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Diversity Partitioning During the Cambrian Radiation
Diversity partitioning during the Cambrian radiation Lin Naa,1 and Wolfgang Kiesslinga,b aGeoZentrum Nordbayern, Paleobiology and Paleoenvironments, Friedrich-Alexander-Universität Erlangen-Nürnberg, 91054 Erlangen, Germany; and bMuseum für Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity at the Humboldt University Berlin, 10115 Berlin, Germany Edited by Douglas H. Erwin, Smithsonian National Museum of Natural History, Washington, DC, and accepted by the Editorial Board March 10, 2015 (received for review January 2, 2015) The fossil record offers unique insights into the environmental and Results geographic partitioning of biodiversity during global diversifica- Raw gamma diversity exhibits a strong increase in the first three tions. We explored biodiversity patterns during the Cambrian Cambrian stages (informally referred to as early Cambrian in this radiation, the most dramatic radiation in Earth history. We as- work) (Fig. 1A). Gamma diversity dropped in Stage 4 and de- sessed how the overall increase in global diversity was partitioned clined further through the rest of the Cambrian. The pattern is between within-community (alpha) and between-community (beta) robust to sampling standardization (Fig. 1B) and insensitive to components and how beta diversity was partitioned among environ- including or excluding the archaeocyath sponges, which are po- ments and geographic regions. Changes in gamma diversity in the tentially oversplit (16). Alpha and beta diversity increased from Cambrian were chiefly driven by changes in beta diversity. The the Fortunian to Stage 3, and fluctuated erratically through the combined trajectories of alpha and beta diversity during the initial following stages (Fig. 2). Our estimate of alpha (and indirectly diversification suggest low competition and high predation within beta) diversity is based on the number of genera in published communities. -
Decoupled Evolution of Soft and Hard Substrate Communities During the Cambrian Explosion and Great Ordovician Biodiversification Event
Decoupled evolution of soft and hard substrate communities during the Cambrian Explosion and Great Ordovician Biodiversification Event Luis A. Buatoisa,1, Maria G. Mánganoa, Ricardo A. Oleab, and Mark A. Wilsonc aDepartment of Geological Sciences, University of Saskatchewan, Saskatoon, SK, Canada S7N 5E2; bEastern Energy Resources Science Center, US Geological Survey, Reston, VA 20192; and cDepartment of Geology, The College of Wooster, Wooster, OH 44691 Edited by Steven M. Holland, University of Georgia, Athens, GA, and accepted by Editorial Board Member David Jablonski May 6, 2016 (received for review November 21, 2015) Contrasts between the Cambrian Explosion (CE) and the Great shed light on the natures of both radiations. Because there is still Ordovician Biodiversification Event (GOBE) have long been recog- controversy regarding Sepkoski’s nonstandardized curves of nized. Whereas the vast majority of body plans were established Phanerozoic taxonomic diversity (13–15), a rarefaction analysis as a result of the CE, taxonomic increases during the GOBE were was performed in an attempt to standardize diversity data. The manifested at lower taxonomic levels. Assessing changes of ichno- aims of this paper are to document the contrasting ichnodiversity diversity and ichnodisparity as a result of these two evolutionary and ichnodisparity trajectories in soft and hard substrate com- events may shed light on the dynamics of both radiations. The early munities during these two evolutionary events and to discuss the Cambrian (series 1 and 2) displayed a dramatic increase in ichnodi- possible underlying causes of this decoupled evolution. versity and ichnodisparity in softground communities. In contrast to this evolutionary explosion in bioturbation structures, only a few Results Cambrian bioerosion structures are known. -
Triassic- Jurassic Stratigraphy Of
Triassic- Jurassic Stratigraphy of the <JF C7 JL / Culpfeper and B arbour sville Basins, VirginiaC7 and Maryland/ ll.S. PAPER Triassic-Jurassic Stratigraphy of the Culpeper and Barboursville Basins, Virginia and Maryland By K.Y. LEE and AJ. FROELICH U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1472 A clarification of the Triassic--Jurassic stratigraphic sequences, sedimentation, and depositional environments UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON: 1989 DEPARTMENT OF THE INTERIOR MANUEL LUJAN, Jr., Secretary U.S. GEOLOGICAL SURVEY Dallas L. Peck, Director Any use of trade, product, or firm names in this publication is for descriptive purposes only and does not imply endorsement by the U.S. Government Library of Congress Cataloging in Publication Data Lee, K.Y. Triassic-Jurassic stratigraphy of the Culpeper and Barboursville basins, Virginia and Maryland. (U.S. Geological Survey professional paper ; 1472) Bibliography: p. Supt. of Docs. no. : I 19.16:1472 1. Geology, Stratigraphic Triassic. 2. Geology, Stratigraphic Jurassic. 3. Geology Culpeper Basin (Va. and Md.) 4. Geology Virginia Barboursville Basin. I. Froelich, A.J. (Albert Joseph), 1929- II. Title. III. Series. QE676.L44 1989 551.7'62'09755 87-600318 For sale by the Books and Open-File Reports Section, U.S. Geological Survey, Federal Center, Box 25425, Denver, CO 80225 CONTENTS Page Page Abstract.......................................................................................................... 1 Stratigraphy Continued Introduction... .......................................................................................... -
Geological Timeline
Geological Timeline In this pack you will find information and activities to help your class grasp the concept of geological time, just how old our planet is, and just how young we, as a species, are. Planet Earth is 4,600 million years old. We all know this is very old indeed, but big numbers like this are always difficult to get your head around. The activities in this pack will help your class to make visual representations of the age of the Earth to help them get to grips with the timescales involved. Important EvEnts In thE Earth’s hIstory 4600 mya (million years ago) – Planet Earth formed. Dust left over from the birth of the sun clumped together to form planet Earth. The other planets in our solar system were also formed in this way at about the same time. 4500 mya – Earth’s core and crust formed. Dense metals sank to the centre of the Earth and formed the core, while the outside layer cooled and solidified to form the Earth’s crust. 4400 mya – The Earth’s first oceans formed. Water vapour was released into the Earth’s atmosphere by volcanism. It then cooled, fell back down as rain, and formed the Earth’s first oceans. Some water may also have been brought to Earth by comets and asteroids. 3850 mya – The first life appeared on Earth. It was very simple single-celled organisms. Exactly how life first arose is a mystery. 1500 mya – Oxygen began to accumulate in the Earth’s atmosphere. Oxygen is made by cyanobacteria (blue-green algae) as a product of photosynthesis. -
Early Cretaceous (Albian) Decapods from the Glen Rose and Walnut Formations of Texas, USA
Bulletin of the Mizunami Fossil Museum, no. 42 (2016), p. 1–22, 11 fi gs., 3 tables. © 2016, Mizunami Fossil Museum Early Cretaceous (Albian) decapods from the Glen Rose and Walnut formations of Texas, USA Carrie E. Schweitzer*, Rodney M. Feldmann**, William L. Rader***, and Ovidiu Fran㶥escu**** *Department of Geology, Kent State University at Stark, 6000 Frank Ave. NW, North Canton, OH 44720 USA <[email protected]> **Department of Geology, Kent State University, Kent, OH 44242 USA ***8210 Bent Tree Road, #219, Austin, TX 78759 USA ****Division of Physical and Computational Sciences, University of Pittsburgh Bradford, Bradford, PA 16701 USA Abstract Early Cretaceous (Albian) decapod crustaceans from the Glen Rose Limestone and the Walnut Formation include the new taxa Palaeodromites xestos new species, Rosadromites texensis new genus, new species, Karyosia apicava new genus new species, Aetocarcinus new genus, Aetocarcinus muricatus new species, and the new combinations Aetocarcinus roddai (Bishop, 1983), Necrocarcinus pawpawensis (Rathbun, 1935) and Necrocarcinus hodgesi (Bishop, 1983). These two formations have yielded a much less diverse decapod fauna than the nearly coeval and proximally deposited Pawpaw Formation. Paleoenvironment is suggested as a controlling factor in the decapod diversity of these units. Key words: Brachyura, Nephropidae, Dromiacea, Raninoida, Etyioidea, North America Introduction deposited in the shallow waters of a broad carbonate platform. Deposition occurred on the southeastern flank of Late Early Cretaceous decapod faunas from the Gulf the Llano Uplift and, on the seaward margin to the Coastal Plain of North America have been well reported northwest, behind the Stuart City Reef Trend. Coral and and described since the early part of the twentieth century rudist reefs, algal beds, extensive ripple marks, evaporites, (Rathbun, 1935; Stenzel, 1945). -
Asteroid Impact, Not Volcanism, Caused the End-Cretaceous Dinosaur Extinction
Asteroid impact, not volcanism, caused the end-Cretaceous dinosaur extinction Alfio Alessandro Chiarenzaa,b,1,2, Alexander Farnsworthc,1, Philip D. Mannionb, Daniel J. Luntc, Paul J. Valdesc, Joanna V. Morgana, and Peter A. Allisona aDepartment of Earth Science and Engineering, Imperial College London, South Kensington, SW7 2AZ London, United Kingdom; bDepartment of Earth Sciences, University College London, WC1E 6BT London, United Kingdom; and cSchool of Geographical Sciences, University of Bristol, BS8 1TH Bristol, United Kingdom Edited by Nils Chr. Stenseth, University of Oslo, Oslo, Norway, and approved May 21, 2020 (received for review April 1, 2020) The Cretaceous/Paleogene mass extinction, 66 Ma, included the (17). However, the timing and size of each eruptive event are demise of non-avian dinosaurs. Intense debate has focused on the highly contentious in relation to the mass extinction event (8–10). relative roles of Deccan volcanism and the Chicxulub asteroid im- An asteroid, ∼10 km in diameter, impacted at Chicxulub, in pact as kill mechanisms for this event. Here, we combine fossil- the present-day Gulf of Mexico, 66 Ma (4, 18, 19), leaving a crater occurrence data with paleoclimate and habitat suitability models ∼180 to 200 km in diameter (Fig. 1A). This impactor struck car- to evaluate dinosaur habitability in the wake of various asteroid bonate and sulfate-rich sediments, leading to the ejection and impact and Deccan volcanism scenarios. Asteroid impact models global dispersal of large quantities of dust, ash, sulfur, and other generate a prolonged cold winter that suppresses potential global aerosols into the atmosphere (4, 18–20). These atmospheric dinosaur habitats. -
Capricious Suntime
[Physics in daily life] I L.J.F. (Jo) Hermans - Leiden University, e Netherlands - [email protected] - DOI: 10.1051/epn/2011202 Capricious suntime t what time of the day does the sun reach its is that the solar time will gradually deviate from the time highest point, or culmination point, when on our watch. We expect this‘eccentricity effect’ to show a its position is exactly in the South? e ans - sine-like behaviour with a period of a year. A wer to this question is not so trivial. For ere is a second, even more important complication. It is one thing, it depends on our location within our time due to the fact that the rotational axis of the earth is not zone. For Berlin, which is near the Eastern end of the perpendicular to the ecliptic, but is tilted by about 23.5 Central European time zone, it may happen around degrees. is is, aer all, the cause of our seasons. To noon, whereas in Paris it may be close to 1 p.m. (we understand this ‘tilt effect’ we must realise that what mat - ignore the daylight saving ters for the deviation in time time which adds an extra is the variation of the sun’s hour in the summer). horizontal motion against But even for a fixed loca - the stellar background tion, the time at which the during the year. In mid- sun reaches its culmination summer and mid-winter, point varies throughout the when the sun reaches its year in a surprising way. -
Day-Ahead Market Enhancements Phase 1: 15-Minute Scheduling
Day-Ahead Market Enhancements Phase 1: 15-minute scheduling Phase 2: flexible ramping product Stakeholder Meeting March 7, 2019 Agenda Time Topic Presenter 10:00 – 10:10 Welcome and Introductions Kristina Osborne 10:10 – 12:00 Phase 1: 15-Minute Granularity Megan Poage 12:00 – 1:00 Lunch 1:00 – 3:20 Phase 2: Flexible Ramping Product Elliott Nethercutt & and Market Formulation George Angelidis 3:20 – 3:30 Next Steps Kristina Osborne Page 2 DAME initiative has been split into in two phases for policy development and implementation • Phase 1: 15-Minute Granularity – 15-minute scheduling – 15-minute bidding • Phase 2: Day-Ahead Flexible Ramping Product (FRP) – Day-ahead market formulation – Introduction of day-ahead flexible ramping product – Improve deliverability of FRP and ancillary services (AS) – Re-optimization of AS in real-time 15-minute market Page 3 ISO Policy Initiative Stakeholder Process for DAME Phase 1 POLICY AND PLAN DEVELOPMENT Issue Straw Draft Final June 2018 July 2018 Paper Proposal Proposal EIM GB ISO Board Implementation Fall 2020 Stakeholder Input We are here Page 4 DAME Phase 1 schedule • Third Revised Straw Proposal – March 2019 • Draft Final Proposal – April 2019 • EIM Governing Body – June 2019 • ISO Board of Governors – July 2019 • Implementation – Fall 2020 Page 5 ISO Policy Initiative Stakeholder Process for DAME Phase 2 POLICY AND PLAN DEVELOPMENT Issue Straw Draft Final Q4 2019 Q4 2019 Paper Proposal Proposal EIM GB ISO Board Implementation Fall 2021 Stakeholder Input We are here Page 6 DAME Phase 2 schedule • Issue Paper/Straw Proposal – March 2019 • Revised Straw Proposal – Summer 2019 • Draft Final Proposal – Fall 2019 • EIM GB and BOG decision – Q4 2019 • Implementation – Fall 2021 Page 7 Day-Ahead Market Enhancements Third Revised Straw Proposal 15-MINUTE GRANULARITY Megan Poage Sr. -
The Geologic Time Scale Is the Eon
Exploring Geologic Time Poster Illustrated Teacher's Guide #35-1145 Paper #35-1146 Laminated Background Geologic Time Scale Basics The history of the Earth covers a vast expanse of time, so scientists divide it into smaller sections that are associ- ated with particular events that have occurred in the past.The approximate time range of each time span is shown on the poster.The largest time span of the geologic time scale is the eon. It is an indefinitely long period of time that contains at least two eras. Geologic time is divided into two eons.The more ancient eon is called the Precambrian, and the more recent is the Phanerozoic. Each eon is subdivided into smaller spans called eras.The Precambrian eon is divided from most ancient into the Hadean era, Archean era, and Proterozoic era. See Figure 1. Precambrian Eon Proterozoic Era 2500 - 550 million years ago Archaean Era 3800 - 2500 million years ago Hadean Era 4600 - 3800 million years ago Figure 1. Eras of the Precambrian Eon Single-celled and simple multicelled organisms first developed during the Precambrian eon. There are many fos- sils from this time because the sea-dwelling creatures were trapped in sediments and preserved. The Phanerozoic eon is subdivided into three eras – the Paleozoic era, Mesozoic era, and Cenozoic era. An era is often divided into several smaller time spans called periods. For example, the Paleozoic era is divided into the Cambrian, Ordovician, Silurian, Devonian, Carboniferous,and Permian periods. Paleozoic Era Permian Period 300 - 250 million years ago Carboniferous Period 350 - 300 million years ago Devonian Period 400 - 350 million years ago Silurian Period 450 - 400 million years ago Ordovician Period 500 - 450 million years ago Cambrian Period 550 - 500 million years ago Figure 2. -
The Cenozoic Era - Nýlífsöld 65 MY-Present Jarðsaga 2 Ólafur Ingólfsson Origin of the Term: the Tertiary Tertiary System
The Cenozoic Era - Nýlífsöld 65 MY-Present Jarðsaga 2 Ólafur Ingólfsson Origin of the Term: The Tertiary Tertiary System. [1760] Named by Giovanni Arduino Period as the uppermost part of his 65-1.8 MY three-fold subdivision of mountains in northern Italy. The Tertiary became a formal period and system when Lyell published his work describing further subdivisions of the Tertiary. The Tertiary Period is divided into five epochs (tímar): Paleocene (65-56 MY), Eocene (56-34 MY), Oligocene (34-24 MY), Miocene (24-5,3 MY), and Pliocene (5,3-1,8 MY). Confusing set of stratigraphic terms... More than 95% of the Cenozoic era belongs to the Tertiary period. During the 18th century the names Primary, Secondary, and Tertiary were given by Giovanni Arduino to successive rock strata, the Primary being the oldest, the Tertiary the more recent. In 1829 a fourth division, the Quaternary, was added by P. G. Desnoyers. These terms were later abandoned, the Primary becoming the Paleozoic Era, and the Secondary the Mesozoic. But Tertiary and Quaternary were retained for the two main stages of the Cenozoic. Attempts to replace the "Tertiary" with a more reasonable division of “Palaeogene” (early Tertiary) and “Neogene” (later Tertiary and Quaternary) have not been very successful. Stanley uses this division. The World at the K/T Boundary Paleocene plate tectonics During the Paleocene, the inland seas of the Cretaceous Period dry up, exposing large land areas in North America and Eurasia. Australia begins to separate from Antarctica, and Greenland splits from North America. A remnant Tethys Sea persists in the equatorial region. -
“Anthropocene” Epoch: Scientific Decision Or Political Statement?
The “Anthropocene” epoch: Scientific decision or political statement? Stanley C. Finney*, Dept. of Geological Sciences, California Official recognition of the concept would invite State University at Long Beach, Long Beach, California 90277, cross-disciplinary science. And it would encourage a mindset USA; and Lucy E. Edwards**, U.S. Geological Survey, Reston, that will be important not only to fully understand the Virginia 20192, USA transformation now occurring but to take action to control it. … Humans may yet ensure that these early years of the ABSTRACT Anthropocene are a geological glitch and not just a prelude The proposal for the “Anthropocene” epoch as a formal unit of to a far more severe disruption. But the first step is to recognize, the geologic time scale has received extensive attention in scien- as the term Anthropocene invites us to do, that we are tific and public media. However, most articles on the in the driver’s seat. (Nature, 2011, p. 254) Anthropocene misrepresent the nature of the units of the International Chronostratigraphic Chart, which is produced by That editorial, as with most articles on the Anthropocene, did the International Commission on Stratigraphy (ICS) and serves as not consider the mission of the International Commission on the basis for the geologic time scale. The stratigraphic record of Stratigraphy (ICS), nor did it present an understanding of the the Anthropocene is minimal, especially with its recently nature of the units of the International Chronostratigraphic Chart proposed beginning in 1945; it is that of a human lifespan, and on which the units of the geologic time scale are based. -
North American Coral Recovery After the End-Triassic Mass Extinction, New York Canyon, Nevada, USA
North American coral recovery after the end-Triassic mass extinction, New York Canyon, Nevada, USA Montana S. Hodges* and George D. Stanley Jr., University of INTRODUCTION Montana Paleontology Center, 32 Campus Drive, Missoula, Mass extinction events punctuate the evolution of marine envi- Montana 59812, USA ronments, and recovery biotas paved the way for major biotic changes. Understanding the responses of marine organisms in the ABSTRACT post-extinction recovery phase is paramount to gaining insight A Triassic-Jurassic (T/J) mass extinction boundary is well repre- into the dynamics of these changes, many of which brought sented stratigraphically in west-central Nevada, USA, near New sweeping biotic reorganizations. One of the five biggest mass York Canyon, where the Gabbs and Sunrise Formations contain a extinctions was that of the end-Triassic, which was quickly continuous depositional section from the Luning Embayment. followed by phases of recovery in the Early Jurassic. The earliest The well-exposed marine sediments at the T/J section have been Jurassic witnessed the loss of conodonts, severe reductions in extensively studied and reveal a sedimentological and paleonto- ammonoids, and reductions in brachiopods, bivalves, gastropods, logical record of intense environmental change and biotic turn- and foraminifers. Reef ecosystems nearly collapsed with a reduc- over, which has been compared globally. Unlike the former Tethys tion in deposition of CaCO3. Extensive volcanism in the Central region, Early Jurassic scleractinian corals surviving the end- Atlantic Magmatic Province and release of gas hydrates and other Triassic mass extinction are not well-represented in the Americas. greenhouse gases escalated CO2 and led to ocean acidification of Here we illustrate corals of Early Sinemurian age from Nevada the end-Triassic (Hautmann et al., 2008).