(Diptera) from Southern Spain and Description of a New Trixoscelis Species from Northern Europe

' Trixoscclidae (Diptera) from Southern Spain and Description of a New Trixoscelis Species from Northern Europe

BY WALTER HACKMAN Zoological Museum, Jarnvagsgatan 13, Helsinki

Ent. scand. 1. 1970. 127-134.

Abstruct Seven species of Trixoscelis are recorded from o11 material from Finland. A key to the Palearctic Southem Spain. Three of them, T. psammophila, species of Trixoscelis is given. Some taxonomic T. Zyneborgi, and T. mendizabali, are described as problems in connection with the present material new. A fourth new species, T. siniilis, is described are discussed.

The Trixoscelidae collected by ihe Danish- ilar to T. obseureh Fallén but differing from Finnish expedition in 1966 to Southern Spain this species in the male genitalia and some (he provinces of Almería and Granada, see other characters. Frontal colour and chaeto- further Hackman (1969) and a collection of taxy as in obscicrella. Eyes oval, longest dia- this family brought together by Mr. J. R. m'etre almost horisontal. Genae broad as in Vockeroth (Canada Department of Agricul- obscurella (slightly more than half of tlie ture, Ottawa) in Granada in 1960 contained vertical diametre of the eye) and entirely al1 together 7 species including three new to white. Antennae as in obscurella. Thoracic science. The Palearctic species of the family chaetotaxy as in obscurella. Mesonotum grey- have not been revised since Czerny (1927) ish yellow, laterally pale grey. Scutellum and ,for the identification of the present brown, basa1 border pale grey. Plcura pale Spanish collections it was necessary to revise grey. h4esopleuron above with a much paler a number of species. During the revision of brown shade than in obscurellu. Wings with material in the collections of the Museum in pattern of dark confluent spots (fig. 3) more Helsinki two further new species were found contrasting than in oúscurellu but not sharply and as one of these (from Finland) is very outlined as in Zaeru Czerny. Halteres white. closely related to one of the new Spanish Legs greyish yellow, femora 1 and 111 with a species its description is included in this paper. grey dorsal shade in the apical third. Last The type material of Trixoscelis frontalis joint of tarsi darkened (mainly by an optical Fallén was kindly lent for study by Mr. P. 1. effect caused by the pilosity). Abdomen above Persson from the Riksmuseum, Stockholm. shiny black, the tergites without any lateral ' The following abbreviations of collector's grey pollinosity. Male genitalia as in figs. 1-2. narnes have been used below: L. L. - L. Lyne- - 9. Very similar to the male in size, colour borg, L. L. et al. - L. Lyneborg, O. Martin and and structure. Of the abdominal tergites only S. Langemark, J. V. - J. R. Vockeroth. five are visible, the last segments being re- The species in the collections from Southern tracted (dried specimens). The dark shades on Spain are the following: the femora are more contrasting than in the male. Trixoscelis psammophila nsp. . Holotype: cf, Spain; Almería: El Albufera Figs. 1-3 near Adra, 14 April 1966 (W. H.), preserved Description. - d'. Body length about z mm, in the collection of the Zoological Museum of wing length 2 mm. In general appearence sirn- the Helsinki University.

Fnt. srnnd. r:a. 1970

...... ># , . . ,; .. . .. ,. .. 1 128 Walter Eackman Trixoscrlidae (1

va E of Motril, 4 d 2 5 I 9, 14April; I cf I 9, I et al.); Rio Giiadalfeo, Or r8 April 1966 [L. L. et al.); Lanjaron, 600 m, I cf, 26 et1960 al.); [J. Granada,V.). 700 m, 4 Distribution. - Souther Africa and USSR: “Daiiric, Taken in Spain in Guadar? Sierra Morena (Ii. Lindbi from Soiithern Spain by (19093.

Trixoscelis sanctiferdinandi (Figs. 10-1 1) Material. - Almeria: P March; I d 3 0,g Marcl W.1%); Cabo de Gata, I 0 jr March 1966 (L. L.); Ric 1966 (W. H.).

.I I Distribution. Spain (t \’! 7 - Fernando near Cadiz) . Remarks. - The origina name is Ceomyza S. Ferdii Czerny & Strobl, rgog).

Trixoscelis frontdis (Fallén, wgs. 16-17, 30,351 (? canescens Loew, 1865; Material. - Almeria: Pec g March 1966 (L. L.); Rioj I cf, 11 March; 2 O, zo h and W. H.); Alhama del P I 9, ZI March 1966 (L. L.) Figs. 1-3. Trixoscelis psammophila n.sp., fig. I male genitalia, caudal view, fig. 2 phallic organ and hypandrium, fig. 3 wing. Figs. 4-5. T. obscureifa Fall. [Finland), fig. 4 renueva E of Motril, I d April; I April; epandrium and surstyli, caudal view, fig. 5 phallic organ. Figs. 6-7. T. iueta Becker 12 9, 14 I (L. L. et al.); Granada, 7c (Morocco), fig. 6 epandrium and surstyli, fig. 7 phallic organ. Figs. 8-9. T. approximata Locw (Spain), male genitalia in caudal and lateral view. 14 July 1960 (J. V.). Distribution. - Obviously in Europe but because c related species many recm Paratypes. - Almeria: Cabo de Gata, I 9, Sarothamnus tussocks on open sand in El are uncertain. 1 have 24 March 1966 (W.H.), I d‘ I 9, 31 March Albufera. SCC. 1966 (L. L.); El Albufera E of Adra, 5 Sweden (Fallén’s types), Fi: d‘ vorossisk, leg. A. Luther), ” z9 1966 (L. ” I4 Triroscelis approximata (Loew, 1865) a April 1966 (W. H.). - Granada: ‘Torrenueva da Foz, leg. H. Lindberg) (Figs. 8-9) Lindberg). T. canescens of E of Motril, 2 d I 0, 14 April 1966 (Martin & Langemark) . Material. - Almería: Cabo de Gata, I d‘, from Yugoslavia belongs her Remarks. - A characteristic species under 26 March 1966 (L. L.). - Granada: Torrenue- Remarks. - The male c

*.”....,-.._-”..-- .. . .._. - .- - , , ...... -.

_..., ,..: ...... , . -. . . - ~ __ . ._ ...... <_. - ..-:U_-.

Trixoscelidae (Diptera) from Southern Spain, and a New Trixoscelis Species 129

va E of Motril, 4 d' z 9, 12 April; 2 d separated from those of other related species I 9, 14 April; I d I 9, 17 April 1966 (L. L. by the following characters: Fore tibia dark et al.) ; Rio Guadalfeo, Orgiva, 300 m, I 9, except at the base, fore basitarsus yellow, third 18 April 1966 (L. L. et al.); Rio Lanjaron near basitarsus dark and moderately swollen (figs. Lanjaron, 600 m, I 3, 26 April 1966 (L. L. 30, 35). Further there are the characters of the et al.); Granada, 700 m, 4 d' 3 9, 14 July male genitalia (figs. 16-17). 1 have not seen 1960 (J. V.). the type of canescens Loew, but it is a female, Distribution. - Southern Europe, North and 1 have not been able to separate females Africa and USSR: "Dauria" (leg. Sahlberg). of frontalis Fall. and similis nsp. (probably Taken in Spain in Guadarama (R. Frey) and frontalis Collin nec Fallén). It seems more Sierra Morena [H. Lindberg) and reported likely that canescens is a synonym to the wide from Southern Spain by Czerny & Strobl spread frontalis. c 1909). Collin (1943) separates two close species from England under the names T. canescens and frontalis. The former has darkened front tibia iri the male and is obviously frontalis Trixosceiis sanctiferdinandi (Czerny, I 909) Fall. The later species with yellow front tibia (Figs. 10-1 1) in the male is probably similis. The presence Material. Almería: Pechina, 4 9, 5 or absence of brownish stripes on the meso- - notum is at least in Finnish specimens not a hgarch; I 3 9, g March 1966 (L. L. and d reliable character for separation of these close W. H.); Cabo de Gata, I 8, 24 March; z 9, species. 31 March 1966 (L. L.); Rioja, I 9, 12 April 1966 (W. H.). Distribution. - Spnin (type locality San Fernando near Cadiz). Reniarks. - The original spelling of the Triuoscelis' lyneborgi n.sp. name is Geoniyza S. Ferdinandi (Czemy in CFigs. 18-19, 29, 34) Czerny & Stiobl, 1909). Description. - Very similiar to T. similis P n.sp. described below. - c?. Body length 2.5 mm, wing length 2.3-2.7 mm. Orbitae and Trixoscelis frontaiis (Fallén, I 823) ocellar triangle pale grey, remaining median (Figs. 16-17, 30, 35) parts of frons yellow. Frontal chaetotaxy and - general shape of head as in frontaiis. Genae (? canescens Loew, 1865) narrow as in frontalis. Antennae as in fronra- Mateiial. - Almería: Pechina, I d' I 9, lis but at least in the 5 males in the present g March 1966 (L. L.); Rioja, I d, 7 March; material the third joint yellow only with a I d, 11 March; 2 9, 20 March 1gG6 (L. L. very faint trace of a dark spot on externa1 aiid W. H.); Alhama del Almería, 5 km W, surface. Mesonotum pale grey, entirely pol- alia, caudal vicw, fig. z I 9, 21 March 1966 (L. L.). - Granada: Tor- linose and at most with very faint trace of a ella Fall. (Finland), fig. 4 renueva E of hiIotri1, I d, 10 April; I 9, brownish sfripe pattern (of the type occurring igs. 6-7. T. laero Becker IZ April; I 9, 14 April; I d,17 April 1966 in the femalo). In the colour of the fore leg Figs. 8-9. T.approxiinnta (L. L. et al.); Granada, 700 m, z d' I 9, (fig. 29) distinctly differing from fronralis: 14 July 1960 (J. V.). Coxa yellow, femur dark grey, tibia entirely Distribution. - Obviously widely distributed greyish yellow, tarsus yellow, last joint slightly in Europe but because of confusion with darkened. Third femur dark, tibia yellow, .ussocks on opcn sand in E1 related spccies many records in the literature basitarsus very slightly swollen (fig. 34) and are uncertain. 1 have seen specimens from slightly darkened. Third tarsus yellow with Sweden (Fallén's types), Finland, USSR (No- apically darkened ultimate joint. Wings clear, vorossisk, leg. A. Luther), Portugal (Figueira not shaded along the costa. Cross-veins ta and roximata (Loew, 1865) da Foz, leg. H. Lindberg) and Corse (leg. H. tp in usual position (as in fronralis). Abdomi- Lindberg). T. canescens of Bequaert (1960) nal tergites dull greyish pollinose, seen from from Yugoslavia belongs here. Almería: Cabo de Gata, I 8, above broadly brownish with pale grey lateral i (Lo. I,.). - Granada: Torrenue- Rcmarks. - The male of T. fronralis is borders. Male genitalia distinctly different !%t. rcaiid. I:Z. 1970 9 p*. , i

Figs. 10-1 1. Trixoscelis sancriferdinandi Czerny, fig. 10 epandrium and surstyli, 'caudal view, fig. 11, phallic organ in profile. Figs. 12-13. T. proxima Séguy (Canary Is.), fig. 12 gonite, fig. 13 penis, ventro-caudal view. Figs. 14-15.T. baliogastra Czerny (Morocco), Fig. 23. Trixoscelis mendiz fig. 14 gonites and penis, fig. 15 penis, ventro-caudal view. Figs. 16-17 T. frontalis Fall. Loew, phallic organ, pro' (Finland), fig. 16, male hypopygium in profile, fig. 17, penis in ventro-caudal view. pedestris, penis, caudal 1 Figs. 18-19. T. Iyneborgi nsp. (Spain), fig. 18 epandrium and surstyli, caudal view, pedestris, fig. 28 T. MC: bT. proxiina. Figs. 32-36. fig. 19 gonites and penis. Figs. 20-21. T. similis nsp., fig. 20, penis in ventro-caudal fig. 33 T. mendizabaii, fig view, fig. 21, male genitalia in profile. Fig. 22. T. sexlineara Frey, gonite and penis.

Fnt. srnnii. i:a, 1970 Ent. scand. I: a. 1970

-. . . .. Trixoscelidae (Diptera) from Southern Spain, and a New Trixoscelis Species 131

16 26 I

.styli, caudal ' IS.), fig. 12 Fig. 23. Trixoscelis mendizabali n.sp. (Spain), niale genitalia, profile. Fig. 24. T.pedesrris . (Morocio), Loew, phallic organ, profile. Fig. 25. T. mendizabali, penis caudal view. Fig. 26. T. witalis Fall. pedestris, penis, caudal view. Figs. 27-31. First leg in Trixoscelis species, fig. 27 T. :auJal view. pedestris, fig. ZS T. mendizabali, fig. 29 T. lyneborgi, fig. 30 T. frotrtalis, fig. 31 2audal view, ' *T.proxima. Figs. 32-36. Third leg of male in Trixorcelis species, fig. 32 T. pedesrris, rentro-caudal fig. 33 T.mendizabali, fig. 34 T. lyncborgi, fig. 35 T. frontalis, fig. 36 T. proxima. :d penis.

Ent. siand. 1:2. 1970 132 Walter Iíackman Trixos

from those of the related species siinilis and pedestris can he recognized anioiig others by Kcy ro tlie Palca frontalis (figs. 18-19). - 9. Very similar to the colour characters of thc fore ieg (sec fig. 'I'hcre are sever: the females of frontalis and similis. Body 27 snd the key given later in this paper). author and thcrefo length ~7-2.9inm, wing length 2.7-3.0 mm. a coriipilation bas, The wing is slightly yellowish without shade 1927, Bequaert 196 along costa. Costa yellow seen from above (in Trixoscelis mendizabali n.sp frontalis dark). First tibia greyish yellow, IFigs. 23, 25, 28, 331 rarely somewhat darkened. First femur and I. Mesonotum blacl. tarsus dark as in the related species. Second This new species, closely related to T. pe- ywgoslavensis Bec and third legs yellow, only third femur darken- destris, is named to tlie honotir of Dr. Manuel - Mesonotum palc ed. First to sixth abdominal tergite greyish Mendizabal, director of the InstitEto de Acli- oclireous yellow pollinose, above brownish. matación in Almería. 2. Wings witli patti Description. - cf. Body length sbout 3 nim, posterior cross-ve Holotype: cf, Spain; Almería: Pechina, g wing length 2.7-2.9 mm. Colour and chaeto- - Wings clear or a March 1966 (L. L.), preserved in the Zoolog- taxy of the head as in pedestris. Width of Posterior cross- ical Museum of the University, Copenhagen. genae about 1/4 of the vertical diametre of brown ...... Paratypes. - Almería: Pechina, I 5 d', the eye. Antennae yellow. Mesonotiim dull 3. Mesociotuiii with March 1969 (W. H.), I I 9, data as the d' pale grey with indistinct brownish longitudinal s tripes ...... holotype; Rioja, 2 cf 2 9, 7 March; I d' stripes inside tlie dorsocentral row and along - Mesonotuni with 2 $2, g March; I II March; z 9, 12 cf, the intra-alar row. This pattern is usually very 4. Mesonotum red(' March 1966 (L. L.). .; Fuerteven- faint or absent in pedestris. Wings yellowish stripes . biliiicnrn tura: Betencuria, I arch 1949 (H. m as in pedesrris: The posterior cross-vein (tp) - Me:onatuin pali Lindberg); Chilegua, I 9, 4-14 March 1949 is in mendizabali longer than the last portion (H. Lindberg); Tenerife: Puerto de la Cruz, median stripe 1 of M3+4. Femur I dark grey, tibia 1 dark in and witli a shor. I 9, February (H. Lindberg). 2-4 1949 the apical third, basitarsus 1 darkened (fig. esch side along Three female specimens from Madeira: 28). Third leg (fig. 33) rather similar to that ...... Deserta Grande (1959, leg. H. Lindberg) a.re of pedestris (fig. 32). The ultimate tarsal (. very similar to T. lyneborgi but may possibly joint (legs 1-111) not so distinctly blackish at 5. Wings dark bri belong to tlie Madeiran species sexlineata apex as in pedesrris. Male genitalia (figs. 23, with only three Frey. 25) distinctly different from those of pe- clear spot reachi destris (fig. 24). - 9.Very difficult to sepa- beyond the mid rate from pedesrris. Body length 3-3.3 mm, other (subapica wing length 3.3-3.5 mm. Third antennal joint and M3+4 . . . . Trixoscelis pedestris (Loew, 1865) with a large dark spot on the externa1 surface - Wings not as a' (Figs. 24, 27, 32) (usually also in pedesrris). Mesonotum grey 6. Clear areas of tl with comparatively distinct brownish stript: sharply by the Material. - Almería: Pechina, 10 d' 3 9, pattern. Posterior cross-vein slightly longer spots. Mesonotu 5 March; 6 cf, g March 1966 (L. L. and W. than the last portion of M3+4 [probably a ...... H.); Alcazaba, 2 3, 18 April; I 9, 24 April reliable character for separating females of - Clear areas of 1966 (W. H.);Rioja, 4 O" 4 9, 7 March; 2 mendizabali from those of pedesrris). At cloud patterii n 9,9-11 March 1966 (L. L. and W. H.); I 9, least the apical third. of fore tibia blackish, almost clcar or 12 April 1966 (W. H.); Alhama del Almería first femur and tarsus black as in the related the cross-veins 5 km W, 2 2 17 March; I ?,ZI March d Q, species. Third femur somewhat darkened at 7. Wings broadiy i 1966 (L. L. and W. H.); Cabo de Gata, 3 d, apex, remaining parts of the leg yellow. Ab- extensively alor 24 March 1966 (L. L.) - Granada: Torrenueva, domen as in pedestris. to great exten! 3 cf I 0,10 April 1966 (L. L. et al.). Holotype. d', Spain; Almería: Pechina, 5 dark brown . . . Distribution. - Southern Europe, Asia March 1966 (W. H.), in the collection of the - Wings narrowt Minor, Canary Islands, North Africa. T. Zoological Museum of the Helsinki University. and with a t pedestris is rapported from Southern Spain by Paratypes. - Almería: Pechina, 2 0, same Abdominal tergi ' Czerny & Strobl (1909). data as the holotype; 2 9, g March 1966 (L. as in figg. 8-9 Remarks. - The females of this species are L.); Rioja, I 9, 11 March; 4 cf, 25 March 8. Abdominal ter; extremely difficult to separate from those of 1966 (L. L.). - Morocco: Ait Melloul near dark. Male genit the following species and there are uncertain Oued Sous, I cf 3 9, 27 Febriiary-2 March ...... cases in the above material. The males of 1961 (H. Lindberg). - Abdoininal ter¿

Ent.sroid. I:Z 1970 Ent. siand. 1:z. 1970 ...

...... -

Trixoscelidae (Diptera) frorn Southern Spain, and a New Trixoscelis Species 133

-d among others by Key ro rhe Palearctic specics of Trixoscelis. pollinose lateral borders. The sixth tergite he fore leg (see fig. There are several species not seen by the with a pale grey pollinose median spot. Scu- .- in this paper). author and therefore the key below is partly tellum unicolorous, pale. Male genitalia as in a compilation based on literature (Czerny figs. 4-5 ...... obscureila Fallén (Europe) 1927, Bequaert 1960). 9. Abdomen above shiny blackish brown .... 10 ip. ! - Abdomen not shiny above, dull grey with or without brownish pattern ...... 11 Mesonotum black, slightly greyish pruinose . . 10. Mesonotum ochreous without longitudinal *ly related to T. pe- yugoslauensis Bequaert (not seen, Macedonia) stripes ...... gentilis Frey (Canary 1s.) mour of Dr. Manuel Mesonotum pale grey, pale brownish or - Mesonotum pale grey with more or less ;ie Instituto de Acli- ochreous yellow ...... z distinct brown stripes along the dorsocentrals Wings with pattern of dark spots oí at least (inside of the rows) and along the intra alars. length about 3 mm, posterior cross-vein bordered with brown . . 3 Male genitalia as in figs. 10-11 ...... Colour and chaeto- i Wings clear or at most shadowed along costa...... sanctiferdinandi Czerny pedestris. Width of Posterior cross-vein never bordered with 11. Wing veins brown ...... 12 vertical diametre of brown ...... 9 - Veins pale yellowish, at most costa brownish w. Mesonotum dull Mesonotum with one more dark longitudinal 12. Abdominal...... ter9 rownish longitudinal stnpes ...... 4 brown'3 ntral along row and Mesonotum without stripe pattern ...... 5 lateral spodale genitaiia as figs. 14-15 ittern is usually very LT Mesonotum reddish grey witli two blackish bnliogastra ,Cz15- and N h Africa) ris. Wings yellowish stripes . bilincata Macquart (not seen, France) - Abdomi tergites not as above. ale genita- Aor cross-vein (tp) Mesonotum pale grey with a dark brown lia as . figs. 12-13 ...... han the iast portion median stripe broadening 011 the scutellum ...... proxima Séguy (Azores Canary 1s.) rey, tibia 1 dark iri and witli a sliort stripe in the presutural arca 13. Portion M, between th5dss-veins short, LIS 1 darkened (fig. each side along the intra-alars ...... +* not double m-ffie last portion of atlier similar to that ...... picncticornis Becker The ultimate tarsal (North Africa and sanarv M3t4...... 14 h) - Portion of between the cross-veins at iistinctly blackish at Wings dark brown by conflutnt spots and M,tz - least double as long the last portion of z genitalia (figs. 23, with only tiiree larger clear areas: one long as '5 from those of pe- clear spot reaching from the base of the wing M3t4...... 'ry difficult to sepa- beyond the middle and situated like the two 14. Genae broad, about half of the vertical dia- y length 3-3.3 mm, other (subapical) clear spots between R4t5 metre of tiie eye. Third antennal joint dark Third antennal joint and M, ...... margindla Fallén (Europe) ...... oedipus Becker (not seen, Tibet) the externa] surface Wings not as above ...... 6 - Genae narrow. Third antennal joint yellow, )). Mesonotum grey Ciear areas of tlie wing separatrd more or less at most with a brown spot at the base of nct brownish stripf: sharply by the brown patterii as four large arista ...... undescribed sp. from kan. vein slightly longer spots. Mesonotum ochreous ...... 15. Males ...... 16 M3t4 (probably a ...... laeta Becker (North Africa) - Fernales ...... 22 parating females of Clcar areas of the wing confluent, browri r6. First tibia almost entirely dark grey. Costal of pedestris). At cloud pattern not sharply outlined, or wings cell slightly shadowed. Costa dark seen from fore tibia blackish, almost clear only with brown border along above (optical effect). Male genitalia as in ick as in the related the cross-veins ...... 7 figs. 16-17 ...... frotiralis Fal!í.n riewhat darkened at Wingc broadly clouded at the cross-veins and - First tibia dusky yellow, at ni- in the the leg yellow. Ab- exrciisively alvng costa. Abdominal tergites apical third. Costa usually yellow seen from to great extent or eritirely shiny black or above ...... 17 4lmcría: Pechina, 5 dark brown ...... 8 17. First tibia rntirely dusky yellow ...... 18 the collectinn of the Wings narrower clouded at ihe cross-veins - Apical third or fourth of first tibia dark . , 20 Helsinki University. and with a brownish shade along costa. 18. Third basitarsus and the following joint Pechina, 2 $2, sanie Ahdorninal tergites duli above. Male genitalia brown. Male genitalia as in fig. 22 ...... ?, 9 March 1966 (L. as in figs. 8-9 ...... approximata Loew ...... sexlineara Frey (Madeira) .ch; 4 8, 25 Marcti Abdominal tergites shiny black. Scutcllum - Third basitarsus at most very slightly darker J: Ait Melloul mar dark. Male genitalia as in figs. 1-2 ...... than third tibia. The second tarsal joint of 7 February-2 March ...... psamrnoplrila nsp. third leg not darker than the following joints Abdominal tergites r-5 shiny brown with ...... 19

Ent.scand. 1:2. 1971 Ent. scsnd. 1:2. igp .? .,

134 Walter Hackman

19. Male genitalia as in figs. 18-19 ...... head as in fronrulis and the above mentioned ...... lyneborgi nsp. related species. Mesonotum pale grey with - Male gcnitalia as in figs. m-21- n.sp. faint brownish stripe pattern [of same type 20. Basitarsus of first leg dusky yellow. End as in frontalis), sometimes the stripes are joints of al1 tarsi blackish in the apical half. absent. Wing veins yellow, costa in some Genitalia as in figs. 24-26 .... pedestris Loew specimens dark seen from above. End portion - Basitarsus of first leg dark ....:...... 21 of M3+4 longer than the posterior cross-vein. 21. First tarsus entirely dark ...... First tibia entirely dusky yellow brown seen ...... coei Bequaert [not seen, Yugoslavia) from above. Second and third legs (except - Second and following joints of first tarsus coxae) yellow. Abdominal tergites grey or dusky yellow, only basitarsus and the apical sometimes medially brownish. Male genitalia half of the end joint darkened. Male genitalia (figs. 20-21) distinctly different from those as in figs. 23, 25 ...... mendizubaíi nsp. of lyneborgi and sexlineata. - 0.The females 22. Costa dark seen from above. Costal cell associated with the males described above are slightly shaded. First leg except of coxa en- so similar to those of fronrulis that 1 have not tirely blackish grey ...... been able to found any certain character for ...... frontulis Fallén and similis nsp. separation. - Costa not dark and costal cell not shaded. Holotype: d, Finland; Ob: Hailuoto (R. First tibia entirely dusky yellow or dark in Frey). the apical half ...... 23 Paratypes from Finland; Al: Jomala, I d' 23. First tibia entirely dusky yellow ...... (R. Frey); Sund, I d' (R. Frey); Finstrom, ...... lyneborgi nsp. 2 d' (R. Frey); N: Masaby, 8 d, 21 July - First tibia dark in the apical half ...... 24 1956 (W. H.); Om: Nykarleby, I d' (R. 24. Posterior cross-vein slightly longer than the Frey). last poriion of M3 ...... metidizabali nsp. The females probably belonging to this - Posterior cross-vein shorter thaii the last por- spccies are taken in N: Masaby, I 9 (W. H.), tion oF M3+,...... 25 Om: Jakobstad, I 9 (R. Frey) and Ob: Hai- 25. End joints of second and third tarsi blackish iuoto, 4 9 (R. Frey). T. frotituiis is (based on in the apical halF ...... pedestris Loew the captures of males) not known from the - End joints of tarsi not as above ...... 'provinces Om and Ob and al1 certain records ...... coci Bequaert are from Soutliern Finland.

The rather dubious species puberulu Zet- terstedt included by Czerny (1927) in Trixo- References scelis has been omitted from the key. sa- T. BEQUAERT,M., 1960. Contribution 5 I'étude des bulicolu Frey (1958) described from Fuerte- Diptkres de Yougoslavie: Trixoscelinae (Helo- ventura (Canary 1s.) is most probably punc- myzidae). - Bull. Ann. Soc. ent. Belg. 94. ricornis Becker. A species fiom the Canary Bruxellcs. 59-68. 1s. has bcen iriterpreted here as proxinta Séguy COLLIN, J. E., 1943. The Britisli species of Helo- describcd from the Azores but 1 have not seen myzidae (Diptera). - Ent. mon. Mag. 79. Lon- the type material. The new species similis don. 234-251. - mentioned repeatedly in this paper is de- CZERNY,L., 1927. Fani. 53. Helomyzidae + Trixo- scribed below : scelidae + Chiromyidae. In LINDNER:Die Fliegen der palaearktischen Region. 5. Stuttgart. 1-56. CZERNY,L. & STROBL,G., rgog. Spanische Dipteren Trixoscelis similis n.sp. . 111. Beitrag. - Verh. 2001.-bot. Ges. Wicn. 59. [Figs. 20-21) Wien. 121-301.

Ent. sranil. ~:i.IWO