Japanese Journal of Systematic Entomology, 26 (1): 11–14. June 30, 2020.

New Taxa described by Miyatake (1985)

Hiroyuki YOSHITOMI and Hiroyuki MURAKAMI Entomological Laboratory, Faculty of Agriculture, Ehime University, 3-5-7 Tarumi, Matsuyama, 790-8566 Japan. E-mail: [email protected]

Abstract We listed 22 taxa (4 genera and 18 species) described by Miyatake (1985).

Introduction Miyatake, 1985 Det. H. Murakami 2013”. Paratype: “HATSUNO, AMAMI IS. 27. VI. 1961 T. Shibata”, “♀”, “11- Miyatake (1985) described 22 new taxa (4 genera and 18 05” (OL), “PARATYPE Callimerus ryukyuensis Miyatake, species) in 6 families in a book’The Coleoptera of Japan in 1985 Det. H. Murakami 2013” (RL). Color, 3’. All the type series of them are preserved in Ehime University Museum (EUMJ), and the specimens figured in Callimerus ishigakiensis Miyatake, 1985: 155, pl. 25-6. the book were attached small orange labels. Unfortunately the Type specimen. Holotype (EUMJ, ser. No. 1288; Fig. 1D), descriptions were brief and short, and the information of type “[RYUKYU] Is. Ishigaki Banna 8. V, 1963 Y. Arita leg.”, series was completely absent. In the present paper, we listed the “HOLOTYPUS”, “HOLOTYPE Callimerus ishigakiensis new taxa described by Miyatake (1985) with the information of Miyatake, 1985 Det. H. Murakami 2013”. Paratype: “Ishigaki type series, photos and comments. Is. Loo-Choo 19. IV. 1963 H. OMURA”, “♀”, “11-06” (OL), “11-06” (OL), “ 右前フ節の第5 節だけ”, “PARATYPE List Callimerus ishigakiensis Miyatake, 1985 Det. H. Murakami The following list are the genera and species described 2013” (RL). by Miyatake (1985). Abbreviations for the label data are as follows: OL: orange label (this label indicates the number of Coptoclerus gressitti Miyatake, 1985: 157, pl. 25-18. plate in Miyatake (1985) attached by himself); RL: red label; Type specimen. Holotype (EUMJ, ser. No. 1290; Fig. 1E), MN: manuscript name. “16 VI 1962 BAEUBIBO”, “ASAHINA”, “11-18” (OL), “HOLOTYPE Coptoclerus gressitti Miyatake, 1985 Det. H. Ciidae Murakami 2014” (RL). Remarks. “Neoennearthron amamense Miyatake” was Remarks. Lim et al. (2015) redescribed the species based appeared in p. 283 (Miyatake, 1985), but this species was on two specimens of female without examination of holotype. not described. Kawanabe (2004) has already mentioned that This species is known from Japan and Korea (Lim et al., 2015) this name was simple mistake of Neoennearthron hisamatsui Miyatake, 1959. Genus Iwawakia Miyatake, 1985: 158. Iwawakia femorata Miyatake, 1985: 158, pl. 25-26. Syncosmetus reticulatus Miyatake, 1985: 280, pl. 46-2. Type specimen. Holotype (EUMJ, ser. No. 1319; Fig. 1F), Type specimen. Holotype (EUMJ, ser. No. 1318; Fig. 1A), “ 岩湧山 河内長野市 Osaka Japan 17. Jul. 1966 Osaka K. D. “[KYUSHU] Mt. Kurodake Ooita Pref. 25. VI. 1983 K. Kido Leg.”, “ キイロモモブトカッコウムシ Iwawakia femorata leg.”, “ コフキサルノコシカケ”, “No. 9”, “C-2” (OL), Miyatake, 1985”, “Holotype” (RL). “HOLOTYPE Syncosmetus reticulatus Miyatake, 1985 DET. Remarks. Sato (2002) redescribed the genus with Yoshitomi, 2020” (RL). examination of holotype. This species is known from Honshu and Shikoku, Japan (Sato, 2002; Waki & Murakami, 2020). Genus Xylographella Miyatake, 1985: 280. Xylographella punctata Miyatake, 1985: 280, pl. 46-4 Neohydnus aritai Miyatake, 1985: 154, pl. 25-2. Type specimen. Holotype (EUMJ, ser. No. 475; Fig. 1B), Type specimen. Holotype (EUMJ, ser. No. 1320; Fig. 1G), “(SHIKOKU) Mt. Tsurugi Tokushima Pref. 9. VIII. 1954 Coll. “[RYUKYU] Is. Ishigaki Takeda 2. V, 1963 Y. Arita leg”, M. Hirai”, “T”, “C-4” (OL), “HOLOTYPE Xylographella “11-02” (OL), “Neohydnus marginatus M. Miyatake” (MN), punctate M. MIYATAKE, 1985” (RL). Paratypes (EUMJ): 24 “HOLOTYPE Neohydnus aritai Miyatake, 1985 Det. H. exs., same data as for the holotype. Murakami 2014” (RL). Remarks. The genus and species were redescribed by Kawanabe and Miyatake (1996). Genus Platytenerus Miyatake, 1985: 158. Remarks. Murakami (2015) redescribed this genus with the Cleridae reevaluation of its systematic position by analysing phylogenetic Callimerus ryukyuensis Miyatake, 1985: 154, pl. 25-5. relationships. This genus is comprised of three species, P. Type specimen. Holotype (EUMJ, ser. No. 1287; Fig. 1C), castaneus (Kôno, 1930), P. iriomotensis Murakami, 2015, and P. “LOOCHOO Is. Okinawa Yona 22. IV. 1962 Y, Arita leg”, sichuanus Murakami et al., 2019 (Murakami et al., 2019). “HOLOTYPUS”, “HOLOTYPE Callimerus ryukyuensis Stigmatium ryukyuense Miyatake, 1985: 157, pl. 25-17.

Ⓒ Japanese Society of Systematic Entomology 12 Yoshitomi, H. and H. Murakami

Fig. 1. Holotypes and their labels of the species described by Miyatake (1985). A, Syncosmetus reticulatus; B, Xylographella punctate; C, Callimerus ryukyuensis; D, Callimerus ishigakiensis; E, Coptoclerus gressitti; F, Iwawakia femorata; G, Neohydnus aritai; H, Stigmatium ryukyuense; I, Xenorthrius elongatus; J, Litargus kyushuensis; K, Parabaptistes irregularis; L, Anoxacis flavomarginata. Scale bars = 1.0 mm.

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Fig. 2. Holotypes and their labels of the species described by Miyatake (1985). A, Anoxacis vittata; B, Allecula nipponica; C, Mycetochara elongata; D, Thymalus amamiensis. Scale bars = 1.0 mm.

Type specimen. Holotype (EUMJ, ser. No. 1289; Fig. 1H), Oedemeridae “(RYUKYUS) Yona Is. Okinawa July 23, 1968 M. Takagi Genus Anoxacis Miyatake, 1985: 407. leg.”, “11-17” (OL), “HOLOTYPE Stigmatium ryukyuense Anoxacis flavomarginataMiyatake, 1985: 407, pl. 69-13. Miyatake, 1985 Det. H. Murakami 2014” (RL). Type specimen. Holotype (EUMJ, ser. No. 1324; Fig. 1L), “13-18” (OL), “JAPAN: SHIKOKU Niyodo-river, Haruno- Xenorthrius elongatus Miyatake, 1985: 156, pl. 25-12. town, Kochi Pref. 2. VII. 1983 Y. Ito leg.”, “Collection of Type specimen. Holotype (EUMJ, ser. No. 1321; Fig. 1I), Ent. Lab. Ehime Univ.”, “♂”, “1”, “HOLOTYPE Anoxacis “(AMAMI IS.) Hatsuno 2. V. 1977 A. Oda leg.”, “11-12” (OL), flavomarginata Miyatake, 1985 DET. Yoshitomi, 2014” (RL). “HOLOTYPE Xenorthrius elongatus Miyatake, 1985 DET. H. Remarks. Akiyama (2000) redescribed the species without Yoshitomi, 2020” (RL). examination of holotype, and Švihla (2008) transferred this Remarks. This species is a secondary junior hononym species to the genus Asessinia Švihla, 1986. Akiyama (2018) of Xenorthrius elongatus (Corporaal, 1926), and is treated recorded this species from Taiwan. as Xenorthrius procerus Gerstmeier & Eberle, 2010 in the present time (Gerstmeier & Eberle, 2010). Anoxacis iriomotensis Miyatake, 1985: 407, pl. 69-15. Type specimen. Holotype (EUMJ, ser. No. 1325). We Mycetophagidae cannot find the holotype in EUMJ collection at the present Litargus kyushuensis Miyatake, 1985: 287, pl. 47-10. time. Type specimen. Holotype (EUMJ, ser. No. 1322; Fig. 1J), Remarks. Akiyama (2000) redescribed the species without “Chôja baru Mt. Kujû Ôita Pref. 17-18. VI. 1984 M. Tsuneoka examination of holotype, and Švihla (2008) transferred this leg.”, “PARATYPE Litargus multisignatus n. sp.” (MN), “9- species to the genus Asessinia Švihla, 1986. Akiyama (2018) 10” (OL), “HOLOTYPE Litargus kyushuensis Miyatake, 1985 recorded this species from Taiwan. DET. Yoshitomi, 2020” (RL). Remarks. Miyatake (1989) gave a short description and Anoxacis vittata Miyatake, 1985: 407, pl. 69-14. key to species. Type specimen. Holotype (EUMJ, ser. No. 1326; Fig. 2A), “Ohdomari Kagoshima Pref. 12. VII. 1969 M. Sakai Parabaptistes irregularis Miyatake, 1985: 288, pl. 47-23. leg.”, “13-19” (OL), “Collection of Ent. Lab. Ehime Type specimen. Holotype (EUMJ, ser. No. 1323; Fig. Univ.”, “HOLOTYPE Anoxacis vittata Miyatake, 1985 1K), “Mt. Kohtsu-zan Tokushima JAPAN”, “9-25” (OL), DET Yoshitomi, 2015” (RL). Paratype: 1 ex., “Is Amami “HOLOTYPE Parabaptistes irregularis Miyatake, 1985 DET. Ehime 小湊 Pref. 26. V. 1982 H. Kusunoki leg.”, “13-20” Yoshitomi, 2020” (RL). (OL), “Collection of Ent. Lab. Ehime Univ.”, “PARATYPE Remarks. Miyatake (1989) gave a short description and Anoxacis vittata Miyatake, 1985 DET Yoshitomi, 2015” (RL). key to species as “P. sp. (= irregularis Miyatake, 1985)”. This Remarks. Akiyama (2000) redescribed the species without species is a secondary junior homonym of Eulagius irregularis examination of holotype, and Švihla (2008) transferred this (Reitter, 1888), and is treated as Eulagius ussuriensis Nikitsky, species to the genus Asessinia Švihla, 1986. 1988 in the present time (Nikitsky, 2008).

June 30, 2020, JJSE 26 (1) 14 Yoshitomi, H. and H. Murakami

Asclera amamiama Miyatake, 1985: 405, pl. 68-25. 357‒377. Type specimen. Holotype (EUMJ, ser. No. 1327). We cannot Akita, K. and K. Masumoto, 2016. The Tenebrionid beetles of Japan. Mushi-Sha’s Iconographic Series of Insects 9. 302 pp., find the holotype in EUMJ collection at the present time. Mushi-Sha, Tokyo. Remarks. Mizota (1999) redescribed this species with Akiyama, H., 2000. Illustrated notes on the family Oedemeridae lectotype designation and figured. (Coleoptera) of Japan. Kanagawa-Chûhô, (132): 1‒53. (In Tenebrionidae Japanese.) Allecula nipponica Miyatake, 1985: 347, pl. 59-12. Akiyama, H., 2018. New records of Asessinia flavomarginata and A. Type specimen. Holotype (EUMJ, ser. No. 1; Fig. 2B), iriomotensis (Coleoptera, Oedemeridae) from Taiwan. Elytra, “(KYUSHU) Mt. Hiko Fukuoka Pref. VII-23, 1979 Y. New series, 8: 289‒290. SEIYAMA leg.”, “HOLOTYPE Allecula nipponica Miyatake” Asakawa, D., Y. Senda, and H. Yoshitomi, 2020. Revision of the genus Thymalus (Coleoptera, Cleroidea, Thymalidae) of (RL), “Ehime Univ. Collection”, “ 日本産ゴミムシダマシ大 Japan. Japanese Journal of systematic Entomology, 26: 69‒83. 図鑑（むし社）掲載標本 Akita et Masumoto 2016”. Gerstmeier, R. and J. Eberle, 2010. Revision of the Indo- Remarks. Akita and Masumoto (2015) redescribed Australian checkered beetle genus Xenorthrius Gorham, 1892 this species with examination of holotype, and Akita and (Coleoptera: Cleridae, Clerinae). Zootaxa, 2584: 1‒121. Masumoto (2016) showed the color figure of the holotype. Kawanabe, M., 2004. An illustrated guide to identification of ciid beetles of Japan, V. Coleopterists’ News, (146): 1‒5. (In Mycetochara elongata Miyatake, 1985: 348, pl. 59. 26. Japanese.) Kawanabe, M. and M. Miyatake, 1996. A redescription of Type specimen. Holotype (EUMJ, ser. No. 2; Fig. 2C), Xylographella punctata (Coleoptera, Ciidae), with description “[SHIKOKU] Mt. Tsurugi 11. VII. 1976 S. Hisamatsu”, of a new tribe. Elytra, 24: 125–130. “EHIME Univ. Collection”, “10-27” (OL), “HOLOTYPE Lim, J., Nomura, S., Lee, S., Y. Han, and B. Lee, 2015. Record Mycetochara elongata Miyatake” (RL), “Mycetochara of Coptoclerus Chapin from Korea with new species and a (Ernocharis) elongata Miyatake, 1985 Det. K. Akita, 2016”, note on the distribution of the genus (Coleoptera: Cleirdae). “ 日本産ゴミムシダマシ大図鑑（むし社）掲載標本 Akita Oriental Insects, 49 (3‒4): 223‒232. et Masumoto 2016”. Miyatake, M., 1985. Trogossitidae [pp. 147‒150, pl. 47], Cleridae Remarks. Akita and Masumoto (2014) redescribed this [pp. 151‒160, pls. 24, 25], Ciidae [pp. 279‒285, pl. 46], Mycetophagidae [pp. 285‒288, pl. 47], Tetratomidae [pp. species with examination of holotype. Akita and Masumoto 345‒346, pl. 59], Alleculidae [pp. 346‒348, pl. 59], Oedemeridae (2016) showed the color figure of the holotype. [pp. 401‒408, pls. 68, 69]. In: Kurosawa Y., S. Hisamatsu & H. Sasaji (eds.) The Coleoptera of Japan in Color, 3. 514 pp. Thymalidae Hoikusha, Osaka. (In Japanese, with English book title.) Thymalus amamiensis Miyatake, 1985: 149, pl. 24-9. Miyatake, M., 1989. A review of Japanese Mycetophagidae. Nature Type specimen. Holotype (EUMJ, ser. No. 1328; Fig. 2D), and Insect, 24(1): 8‒15. (In Japanese, with English title.) “Mt. Yuwan Amami-Oshima 19. IV. 1971 M. Sakai leg.”, “8- Mizota, K., 1999. A revision of the genus Indasclera in Japan (Coleoptera, Oedemeridae). Insecta matsumurana, new series, 06’” (OL), “HOLOTYPE Thymalus amamiensis Miyatake, 56: 69‒94. 1985 DET. Asakawa & Yoshitomi 2019” (RL). Murakami, H., 2015. The genus Platytenerus Miyatake, 1985 Remarks. See Asakawa et al. (2020). (Coleoptera: Cleridae: Neorthopleurinae), with description of a new species from Japan. Zootaxa, 3936 (4): 584–592. Acknowledgements Murakami, H., W.-I., Chou, and F.-M. Shi, 2019. A new species of Platytenerus Miyatake, 1985 from China (Coleoptera: First of all, we wish to express our hearty thanks to the Cleridae: Orthopleurinae). Japanese Journal of Systematic late Dr. Mutsuo Miyatake for his kind support during his life. Entomology, 25: 154‒157. We thank Dr. M. Sakai (EUMJ) and Dr. K. Mizota (Miyagi Nikitsky, N. B., 2008. Family Mycetophagidae Leach, 1815. University of Education) for giving us the information of the Pp. 51‒55. In: Löbl, I. & Smetana, A. (eds): Catalogue of type series. Palaearctic Coleoptera. Volume 5. Tenebrionoidea. 670 pp., Apollo Books, Stenstrup. Sato, M., 2002. Notes on the genus Iwawakia, with description References of a new species from central Japan. Japanese Journal of Akita, K. and K. Masumoto, 2014. New or little-known tenebrionid systematic Entomology, 8: 109–114. species (Coleoptera) from Japan (15) A revisional study of the Švihla, V., 2008. Family Oedemeridae Latreille, 1810. Pp. 45, genus Mycetochara Berthold (Alleculinae, Alleculini). Elytra, 353–369. In: Löbl, I. & Smetana, A. (eds): Catalogue of new series, 4: 213‒235. Palaearctic Coleoptera. Volume 5. Tenebrionoidea. 670 pp., Akita, K. and K. Masumoto, 2015. New or little-known tenebrionid Apollo Books, Stenstrup. species (Coleoptera) from Japan (16) Descriptions of eight Waki, Y. and H. Murakami, 2020. A new record of Iwawakia new species of genera Allecula and Borboresthes (Alleculinae, femorata Miyatake, 1985 from Shikoku. Sayabane, new Alleculini), and redescription of the known species with series, (36): 73. (In Japanese.) proposal for two new synonyms. Elytra, new series, 5: [Received: November 1, 2019; accepted: April 1, 2020]

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