1); AD-89, NAD-22A (1); AD-149, NAD-15 (1); AD-1.54 NAD-26 (1). Remarks: Terebellides stroemii has characteristically lamellate branchiae in which all the stems appear to have fused medially. The prostomium is a large folded membrane that can be partially withdrawn into the mouth; this makes individual specimens appear strikingly different. Occurrence: Possibly cosmopolitan. FAMILY SABELLIDAE MALMGREN, 1867 KEY TO SPECIES la. Thoracic neuropodial companion setae present 2 b. Thoracic neuropodial companion setae absent 3 2a. Last 8 to 12 setigers partially fused, forming a modified anal plaque Euchone analis b. All abdominal setigers free from one another, anal plaque absent Chone gracilis 3a. Collar nearly or completely covering the base of the tentacular crown on all sides Potamethus mucronatus b. Collar deeply cut away on the dorsal side, leaving bare the base of the tentacular crown and a large vascular coil Fabrisabella similis 44 Allan Hancock Foundation Monograph No. 11 Chone gracias Moore, 1906 Chone gracilis Moore, 1906, pp. 257-259, pl. 12, figs. 62-66; Hartman, 1969, pp. 665-666,4 figs,; Banse, 1972, pp. 470-472, fig. 4a-e. Records: AD-6, NAD-17 (3); AD-13, NAD-18 (3); AD-17. NAD-17 (1). Remarks: Chone gracilis has a ventrally indented collar and a very long tentacular crown combined with a very strongly glandularized epithelium. The setae specified as diagnostic by Hartman (1969) are, as remarked by Banse, not limited to this species. Occurrence: Alaska; other records appear doubtful. Euchone analis (Kröyer, 1856) Sahella analis Kröyer, 1856, p. 17. Euchone analis: Banse, 1972, pp. 482-483, fig. 9a-c. Records: ?AD-7. NAD-16 (3); AD-13, NAD-18 (1); AD-17, NAD-17 (1). Remarks: The present specimens agree with Euchone analis in the structure of the setae as well as in the number of setigers in the abdomen.The predepressional area has from 17 to 19 setigers and the depressed area has about 10 setigers. Occurrence: Greenland and Alaska. Fabrisabella similis Fauchald, 1972 Fahrisahella similis Fauchald, 1972a, pp. 329-330, pl. 69, figs. a-f. Record: AD-148, NAD-12 (1). Remarks: In Fahrisahella similis the collar is separated into two distinct parts, both ventrally and dorsally, and the collar is deeply cut away dorsally, leaving bare a large vascular coil on either side. The tentacular crown is very short and basally fused in the present specimen and resembles the condition of the specimen originally described. This may, in fact, represent the normal condition of the tentacular crown in this species, rather than a regeneration as suggested by Fauchald (1972a, p. 330). Occurrence: Western Mexico in bathyal depths. Potamethus mucronatus (Moore, 1923) Notaulax mucronata Moore, 1923, pp. 243-245, pl. 18, figs. 43-44. Potamethus mucronatus: Hartman, 1969, pp. 719-720, 4 figs. Records: AD-6, NAD-17, (3); AD-17, NAD-17 (2). Remarks: Potamethus mucronatus has long, straight, acicular uncini in the thoracic neuropoda; companion setae are present. Occurrence: Bathyal depths off southern California. Deep-Water Polychaetes 45 a. o CíO !), p. 29,as/^n: 7Z CL 00 c ra c r*! c» « Q£ u -' m cu í. • p :^-d ^ •y, OÍ c 4; - u u Ib £ O. i: < c i 2 "0 D.-0 Q I 1 CL 9 c !S 3 S Z 6' X :ra¿ le O E c/o O ^^ > CT> u 4•1 5 S c 11, t ^ Oí C U 1>4 "O X X c^ X 05 n¡ 7!' ín •Ji SM r^ O) ^ ^ ^ >, ?5 •/. 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" E ¡M' U. ¿_ C • 2 ^ 1 ai .g .5 1 • ^ '^ ^•# a> C bo 1972, p. 439, both as Stemasp •3 ribrnnchiata (Moore, 1923); th /o.Mor Stimpson, 1854; Hartma rey, 1972, p. 439. nformis délie Ciiiaje, 1841 ; Har 'osa (Rathke, 1843); Hartman p. 35. •ijlata (Treadweli, 1914); Hart Co (^' -- re S '<^ .:^ , •Ci. -C:- •S. -^ re ^ ^ '1 •S. c^ 0 U 0 re ir re • S -5 .0 10 2- ^ 1 x' X re CO ;; 1 CO CO U "^ CO ^ CO ^ 2 S 3 •^"l S "* ? s CO :§ 0 aq OQ 5^ H; S S; ^ t£ t2 it. -o Allan Hancock Foundation Monograph No. 11 i_- o CO a. a. o t^ • . c-Ci • r^ o o ^' • S ¡£ " ~ D- ^ ,n ,o • a. 9- . J: 00 ^ n • _ ^ o 3 C-.• t« í^ »o *" ^ ic • .^'ü TT ^ e-Cj: ^' 'S O-. - a¿ as _. eo SM cr. , ^ ^ t^ áü ,¿ -••" -z •" iJ 'f .¡£ CL Ä •- • :r-c -r 73 = ^ _ .33-p w^^ïc^i- PT3 UJ <; t~-u a Qj -y- '^ • :$ a í^' "í 2 42 ^ J3 £ ;5 s ^ ^ S 5 .-^ •_! ^ .- E " ~~" "^ • f^^^ -^ •' '•'t^ p IJ 1 ;- ^ **•. 5 u m c/î t^ 2 -S ri- ¿-^.£i s ë gîl^^' :c U s c o^•-" «¡i^ -2-f'-^ •~ tA :r ;;: ,Ä t^ . 5i S-1 -^^ -r5 '•• -^•^. n0 ^5 - u -^ >.. « c ~ o ., .-s • -^• rs.aS"^ .,._'~S'Ci-i:<: ^4;r- i:£_ s uO r; " TT O û • •^ • •y. Tf H '3 . ai ai ai a.• d. 00 S 0' •p - a. c 0" d. •5 cr> ^- CM' ^ß t'- -n "5 Si x' ai 3 t/3 a: -* ^' X E {/3 bu •"^ au c si V < ai Q a. < « , p- 43 a. 972, p. 2; Har u . .. d Reisi E !^ X U a 0 en j E j ,->u < u ai CQ CQ in 0 a¿ iH o 3: er. u E 1874; this pape 835; Hartman ; 818); Hartman 0); Carey, 1972 . 439; this pape d Berkeley, 194 1901; Hartman; almgren, 1866; amberlin, 1919: nson, 1901); Ha . - , 1776): Carey, 1 ^ )9; Hartman an 1^ r- iges, 1865); this 2 £ his paper, '*> 1• ."j S P j« 5 t so i = 1 -5 • S u. . a il £ S SS •S. 5 ai -5 .•s -g -5-A s >^ •2 0 -0 elongata Moore, 19 a medusa (Savigny, fasciata (Cirnbe, 18 cristata (O.K. Miillci parifica Berkeley ai 150, p. 45. mfiliitrite robusta (Jol '50. p. 44. •S. 5, 5 "^ ^ ce ~5' CTl d. ,S 1 5 5 en 1 S ^^ 1^ 'î: ¿j "-1 ^ Í iC í 5: S s '^ u. K f-. Deep-Water Polychaetes S3 STATION LIST AD-5, NAD-10, 22 June 1962: start 44°38.8' N, 124°54.9' VV; finish 44°38.4' N. 124°55.4' W, 600 m AD-6, NAD-17, 6 |iine 1963: start 44°33.5' N, I25°4.6' W; finish -, 2000 in AD-7, NAD-16, 13 August 1962: start 44°38,8' N, 125°12.r \V; finish -, 1800 m AD-9, NAD-21, 13 August 1962: start 44°36.4' N, 125°24.8' VV, finish -, 2800 m AD-11, NAD-15, 5 September 1962: start 44°39.2' N, 12.5°11.0' VV: finish -, 1600 m AD-13, NAD-18, ö September 1962: start 44°39.0' N, 125°13.2' VV; finish -, 2200 m AD-16,NAD-13, 4 October 1962: start 44°39.0'N, 12.5°10.0'VV; finish 44°38.0'N, 125°10.0'W, 1200 m AD-17, NAD-17, 4 October 1962: start 44°39.r N, 125°19.6'W; finish 44 °39.r N, 125°I8.8'VV, 2000 m AD-18, NAD-23, .T October 1962: start 44°39.r N, I26°31.0' VV; finish 44°36.5' N, 126°31.8' W, 2900 m AD-19, NAD-22, 6 October 1962: start 44°39.7' N. 126°0.03' VV; finish -, 2900 m AD-22. NAD-1 1, 4 December 1962; start 44°39.7' N, 124°58.0' VV; fniish 44°39.6' N, 124°58.0' W, 800 m AD-29, NAD-1 1, 24 January 1963?: start 44°39.3' N, 124°57.0' VV; finish -, 800 m AD-30, NAD-11 25 January 1963: start 44°39.3' N, 124°57.4' VV; finish -, 800 m AD-32, NAD-19, 25 January 1963: start 44°38.6' N, 125°20.1'VV; finish 44°37.6' N, 125°21.0' VV, 2400 m AD-33, NAD-21, 25 January 1963: start 44°39.0' N, 125°34.0' VV; finish 44°39.0' N, 125°33.2' VV, 2800 m AD-38, NAD-11, 27'April 1963: start -; finish -, 800 m AD-41, NAD-21, 1 June 1963: start 44°39.3' N, 125°34.2' VV; finish 44°40.9' N, I25°35.2' VV, 2800 m AD-42,NAD-21, IJune 1963: start 44°40.6'N, 125°35.5'VV; finish 44°43.3'N, 125°36.0'VV, 2800 m AD-43, NAD-22, I June 1963: start 44°40.0' N, 126°03.0' VV; finish 44°38.0' N, 126°03.0' VV, 2800 m AD-44, NAD-22, 1 June 1963: start 44°38.0' N, 126°03.0' VV; finish 44°38.5' N, r26°03.8' VV, 2800 m AD-50, NAD-10, 16 June 1963: start 44°32.9' N, 124°53.4' VV; finish -, 600 m AD-53. NAD-26, HAugust 1963; start 44°39.ñ'N, 127°.54.3'VV; finish 44°41.3'N, 127°51.«'VV, 2850 m AD-55, NAD-25, 15 August 1963: start 44°37.4' N, 127°28.0' VV; 2600 m AD-.56, NAD25, 15 August 1963: start 44°38.6' N, 127°28.2' VV; finish 44°38.8' N, 127°25.5' VV, 2600 m AD-59, NAD-11, 29 October 1963; start 44°40.0' X, 125°05.0' VV; finish -, 800 m AD-61, NAD-17(?), 30 October 1963: start 44°39.2° N, 125°1 1.0' VV; fini.sh -, 1400 in AD64, NAD-21, 28 December 1963: start 44°39.5' N, 125°35.9' VV; finish 44°37.9' N, 125°38.9' VV, 3000 m AD-65, NAD-21, 29 December 1963: start 44°42.0' N, 125°37.8' VV; finish 44°40.9' N 125°36.9' VV, 2750 m AD-68,NAD-6, 18Fehiuarv 1964: start 44°39.0'N, 124°33.1'VV; finish 44°38.4'N, 124°33.1'VV, 1.50m AD-74, NAD-14, 20 Febiuai v 1964; start •; finish-, 1400 m AD-86, NAD-2 1, 19 May 1964: start 44°38.5' N, 125°35.0' VV; finish 44°38.4' N, 125°.36.3' VV, 2865 m AD-87, NAD-21, 19 May 1964: start 44°39.8' N, I25°57.6' VV; finish 44°39.2' N, 125°.54.6' VV, 2800 m AD-88, NAD-22A, 20 May 1964: start 44°39.r N, 126°16.8' VV; finish 44°39.0' N, 126°17.8' VV, 2860 m AD-89, NAD-22A, 20 May 1964: start 44°38.5' N, 126°16.r VV; finish 44°38.r N, 126°16.4' VV, 2860 m AD-90, NAD-22, 21 March 1964: start 44°38.3'N, 126°01.0'VV; finish 44°38.9'N, 126°01.4'VV, 2860 m AD-96, NAD-4, 5 June 1964: start 44°44.6' N, 124°18.3' VV; finish 44°44.8' N, 124°18.4' VV, 100 m AD-97, NAD-4, 1.5 June 1964: start 44°44.5'N, 124°17.9'VV: finish 44°44.7'N, 124°17.9'VV, 100 ni AD-101, NAD-10, 16 June 1964: start 44°38.4', 124°.54.0' VV: finish 44°39.5' N, 124°.54.5' VV,600 m AD-105. NAD-12, IS'june 1964: start 44°38.4' N, 125°09.r VV; finish 44°39.r N, 125°09.7' VV, 1000 m AD-106, NAD-13, isjune 1964: start 44°37.6' N, 125°09.7' VV; finish 44°38.4' N, 125°09.6' VV, 1200 m AD-110, NAD-21, 11 August 1964: start 44°40.r N, 125°34.0' VV; finish 44°40.0' N, 125°35.0' VV, 2798 m AD-119, NAD-22, 13 January 1965: start 44°38.0'N, 126°02.2'VV; finish 44°38.0'N, 16°06.0'VV, 2800 m AD-139, NAD-24, 10 February 1965; start 44°39.4' N, 126°59,r VV; finish 44°39.8' N, 126°59.2' VV. 2800 m AD-141 NAD-1 IB, 8 April, 1965: start 44°29.7' N, 125°06.2' VV; finish 44°29.4' N, 125°06.2' VV, 1250 m AD-145, NAD-23, 9 April 1965: start 44°38.4' N, 126°30.2' VV; finish 44°38.3' N, 126°30.9' VV, 2800 m AD-148, NAD-12, 5 June 1965: start 44°4().7' N, 125°10.0'VV; finish 44°41.r N, 125°10.0 VV, 1000 m AD-149, NAD-15, 5'june 1965; start 44°4 1.2'N, 125°15.0'VV; finish 44°4 1.9'N, 125°15.1'W, 1600m AD-150, NAD-26, 21 October 1965: start 44°39.r N, 127°55.5' VV; finish 44°39.0' N, 17°56.6' VV, 2:560 m AD-154, NAD-26, 27 January 1966: start 44°34.5' N, 127°58.3' VV; finish 44°34.2' N, r27°57.9' VV, 1400 m AD-155, NAD-24, 26 March 1966: start 44°38.1' N, 126°59.7'W; finish 44°38.4' N, 127°03.0'2, 2700 m 54 Allan Hancock Foundation Monograph No. 11 PLATE 1 Aedicira (iregnnesis, new species a. Anterior end, dorsal view, lOOx Paraoni'lla (ihrancliidta. new species b. Anterior end, dorsal view, 95x c. Posterior end, dorsal view, 160x Cossura módica, new species d. Anterior end, dorsal view, 50x Deep-Water Polychaetes 55 C Allan Hancock Foundation Monograph No. 11 56 PLATE 2 Tharyx, near monüaris Hartman, 1960 a. Anterior end, lateral view, 50x b. Distal end of seta, median parapodiiim, 950x Minuspio minor, new species c. Anterior end, lateral view, 50x d. Neuiopodial hook, median setiger, 950x TravLsia oregonensis, new species e. Anterior end, dorsolateral view, lOx f. Posterior end, dorsolateral view, lOx Deep-Water Polychaetes 57 B ^^ Allan Hancock Foundation Monograph No. 11 PLATE 3 Mucihregma spinosa, new genus, new species a. Anterior end, dorsal view, 25x b. Anterior end, ventral view, 25x AncistrosyÜK breviceps, Hartman, 1963 c. Anterior end, pharynx everted, 50x Phnloe caeca Uschakov, 1950 d. Anterior end, dorsal view, 45x e. Notoseta, median parapodium, 385x f. Neuroseta, median parapodium, 385x Deep-Water Polychaetes 59 Allan Hancock Foundation Monograph No. 11 60 PLATE 4 Chaetoparifi careyi, new species a. Anterior end, dorsal view, 50x b. Anterior end, ventral view, 50x c. Diagram of posterior parapodium d. Seta, first setiger, 950x e. Seta, first setiger, 950x f. Seta, second setiger, 585x g. Seta, median setiger, 950x Deep-Water Polychaetes «i Allan Hancock Foundation Monograph No. 11 62 PLATE 5 Gyptis hians, new species a. Anterior end, cioisal view, 25x b. Notoseta, profile of median ¡jortion, 950x c. Notoseta, en face view ot median portion, 95()x d. Median parapodium, anterior view, 50x e. Ventralmost neuroseta, median parapodium, 950x Deep-Water Polychaetes 63 1 I i B C 64 Allan Hancock Foundation Monograph No. 11 PLATE 6 Paramphiname pacifun. new species a. Anterior end, dorsal view, 24x b. Median neuroseta, SS.'ix c. Tip of the above, 950x d. Median parapodium, anterior view, .50x Uncophenisa Infida, new genus, new species e. Anterior end, dorsal view, 50x f. Posterior end, dorsal view, 25x g. Hook trom setiger 20, 385x h. Hook from setiger 27, 385x Deep-Water Polychaetes 65 B C 66 Allan Hancock Founciation Monograph No. 11 PLATE 7 Nothria mixta, new species a. Anterior end, dorsal view. 25x b. Distal part of ventralmost composite hook, first setiger, 950x c. Distal part of median composite hook, first setiger, 950x d. First parapodium, anterior view, 53x e. Subacicular hook, median parapodium, 385x f. Pecdnate seta, median parapodium, 950x Fauveliopsis armata, new species g. Whole animal, dorsal view, 25x h. Far posterior notopodium, 385x i. Anterior end, lateral view, 50x Deep-Water Polychaetes 67 68 Allan Hancock Foundation Monograph No. 11 PLATE 8 Jugamphicteis paléala, new genus, new species a. Anterior end, dorsal view, 20x b. Anterior end, ventral view, setae omitted, 20x c. Posterior end, lateral view, 20x d. Junction between thorax and abdomen, lateral view, 20x e. Abdominal iincinus, yöOx f. Thoracic uncinus, 950x g. Junction between thorax and abdomen, dorsal view, 20x Deep-Water Polychaetes 69 70 Allan Hancock Foundation Monograph No. 11 LITERATURE CITED ANNENKOVA, N. 1934. Meeres Paraoniden in CLARK, R.B. and M.L.JONES. 1955. Two newXeph- fernen osten der USSR [In Rus.sian, German sum- ly\ (.\imelida, Polychaeta) from San Francisco Bay. mary]. Dokladv Akad. Nauk SSSR (C.R. Acad. Sei. J. Washington Acad. Sei. 45(5): 143-146, figs. 1-2. URSS). 3(8-9): 656-661, figs. 1-3. Dl I LEVSEN, H. 1917. Annelids. I. Vol. IV. pt. 4 m AUDOUIN, J.V. and H. MILNE EDWARDS. 1833. Fhe Danish Ingolf-Expedition. H. Hagerup, Classification des Annélides, et description de celles Copenhagen, 71 pp., 6 pis. qui habitent les côtes de la France. Ann. Sei. Nat., EHLERS, E. 1868.Die Borstenwiarmer (Annelida Paris. Ser. I, 29: 195-269, 388-412. (Chaetopoda) nach systematischen und anatomis- . 1834. Annélides. Vol. 2 in Recherches pour chen Untersuchungen. Wilhelm Engelmann, servir a l'histoire naturelle du littoral de la France, Leipzig, 748 pp, pis. 1-24. ou recueil de mémoires sur l'anatomie, la F.^BRICIUS, O. 1780. Fauna Groenlandica, system- physiologie, le classification et les moeurs des atice sislens, .Animalia Groenlandiae occidentalis animaux des nos côtes. Crochard Libraire, Paris, hactenus indagata, quoad nomen specificum, triv- 290, pp.. pis. 1-8. iale, vernacuhimque; Synonyma auctorum BANSE, K. 1972. Redescription of some species of plurium, descriptionem, locum, victum, genera- Chorif Kröyer and Euchone Malmgren and three tioncm, mores, usum, capturamque singuli; piout new species (Sabellidae, Polvchaeta). Fishery Bull. detegendi occasio fuit, maximaque parti secundum 70(2): 459-495. jjroprias observationes. Hafniae et Lipsiae, xvi -I- BANSE, K. and K.D. HOB.SON. 1974. Benthic er- 452 pp. rantiatepolychaetes of British Columbia and Wash- FAUCHALD, K. 1968. Onuphidae (Polychaeta) from ington. Bull. Fish Res. Bd. Canada. 185: HI pj), western Mexico. Allan Hancock Monogr. Mar. Biol. BERGSTRÖM, E. 1916.Die Polynoiden der Schwedis- 3: 1-82, pis. 1-12. chen Südpolarexpedition 1901-1903. Zool. Bidr., . 1969. A revision of six species of the Upp.sala. 4: 269-304, pis. 2-5. flavus-bidentatus group of Eunice (Eunicidae: BERKELEY, E. and C. BERKELEY. 1941. On a col- Polychaeta). Smith.son. Contrib. Zool. 6:1-15, figs lection of Polychaeta from southern California. 1-6. Bull. So. Calif. Acad. Aci. Vol. 40, pt. 1, pp. 16-60, 1970. Polychaetous annelids of the families pi. 5. Eunicidae, Lumbrineridae, Iphitimidae, Arabel- 1945. Notes on Polychaeta from the coast of lidae, Lysaretidac and Dorvilleidae from western western Canada. •III. Further notes on Syllidae Mexico. Allan Hancock Monogr. Mar. Biol 5- and some observations on other Polychaeta er- 1-335, pis. 1-27. rantia. Ann. Mag. Nat. Hist. Ser. 11, 12: 316-335. 1972a. Benthic polychaetous annelids from 1956. Notes on Polvchaeta from the east deep water off western Mexico and adjacent areas coast of Vancouver Island and from adajcent wat- in the eastern Pacific ocean. Allan Hancock ers, with a description of a new species oiAnciden. ]. Monogr. Mar. Biol. 7: 1-575, pis. 1-69. Fish. Res. Bd. Canada 13(4): .541-.546. . 1972b. Some polychaetous annelids from CAREY, A.G., Jr. 1972. Ecological observations on the the deep basins in Sognefjorden, western Norway benthic invertebrates froin the central Oregon Sarsia. 49: 89-106, figs. 1-4. continental shelf. Chapter 20, pi>. 422-443, in À.T. 1974. Deep-water errant polychaetes from Prater and D.L. Alverson, Fhe Columbia River Es- Hardangerfjorden, western Norwav. Sarsia. 57- tuary and Adjacent Ocean Waters: Bioenviron- 1-32, figs. 1-5. menlal Studies. 1976. The polychaete worms•definitions CAREY, A.G., Jr. and D.R. HANCOCK. 1965. An and keys to orders, families, and genera. Los An- anchor-box dredge for deep-sea sampling. Deep- geles County Museum of Natural History Science Sea Res. 12: 983-984. Series, 28, IX and 188 pp. CAULLERY, M. 1944. Polychetes .sédentaires de FAUVEL, P. 1923. Polychetes errantes. Faune de l'expédition du Siboga: Ariciidae, Spionidae, France. 5: 1-488, figs. 1-181. Chaetopteridae, Chlorphaemidae, Ophcliidae, 1927. Polychetes sédentaires. Addenda aux Oweniidae, Sabeariidae, Sternaspidae, Amphic- errantes, Archiannclides, .Myzostomaires. Faune de tenidae, Ampharetidae, Tercbellidae. Monogr. 24^ France. 16: 1-494, figs. 1-152. bis m Siboga-Expeditie. E.J. Brill, Leyden, 204 ijp 1936. Polychetes. Resultats du voyage de la 157 figs. Bélgica en 1897-99 sous le commandement de A. de CHAMBERLIN, R.V. 1919. The Annelida Gerlache de Goinery. J.-E. Buschmann, Anvers, Polvchaeta. Mem. Mus. Comp. Zool., Harvard Coll pp. 1-44, pi. I. 48: 1-514, pis. 1-80. FOSTER, N.M. 1971. Spionidae (Polychaeta) of the CLAPARÈDE, É. 1864. Cilanines z(M)tomiciues parmi Gulf of Mexico and the Caribbean Sea. 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Submarine canyons of southern Cali- 16: 157-170, figs. 1-9. fornia, Part HI. Systematics: Polychaetes. Allan GRUBE. A.-E. 1860. Beschreibung neuer oder wenig Hancock Pacific Expeditions. 27(3): 1-93, figs. 1-4. bekannter Anneliden. Arch. f. Naturgesch. 26(1): 1965. Deep-water benthic polychaetous an- 71-118, pis. 3-5. nelids off New England to Bermuda and other . 1870. Beschreibungen neuer oder weniger North-Atlantic areas. Allan Hancock Foundation liekannter von Hrn.Ehrenberg gesammelter An- Publications, Occas. Paper No. 28, 378 pp., pis. neliden des rothen Meeres. Akad. Wiss. Berlin, 1-52. Monatsber., Sitz, phvs.-math. Kl., 21 June 1869, pp. 1967. Polychaetous annelids collected by 484-521. the USNS Eltanin and Staten Island cruises, chiefly GUSTL'S, R.M. 1972. A species of the genus Eunice from Antarctic Seas. Allan Hancock Monogr. Mar. (Polvchaeta) from the Pacific Northwest Coast. Biol. 2: 1-387, pis. 1-51. Northwest Sei. 46(4): 257-269, figs. 1-7, tables 1-4. 1968, Atlas of the errantiate polychaetous HANCOCK, D.R. 1969. Bathyal and abyssal annelids from California. Allan Hancock Founda- polychaetes (annelids) from the central coast of tion, Univ. So. Calif., Los Angeles. 828 pp., numer- Oregon. Masters thesis, Oregon State University, ous illustrations. 121 pp. 1969. Atlas of the sedentariaie polychaet- HARTMAN. O. 1936. New species of Spionidae (An- ous annelids from California. Allan Hancock nelida Polvihaeta) from the coast of California. Foundation, Univ. So. Calif., Los Angeles. 812 pp., Univ. Calif. Publ. Zool. 41(6): 45-2, figs. 1-22. numerous illustrations. 1938. Descriptions of new species and new . 1971. Abyssal polychaetous annelids from generic records of polychaetous annelids from Cal- the Mozambicjue Basin off southeast Africa, with a ifornia of the families Glyceridae, Eunicidae, compendium of abyssal |3olychaetoiis annelids Stauronereidae, and Opheliidae. Univ. Calif. 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